Eutheria

From New World Encyclopedia
Revision as of 22:00, 28 April 2014 by Rick Swarts (talk | contribs)
Eutherians
Fossil range: Late JurassicHolocene, 160–0Ma
Fossil specimen of Eomaia scansoria, an early eutherian
Fossil specimen of Eomaia scansoria, an early eutherian
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Infraclass: Eutheria
Thomas Henry Huxley, 1880
Subgroups
  • Acristatherium
  • Adapisoriculidae
  • Asioryctitheria
  • Bobolestes
  • †Cimolesta[1]
  • Didymoconidae
  • Deltatherium
  • Endotherium
  • Eomaia
  • Juramaia
  • Leptictida
  • Montanalestes
  • Murtoilestes
  • Prokennalestes?
  • Zalambdalestidae
  • Zhelestidae
  • Placentalia (placental mammals)


Eutheria is one of two mammalian clades with extant members that diverged in the Early Cretaceous or perhaps the Late Jurassic. The other is the Metatheria, which includes marsupials, most of whom accommodate their neonates in pouches. Except for the Virginia opossum, which is a metatherian, all mammals indigenous to Europe, Africa, Asia, and North America north of Mexico are eutherians. Extant eutherians, their last common ancestor, and all extinct descendants of that ancestor are placentals, in the group Placentalia.

  • Clade Australosphenida
  • Order Eutriconodonta
  • Order Gobiconodonta
  • Order Multituberculata
  • Clade Trechnotheria

(from Greek ευ-, eu- "true/good" and θηρίον, thērion "beast" hence "true beasts")


Eutherians are distinguished from noneutherians by various features of the feet, ankles, jaws and teeth. One of the major differences between placental and nonplacental eutherians is that placentals lack epipubic bones, which are present in all other fossil and living mammals (marsupials and monotremes).

The oldest known eutherian species is Juramaia sinensis, dated at 160 million years ago from the Jurassic in China.[2]

The name "Eutheria" was introduced by Thomas Henry Huxley in 1880, meant to encompass a more broadly defined group than Placentalia.

Overview

Mammals (class Mammalia {{#invoke:IPAc-en|main}}{{#invoke:IPAc-en|main}}{{#invoke:IPAc-en|main}}/mli.ə/) are a clade of endothermic amniotes distinguished from the reptiles and the birds by the possession of hair, three middle ear bones, mammary glands in females, and a neocortex (a region of the brain). The mammalian brain regulates body temperature and the circulatory system, including the four-chambered heart. The mammals include the largest animals on the planet, the rorquals and some other whales, as well as some of the most intelligent, such as elephants, some primates and some cetaceans. The basic body type is a four-legged land-borne animal, but some mammals are adapted for life at sea, in the air, in the trees, or on two legs. The largest group of mammals, the placentals, have a placenta which feeds the offspring during pregnancy. Mammals range in size from the {{safesubst:#invoke:convert|convert}} bumblebee bat to the 33-meter (110 ft) blue whale.

In some classifications, the mammals are divided into two subclasses (not counting fossils): the Prototheria (order of Monotremata) and the Theria, the latter composed of the infraclasses Metatheria and Eutheria. The marsupials constitute the crown group of the Metatheria and therefore include all living metatherians as well as many extinct ones; the placentals likewise constitute the crown group of the Eutheria.


While the classification of mammals at the family level has been relatively stable, different treatments at higher levels—subclass, infraclass, and order—appear in contemporaneous literature, especially for the marsupials. Much recent change has reflected the results of cladistic analysis and molecular genetics. Results from molecular genetics, for example, have led to the adoption of new groups such as the Afrotheria and the abandonment of traditional groups such as the Insectivora.


Standardized textbook classification

A somewhat standardized classification system has been adopted by most current mammalogy classroom textbooks. The following taxonomy of extant and recently extinct mammals is taken from Vaughan et al. (2000). This approach emphasizes an initial split between egg-laying prototherians and live-bearing therians. The therians are further divided into the marsupial Metatheria and the "placental" Eutheria. No attempt is made here to further distinguish among the orders within these subclasses and infraclasses. This system also makes no note of the position of entirely fossil groups.

In this and later taxonomies listed here, families are merely listed under the order to which they belong. Please see the pages associated with specific orders to see more detailed relationships among families in that order.

Subclass Prototheria

  • Order Monotremata
    • Family Tachyglossidae (echidnas)
    • Family Ornithorhynchidae (platypuses)

Subclass Theria

  • Infraclass Metatheria (marsupials and their nearest ancestors)
    • Order Didelphimorphia
      • Family Didelphidae (opossums, etc.)
    • Order Paucituberculata
      • Family Caenolestidae (shrew opossums)
    • Order Microbiotheria
      • Family Microbiotheriidae (monito del montes)
    • Order Dasyuromorphia (most carnivorous marsupials)
      • Family Thylacinidae (Tasmanian tigers)
      • Family Myrmecobiidae (numbats)
      • Family Dasyuridae (Tasmanian devils, quolls, dunnarts, planigale, etc.)
    • Order Peramelemorphia (bandicoots, bilbies, etc.)
      • Family Peramelidae
      • Family Peroryctidae
    • Order Notoryctemorphia (marsupial moles)
      • Family Notoryctidae
    • Order Diprotodontia
      • Family Phascolarctidae (koalas)
      • Family Vombatidae (wombats)
      • Family Phalangeridae (brushtail possums and cuscuses)
      • Family Potoroidae (bettongs, potaroos and rat kangaroos)
      • Family Macropodidae (kangaroos, wallabies, etc.)
      • Family Burramyidae (pygmy possums)
      • Family Pseudocheiridae (ringtailed possums, etc.)
      • Family Petauridae (Striped Possum, Leadbeater's Possum, Yellow-bellied Glider, Sugar Glider, Mahogany Glider and Squirrel Glider)
      • Family Tarsipedidae (honey possum)
      • Family Acrobatidae (Feathertail Glider and Feather-tailed Possum)
  • Infraclass Eutheria

McKenna/Bell classification

In 1997, the mammals were comprehensively revised by Malcolm C. McKenna and Susan K. Bell, which has resulted in the "McKenna/Bell classification".

McKenna and Bell, Classification of Mammals: Above the species level, (McKenna & Bell, 1997) is the most comprehensive work to date on the systematics, relationships, and occurrences of all mammal taxa, living and extinct, down through the rank of genus. The new McKenna/Bell classification was quickly accepted by paleontologists. The authors worked together as paleontologists at the American Museum of Natural History, New York. McKenna inherited the project from Simpson and, with Bell, constructed a completely updated hierarchical system, covering living and extinct taxa that reflects the historical genealogy of Mammalia.

The McKenna/Bell hierarchical listing of all of the terms used for mammal groups above the species includes extinct mammals as well as modern groups, and introduces some fine distinctions such as legions and sublegions and (ranks which fall between classes and orders) that are likely to be glossed over by the layman.

The published re-classification forms both a comprehensive and authoritative record of approved names and classifications and a list of invalid names.

Click on the highlighted link for a table comparing the traditional and the new McKenna/Bell classifications of mammals

Extinct groups are represented by †.

Subclass Prototheria

(monotremes)

  • Order Platypoda: platypuses
    • Family Ornithorhynchidae: platypuses
  • Order Tachyglossa: echidnas (spiny anteaters)
    • Family Tachyglossidae: echidnas

Subclass Theriiformes

  • Infraclass †Allotheria
    • Order †Multituberculata: multituberculates
      • Family †Plagiaulacidae
      • Family †Bolodontidae
      • Family †Hahnodontidae
      • Family †Albionbaataridae
      • Family †Arginbaataridae
      • Family †Kogaionidae
      • Family †Sloanbaataridae
      • Family †Cimolodontidae
      • Family †Ptilodontidae
      • Family †Cimolomyidae
      • Family †Eucosmodontidae
      • Family †Taeniolabididae
      • Family †Ferugliotheriidae
      • Family †Sudamericidae
  • Infraclass †Triconodonta
    • Family †Austrotriconodontidae
    • Family †Amphilestidae
    • Family †Triconodontidae
  • Infraclass Holotheria
    • Family †Chronoperatidae
    • Superlegion †Kuehneotheria
      • Family †Kuehneotheriidae
      • Family †Woutersiidae
    • Superlegion Trechnotheria
      • Legion †Symmetrodonta
        • Family †Shuotheriidae
        • Order †Amphidontoidea
          • Family †Amphidontidae
        • Order †Spalacotherioidea
          • Family †Tinodontidae
          • Family †Spalacotheriidae
          • Family †Barbereniidae
      • Legion Cladotheria
        • Sublegion †Dryolestoidea
          • Order †Dryolestida
            • Family †Dryolestidae
            • Family †Paurodontidae
            • Family †Donodontidae
            • Family †Mesungulatidae
            • Family †Reigitheriidae
            • Family †Brandoniidae
          • Order †Amphitheriida
            • Family †Amphitheriidae
        • Sublegion Zatheria
          • Family †Arguitheriidae
          • Family †Arguimuridae
          • Family †Vincelestidae
          • Infralegion †Peramura
            • Family †Peramuridae
          • Infralegion Tribosphenida
            • Family †Necrolestidae
            • Supercohort †Aegialodontia
              • Family †Aegialodontidae
            • Supercohort Theria: therian mammals
              • Family †Pappotheriidae
              • Family †Holoclemensiidae
              • Family †Kermackiidae
              • Family †Endotheriidae
              • Family †Picopsidae
              • Family †Potamotelsidae
              • Family †Plicatodontidae
              • Order †Deltatheroida
                • Family †Deltatheridiidae
                • Family †Deltatheroididae
              • Order †Asiadelphia
                • Family †Asiatheriidae
              • Cohort Marsupialia: marsupials
                • Family †Yingabalanaridae
                • Family †Stagodontidae
                • Family †Pediomyidae
                • Magnorder Australidelphia
                  • Superorder Microbiotheria
                    • Family Microbiotheriidae: monito del monte
                  • Superorder Eometatheria
                    • Order †Yalkaparidontia
                      • Family †Yalkaparidontidae
                    • Order Notoryctemorphia: marsupial moles
                      • Family Notoryctidae: marsupial moles
                    • Grandorder Dasyuromorphia: marsupial carnivores
                      • Family †Thylacinidae: recently extinct Tasmanian tiger and relatives
                      • Family Dasyuridae: Tasmanian devil, quolls, etc.
                      • Family Myrmecobiidae: numbat
                    • Grandorder Syndactyli: syndactylous marsupials
                      • Order Peramelia: bandicoots
                        • Family Peramelidae
                        • Family Peroryctidae
                      • Order Diprotodontia
                        • Family †Palorchestidae
                        • Family †Wynardiidae
                        • Family †Thylacoleonidae
                        • Family Tarsipedidae: honey possum
                        • Family †Ilariidae
                        • Family †Diprotodontidae
                        • Family Vombatidae: wombats
                        • Family Phalangeridae: phalangers
                        • Family Burramyidae: pygmy possums
                        • Family Macropodidae: rat kangaroos, kangaroos and wallabies
                        • Family Petauridae: gliders
                        • Family †Ektopodontidae
                        • Family Phascolarctidae: koala
                        • Family †Pilkipildridae
                        • Family †Miralinidae
                        • Family Acrobatidae: feather-tail glider, pen-tailed phalanger
                • Magnorder Ameridelphia
                  • Order Didelphimorphia: opossums
                    • Family Didelphidae: opossums
                    • Family †Sparassocynidae
                  • Order Paucituberculata
                    • Family †Sternbergiidae
                    • Family Caenolestidae: rat or shrew opossums
                    • Family †Paleothentidae
                    • Family †Abderitidae
                    • Family †Sillustaniidae
                    • Family †Polydolopidae
                    • Family †Prepidolopidae
                    • Family †Bonapartheriidae
                    • Family †Argyrolagidae
                    • Family †Patagoniidae
                    • Family †Groeberiidae
                    • Family †Glasbiidae
                    • Family †Caroloameghiniidae
                  • Order †Sparassodonta
                    • Family †Mayulestidae
                    • Family †Hondadelphidae
                    • Family †Borhyaenidae
              • Cohort Placentalia: placentals
                • Order †Bibymalagasia
                • Magnorder Xenarthra: edentates
                  • Order Cingulata: armadillos and relatives
                    • Family Dasypodidae: armadillos
                    • Family †Peltephilidae
                    • Family †Pampatheriidae
                    • Family †Palaeopeltidae
                    • Family †Glyptodontidae: glyptodonts
                  • Order Pilosa: anteaters, sloths, and relatives
                    • Family †Entelopidae
                    • Family Myrmecophagidae: giant anteaters and relatives
                    • Family Cyclopedidae: pygmy anteater
                    • Family †Rathymotheriidae
                    • Family †Scelidotheriidae
                    • Family †Mylodontidae
                    • Family †Megatheriidae: ground sloths
                    • Family Megalonychidae: two-toed sloths
                    • Family Bradypodidae: three-toed sloths
                • Magnorder Epitheria: epitheres
                  • Superorder †Leptictida
                    • Family †Gypsonictopidae
                    • Family †Kulbeckiidae
                    • Family †Didymoconidae
                    • Family †Leptictidae
                  • Superorder Preptotheria
                    • Grandorder Anagalida
                      • Family †Zambdalestidae
                      • Family †Anagalidae
                      • Family †Pseudictopidae
                      • Mirorder Macroscelidea: elephant shrews
                      • Mirorder Duplicidentata
                        • Order †Mimotonida
                          • Family †Mimotonidae
                        • Order Lagomorpha
                      • Mirorder Simplicidentata
                        • Order †Mixodontia
                          • Family †Eurymylidae
                        • Order Rodentia: rodents
                          • Family †Alagomyidae
                          • Family †Laredomyidae
                          • Family †Ischyromyidae
                          • Family †Allomyidae
                          • Family Aplodontiidae: mountain beaver
                          • Family †Mylagaulidae
                          • Family †Theridomyidae
                          • Family †Reithroparamyidae
                          • Family Sciuridae: squirrels
                          • Family †Eutypomyidae
                          • Family Castoridae: beavers
                          • Family †Rhizospalacidae
                          • Family †Protoptychidae
                          • Family †Armintomyidae
                          • Family Dipodidae: jumping mice, jerboas
                          • Family †Simimyidae
                          • Family Muridae: rats, mice, and relatives
                          • Family Myoxidae: dormice
                          • Family †Eomyidae
                          • Family †Florentiamyidae
                          • Family Geomyidae: pocket gophers, pocket mice, and kangaroo rats
                          • Family Pedetidae: springhaas
                          • Family †Parapedetidae
                          • Family †Zegdoumyidae
                          • Family Anomaluridae: scaly-tailed squirrels
                          • Family †Ivanantoniidae
                          • Family †Sciuravidae
                          • Family †Chapattimyidae
                          • Family †Cylindrodontidae
                          • Family Ctenodactylidae: gundis
                          • Family †Tsaganomyidae
                          • Family Hystricidae: Old World porcupines
                          • Family Erethizontidae: New World porcupines
                          • Family †Myophiomyidae
                          • Family †Diamantomyidae
                          • Family †Phiomyidae
                          • Family †Kenyamyidae
                          • Family Petromuridae: rock rats
                          • Family Thryonomyidae: cane rats
                          • Family Bathyergidae: mole-rats
                          • Family †Bathyergoididae
                          • Family Agoutidae: agoutis and pacas
                          • Family †Eocardiidae
                          • Family Dinomyidae: pacarana
                          • Family Caviidae: cavies
                          • Family Hydrochoeridae: capybara
                          • Family Octodontidae: degus, tuco-tucos
                          • Family Echimyidae: spiny rats, nutria
                          • Family Capromyidae: hutias
                          • Family †Heptaxodontidae
                          • Family Chinchillidae: chinchillas, viscachas
                          • Family †Neoepiblemidae
                          • Family Abrocomidae: rat chinchillas
                    • Grandorder Ferae
                      • Order Cimolesta - pangolins and relatives
                        • Family †Palaeoryctidae
                        • Family †Cimolestidae
                        • Family †Apatemyidae
                        • Family †Stylinodontidae
                        • Family †Tillotheriidae
                        • Family †Wangliidae
                        • Family †Harpyodidae
                        • Family †Bemalambdidae
                        • Family †Pastoralodontidae
                        • Family †Titanoideidae
                        • Family †Pantolambdidae
                        • Family †Barylambdidae
                        • Family †Cyriacotheriidae
                        • Family †Pantolambdodontidae
                        • Family †Coryphodontidae
                        • Family †Pantolestidae
                        • Family †Paroxyclaenidae
                        • Family †Ptolemaiidae
                        • Family †Epoicotheriidae
                        • Family †Metacheiromyidae
                        • Family Manidae: pangolins
                        • Family †Ernanodontidae
                      • Order †Creodonta: creodonts
                        • Family †Hyaenodontidae
                        • Family †Oxyaenidae
                      • Order Carnivora
                        • Family †Viverravidae
                        • Family †Nimravidae
                        • Family Felidae: cats
                        • Family Viverridae: civets, Asiatic palm civets
                        • Family Herpestidae: mongooses
                        • Family Hyaenidae: hyaenas, aardwolf
                        • Family Nandiniidae: African palm civets
                        • Family †Miacidae
                        • Family Canidae: dogs
                        • Family †Amphicyonidae
                        • Family Ursidae: bears
                        • Family †Hemicyonidae
                        • Family Otariidae: eared seals
                        • Family Phocidae: seals, walrus
                        • Family Mustelidae: weasels, skunks, and relatives
                        • Family Procyonidae: ringtails, olingos, kinkajou, raccoons, coatis, red panda
                    • Grandorder Lipotyphla
                      • Family †Adapisoriculidae
                        • Order Chrysochloridea
                        • Order Erinaceomorpha
                          • Family †Sespedectidae
                          • Family †Amphilemuridae
                          • Family †Adapisoricidae
                          • Family †Creotarsidae
                          • Family Erinaceidae: hedgehogs and relatives
                          • Family †Proscalopidae
                          • Family Talpidae: moles
                          • Family †Dimylidae
                        • Order Soricomorpha
                          • Family †Otlestidae
                          • Family †Geolabididae
                          • Family †Nesophontidae: recently extinct west Indian shrews
                          • Family †Micropternodontidae
                          • Family †Apternodontidae
                          • Family Solenodontidae: solenodons
                          • Family †Plesiosoricidae
                          • Family †Nyctitheriidae
                          • Family Soricidae: shrews
                          • Family Tenrecidae: tenrecs
                    • Grandorder Archonta
                      • Order Chiroptera: bats
                        • Family Pteropodidae: flying foxes
                        • Family †Archaeonycteridae
                        • Family †Paleochiropterygidae
                        • Family †Hassianycterididae
                        • Family Emballonuridae: sac-winged bats
                        • Family Rhinopomatidae: mouse-tailed bats
                        • Family Craseonycteridae: bumblebee bats
                        • Family Megadermatidae: false vampire bats
                        • Family Nycteridae: hispid bats
                        • Family Rhinolophidae: horseshoe and Old World leaf-nosed bats
                        • Family Mystacinidae: New Zealand short-tailed bats
                        • Family Noctilionidae: fishing bats
                        • Family Mormoopidae: spectacled bats
                        • Family Phyllostomidae: New World leaf-nosed and vampire bats
                        • Family †Philisidae
                        • Family Molossidae: free-tailed bats
                        • Family Natalidae: funnel-eared bats
                        • Family Furipteridae: smoky bats
                        • Family Thyropteridae: New World sucker-footed bats
                        • Family Myzopodidae: Old World sucker-footed bats
                        • Family Vespertilionidae: common bats
                      • Order Primates: primates
                        • Family †Purgatoriidae
                        • Family †Microsyopidae
                        • Family †Micromomyidae
                        • Family †Picromomyidae
                        • Family †Plesiadapidae
                        • Family †Palaechthonidae
                        • Family †Picrodontidae
                        • Family †Paramomyidae
                        • Family †Plagiomenidae
                        • Family †Mixodectidae
                        • Family Galeopithecidae: colugos
                        • Family †Plesiopithecidae
                        • Family Daubentoniidae: aye-aye
                        • Family †Adapidae
                        • Family Lemuridae: lemurs
                        • Family Lorisidae: lorises and galagos
                        • Family Cheirogaleidae: dwarf lemurs
                        • Family †Archaeolemuridae
                        • Family †Palaeopropithecidae
                        • Family Indriidae: indris and sifakas
                        • Family †Carpolestidae
                        • Family †Omomyidae
                        • Family †Microchoeridae
                        • Family †Afrotarsiidae
                        • Family Tarsiidae: tarsiers
                        • Family †Eosimiidae
                        • Family †Parapithecidae
                        • Family †Pliopithecidae
                        • Family Cercopithecidae: Old World monkeys including colobuses
                        • Family Hominidae: humans, greater apes, lesser apes
                        • Family Callitrichidae: marmosets
                        • Family Atelidae: New World monkeys
                      • Order Scandentia
                        • Family Tupaiidae: tree shrews
                    • Grandorder Ungulata: ungulates
                      • Order Tubulidentata
                        • Family Orycteropodidae: aardvark
                      • Order †Dinocerata
                        • Family †Uintatheriidae
                      • Mirorder Eparctocyona
                        • Order †Procreodi
                          • Family †Oxyclaenidae
                          • Family †Arctocyonidae
                        • Order †Condylarthra
                          • Family †Hyopsodontidae
                          • Family †Mioclaenidae
                          • Family †Phenacodontidae
                          • Family †Periptychidae
                          • Family †Peligrotheriidae
                          • Family †Didolodontidae
                        • Order †Arctostylopida
                          • Family †Arctostylopidae
                        • Order Cete: whales and relatives
                          • Family †Triisodontidae
                          • Family †Mesonychidae: mesonychids
                          • Family †Hapalodectidae
                          • Family †Basilosauridae
                          • Family †Protocetidae
                          • Family †Remingtonocetidae
                          • Family †Agorophiidae
                          • Family †Squalodontidae
                          • Family †Rhabdosteidae
                          • Family †Aetiocetidae
                          • Family †Mammalodontidae
                          • Family †Cetotheriidae
                          • Family Balaenopteridae: rorquals and grey whales
                          • Family Balaenidae: right and bowhead whales
                          • Family Physeteridae: sperm whales
                          • Family Hyperoodontidae: beaked whales
                          • Family Platanistidae: river dolphins
                          • Family Delphinidae: dolphins
                          • Family Pontoporiidae: La Plata River dolphin
                          • Family Lipotidae: baiiji
                          • Family Iniidae: Amazon River dolphin
                          • Family †Kentridontidae
                          • Family Monodontidae: beluga and narwhal
                          • Family †Odobenocetopsidae
                          • Family †Dalpiazinidae
                          • Family †Acrodelphinidae
                          • Family Phocoenidae: porpoises
                          • Family †Albireonidae
                          • Family †Hemisyntrachelidae
                        • Order Artiodactyla: even-toed ungulates
                          • Family †Raoellidae
                          • Family †Choeropotamidae
                          • Family Suidae: pigs
                          • Family Tayassuidae: peccaries
                          • Family †Santheriidae
                          • Family Hippopotamidae: hippos
                          • Family †Dichobunidae
                          • Family †Cebochoeridae
                          • Family †Mixtotheriidae
                          • Family †Helohyidae
                          • Family †Haplobunodontidae
                          • Family †Anthracotheriidae
                          • Family †Dacrytheriidae
                          • Family †Anoplotheriidae
                          • Family †Cainotheriidae
                          • Family †Agriochoeridae
                          • Family †Oreodontidae
                          • Family †Entelodontidae
                          • Family †Xiphodontidae
                          • Family Camelidae: camels and llamas
                          • Family †Oromerycidae
                          • Family †Protoceratidae
                          • Family †Amphimerycidae
                          • Family †Hypertragulidae
                          • Family Tragulidae: mouse deer
                          • Family †Leptomerycidae
                          • Family †Bachitheriidae
                          • Family †Lophiomerycidae
                          • Family †Gelocidae
                          • Family Moschidae: musk deer
                          • Family Antilocapridae: pronghorn
                          • Family †Palaeomerycidae
                          • Family †Hoplitomerycidae
                          • Family Cervidae: deer
                          • Family †Climacoceratidae
                          • Family Giraffidae: giraffe and okapi
                          • Family Bovidae: cattle, antelope, and relatives
                      • Mirorder †Meridiungulata
                        • Family †Perutheriidae
                        • Family †Amilnedwardsiidae
                        • Order †Litopterna
                          • Family †Protolipternidae
                          • Family †Macraucheniidae
                          • Family †Notonychopidae
                          • Family †Adianthidae
                          • Family †Proterotheriidae
                        • Order †Notoungulata: notoungulates
                          • Family †Henricosborniidae
                          • Family †Notostylopidae
                          • Family †Isotemnidae
                          • Family †Leontiniidae
                          • Family †Notohippidae
                          • Family †Toxodontidae
                          • Family †Homalodotheriidae
                          • Family †Archaeopithecidae
                          • Family †Oldfieldthomasiidae
                          • Family †Interatheriidae
                          • Family †Campanorcidae
                          • Family †Mesotheriidae
                          • Family †Archaeohyracidae
                          • Family †Hegetotheriidae
                        • Order †Astrapotheria
                          • Family †Eoastrapostylopidae
                          • Family †Trigonostylopidae
                          • Family †Astrapotheriidae
                        • Order †Xenungulata
                          • Family †Carodniidae
                        • Order †Pyrotheria
                          • Family †Pyrotheriidae
                      • Mirorder Altungulata
                        • Order Perissodactyla: odd-toed ungulates
                          • Family Equidae: horses
                          • Family †Palaeotheriidae
                          • Family †Brontotheriidae
                          • Family †Anchilophidae
                          • Family †Eomoropidae
                          • Family †Chalicotheriidae
                          • Family †Hyracodontidae
                          • Family Rhinocerotidae: rhinoceroses
                          • Family †Helaletidae
                          • Family †Isectolophidae
                          • Family †Lophiodontidae
                          • Family †Deperetellidae
                          • Family †Lophialetidae
                          • Family Tapiridae: tapirs
                        • Order Uranotheria: elephants, manatees, hyraxes, and relatives
                          • Family †Pliohyracidae
                          • Family Procaviidae: hyraxes
                          • Family †Phenacolophidae
                          • Family †Arsinoitheriidae
                          • Family †Prorastomidae
                          • Family Dugongidae: dugongs
                          • Family Trichechidae: manatees
                          • Family †Desmostylidae
                          • Family †Anthracobunidae
                          • Family †Moeritheriidae
                          • Family †Numidotheriidae
                          • Family †Barytheriidae
                          • Family †Deinotheriidae
                          • Family †Palaeomastodontidae
                          • Family †Phiomiidae
                          • Family †Hemimastodontidae
                          • Family †Mammutidae: mastodons and relatives
                          • Family †Gomphotheriidae: gomphotheres
                          • Family Elephantidae: modern elephants


Characteristics

The features of Eutheria that distinguish them from metatherians, a group that includes modern marsupials, are:

  • an enlarged malleolus ("little hammer") at the bottom of the tibia, the larger of the two shin bones.[3]
  • the joint between the first metatarsal bone and the entocuneiform bone in the foot is offset further back than the joint between the second metatarsal and middle cuneiform bones – in metatherians these joints are level with each other.[3]
  • various features of jaws and teeth.[3]

Evolutionary history

Eutheria contains several extinct genera and larger groups, many with complicated taxonomic history that is still unresolved. Members of the Adapisoriculidae, Cimolesta and Leptictida have been previously placed within the out-dated placental group "Insectivora", while Zhelestids have been considered primitive ungulates.[4] However, more recent studies have suggested these enigmatic taxa represent stem group eutherians, more basal to Placentalia.[5][6]

The fossil eutherian species believed to be the oldest known is Juramaia sinensis, which lived about 160 million years ago.[2] Montanalestes was found in North America, while all other nonplacental eutherian fossils have been found in Asia. The earliest known placental fossils have also been found in Asia.[3]
  • Cynodonts
    • † Other Cynodonts
    • † Tritylodontids
    • Mammaliaforms
      • † Other mammaliaforms
      • Hadrocodium
      • Crown-group mammals
        • † Other crown-group mammals
        • Australosphenids
        • Theria
          • † Other Theria
          • Metatheria
          • Eutheria
            • † Other Eutheria
            • Placentals
Simplified, non-systematic, outline of evolution of eutheria from cynodont therapsids.[3]
† = extinct

References
ISBN links support NWE through referral fees

  1. D. L. Rock, and J. P. Hunter, "Rooting Around the Eutherian Family Tree: The Origin and Relations of the Taeniodonta," Journal of Mammalian Evolution 21(issue 1)(2014): 75-91.
  2. 2.0 2.1 Luo, Z. and C. Yuan, Q. Meng, and Q. Ji (2011). A Jurassic eutherian mammal and divergence of marsupials and placentals. Nature 476 (7361): 42–45.
  3. 3.0 3.1 3.2 3.3 3.4 Ji, Q., Luo, Z-X., Yuan, C-X.,Wible, J.R., Zhang, J-P. and Georgi, J.A. (April 2002). The earliest known eutherian mammal. Nature 416 (6883): 816–822.
  4. Rose, Kenneth D. (2006). The beginning of the age of mammals. Baltimore: Johns Hopkins University Press. ISBN 9780801892219. 
  5. Wible, J. R. and Rougier, G. W.; Novacek, M. J.; Asher, R. J. (2007). Cretaceous eutherians and Laurasian origin for placental mammals near the K/T boundary. Nature 447 (7147): 1003–1006.
  6. Wible, John R. and Rougier, Guillermo W.; Novacek, Michael J.; Asher, Robert J. (2009). The Eutherian Mammal Maelestes gobiensis from the Late Cretaceous of Mongolia and the phylogeny of cretaceous eutheria. Bulletin of the American Museum of Natural History 327: 1–123.
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Mammals
Monotremata (platypus, echidnas)

Marsupialia: | Paucituberculata (shrew opossums) | Didelphimorphia (opossums) | Microbiotheria | Notoryctemorphia (marsupial moles) | Dasyuromorphia (quolls and dunnarts) | Peramelemorphia (bilbies, bandicoots) | Diprotodontia (kangaroos and relatives)

Placentalia: Cingulata (armadillos) | Pilosa (anteaters, sloths) | Afrosoricida (tenrecs, golden moles) | Macroscelidea (elephant shrews) | Tubulidentata (aardvark) | Hyracoidea (hyraxes) | Proboscidea (elephants) | Sirenia (dugongs, manatees) | Soricomorpha (shrews, moles) | Erinaceomorpha (hedgehogs and relatives) Chiroptera (bats) | Pholidota (pangolins)| Carnivora | Perissodactyla (odd-toed ungulates) | Artiodactyla (even-toed ungulates) | Cetacea (whales, dolphins) | Rodentia (rodents) | Lagomorpha (rabbits and relatives) | Scandentia (treeshrews) | Dermoptera (colugos) | Primates |

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