Difference between revisions of "Dog" - New World Encyclopedia

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| trinomial = ''Canis lupus familiaris''
 
| trinomial = ''Canis lupus familiaris''
 
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{{Portal|Dogs|HappyMutt.jpg}}
 
[[Image:Dog.jpg|thumb]]
 
  
The '''dog''' (''Canis lupus familiaris'') is a domestic [[subspecies]] of the [[Gray Wolf|wolf]], a [[mammal]] of the [[Canidae]] family of the order [[Carnivora]]. The term encompasses both [[pariah dog|feral]] and [[pet]] variants. It is also sometimes used to describe [[wild dog|wild canid]]s of other subspecies or species.  
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The '''dog''' ''(Canis lupus familiaris'' or ''Canis familiaris)'' is usually considered to be humankind's first, and perhaps most important, [[domestic animal]]. It generally is classified as a [[subspecies]] of the [[gray wolf]] ''(Canis lupus)'', a [[mammal]] of the [[Canidae]] family of the order [[Carnivora]].
  
Over time, the dog has developed into hundreds of [[Dog breed|breeds]] with a great degree of variation.  For example, heights at the [[withers]] range from just a few inches (such as the [[Chihuahua (dog)|Chihuahua]]) to roughly three feet (such as the [[Irish Wolfhound]]); colors vary from white through grays (usually called ''blue'') to black, and browns from light (tan) to dark ("red" or "chocolate") in a tremendous variation of patterns; and [[Coat (dog)|coat]]s can be anything from very short to several centimeters long, from coarse hair to something akin to wool, straight or curly, or smooth.
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The word "dog" also is used to refer to some other members of the Canidae family, for instance the bush dog ''(Speothos venaticus)'' of [[South America]] and the raccoon dog ''(Nyctereutes procyonoides)'' of [[Asia]].  The family Canidae itself often is called the "dog family" and includes [[jackal]]s and [[coyote]]s, which are closely related to dogs and wolves, as well as [[fox]]es, which are less closely related.  Members of the Canidae family are called "canids."
  
== Overview ==
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The trainability, devotion, playfulness, and ability to fit into human households have earned dogs a unique position in the realm of interspecies relationships. Dogs have lived and worked with humans in so many roles—hunting, herding and protecting [[sheep]] and [[cattle]], guarding people and property, rescuing people, companionship, and so forth—that their loyalty has earned them the unique sobriquet, "man's best friend." Their actions mimic the human idea of [[love]] and friendship, leading many dog owners to view their pets as family members.
=== Origins ===
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{{toc}}
{{main|Origin of the domestic dog}}
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In a process generally thought to span at least 14,000 years, humans have domesticated the dog, produced innumerable breeds, and bred and trained them to perform a great variety of useful services and sometimes dubious actions, including as guard dogs trained to instill fear in people and as sports dogs taught to fight other dogs to the death.
 +
[[File:Dog anatomy lateral skeleton view.jpg|thumb|Lateral view of a dog's bone structure.]]
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== Characteristics ==
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Dogs, like [[wolf|wolves]] and other canids, are specialized for a life of hunting and eating other animals. Their bodies are compact and muscular and their legs are long, which gives them speed and endurance for chasing prey. Their jaws are large and their teeth are well suited for grasping, biting, and tearing flesh.  Their [[digestive system]]s, however, are suited for some [[plant]] foods as well as flesh.
  
Some research appears to show that dogs were [[Domestication|domesticated]] from [[Gray Wolf|wolves]] as recently as 15,000 years ago,<ref>{{cite web
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Most dogs are covered with a thick coat of [[fur]], which protects them from the elementsAlaskan malamutes and Siberian huskies, which originated in the [[Arctic]], are able to live outside without shelter in temperatures far below freezing. Dogs' coats come in a wide range of colors, including black, white, brown, yellow, red, and gray. They come in a range of lengths and textures from very short to long, either straight or curly (AKC 2006).   
| url = http://news.bbc.co.uk/2/hi/science/nature/2498669.stm
 
| title = Origin of dogs traced
 
| accessdate = 2006-11-29
 
| last = McGourty
 
| first = Christine
 
| date = [[2002-11-22]]
 
| publisher = BBC News
 
}}</ref> or perhaps as early as 100,000 years ago based upon recent genetic, [[fossil]] and [[DNA]] evidence.<ref>Vilà, C. et al. (1997).</ref> Other research suggests that dogs have only been domesticated for a much shorter amount of time and were domesticated from populations of wild dogs, which had previously diverged from wolves.<ref>{{cite journal
 
| last = Wayne
 
| first = Robert K.
 
| coauthors = Carles Vila, Peter Savolainen, Jesus E. Maldonado, Isabel R. Amorim, John E. Rice, Rodney L. Honeycutt, Keith A. Crandall, Joakim Lundeberg
 
| date = [[1997-01-30]]; accepted [[1997-04-14]]
 
| title = Multiple and ancient origins of the domestic dog
 
| journal = Science
 
| volume = 276
 
| pages = 1687-1689
 
| url = http://www.mnh.si.edu/GeneticsLab/StaffPage/MaldonadoJ/PublicationsCV/Science_Dog_Paper.pdf
 
| format = pdf
 
| accessdate = 2006-12-09
 
}}</ref><ref>{{cite journal
 
| last = Kerstin
 
| first = Lindblad-Toh
 
| coauthors = Claire M Wade, Tarjei S. Mikkelsen, Elinor K. Karlsson, David B. Jaffe, Michael Kamal, Michele Clamp, Jean L. Chang, Edward J. Kulbokas III, Michael C. Zody, Evan Mauceli, Xiaohui Xie, Matthew Breen, Robert K. Wayne, Elaine A. Ostrander, Chris P. Ponting, Francis Galibert, Douglas R. Smith, Pieter J. deJong, Ewen Kirkness, Pablo Alvarez, Tara Biagi, William Brockman, Jonathan Butler, Chee-Wye Chin, April Cook, James Cuff, Mark J. Daly, David DeCaprio, Sante Gnerre, Manfred Grabherr, Manolis Kellis, Michael Kleber, Carolyne Bardeleben, Leo Goodstadt, Andreas Heger, Christophe Hitte, Lisa Kim, Klaus-Peter Koepfli, Heidi G. Parker, John P. Pollinger, Stephen M. J. Searle, Nathan B. Sutter, Rachael Thomas, Caleb Webber
 
| date = [[2005-12-08]]
 
| title = Genome sequence, comparative analysis and haplotype structure of the domestic dog
 
| journal = Nature
 
| volume = 438
 
| pages = 803-819
 
| url = http://www.nature.com/nature/journal/v438/n7069/abs/nature04338.html
 
}}</ref>  
 
  
New evidence suggests that dogs were first domesticated in East Asia, possibly China,<ref name=Savolainen_et_al_2002>{{cite journal
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A dog's [[heart]] and [[lung]]s are oversized relative to its body and its normal everyday needs. A dog also has relatively more [[red blood cell]]s than a human. Most of the time the dog will keep the extra red blood cells stored in its [[spleen]]. When the animal enters into a situation where its full [[metabolism]] is required, such as play, catching game, or fighting other dogs, the extra cells are released into the bloodstream.  The "oversized" heart and lungs would then be running at full capacity, and the animal would have an enhanced ability to engage in aerobic activity. This activity produces internal heat. Dogs, being covered in fur, are limited in their ability to cool down. After a short time the animal must either cease its athletic activity or risk harming itself from overheating. One can easily observe this pattern of intense activity followed by rest periods in puppiesDuring the rest phase, the spleen collects red blood cells and the animal may pant to cool down.
| first = Peter
 
| last = Savolainen
 
| coauthors = Ya-ping Zhang, Jing Luo, Joakim Lundeberg, and Thomas Leitner
 
| date = [[2002-11-22]]
 
| title = Genetic Evidence for an East Asian Origin of Domestic Dogs
 
| journal = Science
 
| volume = 298
 
| issue = 5598
 
| pages = 1610 - 1613
 
| doi = 10.1126/science.1073906
 
| url = http://adsabs.harvard.edu/abs/2002Sci...298.1610S
 
}}</ref> and the first peoples to enter North America took dogs with them from Asia. Genetic research has identified 14 [[ancient dog breeds]], with the oldest being the [[Chow Chow]], [[Shar Pei]], [[Akita Inu]], [[Shiba Inu]] and [[Basenji]]Because many of the 14 breeds are associated with China and Japan, the theory that the dog originated in Asia seems to be likely.<ref name=Savolainen_et_al_2002/>
 
  
As humans migrated around the planet a variety of dog forms migrated with them.  The [[agricultural revolution]] and subsequent [[urban revolution]] led to an increase in the dog population and a demand for [[Specialization (functional)|specialization]].  These circumstances would provide the opportunity for [[selective breeding]] to create specialized [[working dog]]s and [[pet]]s.
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Dogs, like most [[carnivore]]s, are naturally active only 4 to 6 hours each day and prefer to sleep the rest of the time (Kruuk 2002).
  
=== Human relationships ===
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Dogs, like other canids, are considered to be among the most intelligent of all animals, as shown by their ability to adapt to changing conditions in human society.  
[[Image:Sheltie whisky.jpg|thumbnail|[[Shetland Sheepdog]]s are well known for their loyal companionship and intelligence.]]
 
Dogs are highly [[social animal]]s due to selective breeding. This can account for their trainability, playfulness, and ability to fit into human households and social situations.  This similarity has earned dogs a unique position in the realm of interspecies relationships.
 
  
The loyalty and devotion that dogs demonstrate as part of their natural instincts as pack animals closely mimics the human idea of love and friendship, leading many dog owners to view their pets as full-fledged family membersConversely, dogs seem to view their human companions as members of their pack, and make few, if any, distinctions between their owners and fellow dogs.  Dogs fill a variety of roles in [[human society]] and are often trained as [[working dog]]s.  For dogs that do not have traditional jobs, a wide range of [[dog sports]] provide the opportunity to exhibit their natural skills.  In many countries, the most common and perhaps most important role of dogs is as [[companion dog|companions]].
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The dog's ancestor, the gray wolf, is the largest canid; with adults weighing between 18 and 80 kg (40 to 175 lbs)Modern dogs can weigh anywhere from 1 to over 100 kg (2.2 to 220 lbs). Dingos, wild dogs of [[Australia]] which are descended from domestic dogs, average 10 to 20 kg (22 to 44 lbs) in weight (Nowak 1983).
  
Dogs have lived and worked with humans in so many roles that their loyalty has earned them the unique [[sobriquet]] "man's best friend".<ref>{{cite web
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===Senses===
| url = http://www.almostheaven-golden-retriever-rescue.org/old-drum.html
 
| title = The Story of Old Drum
 
| accessdate = 2006-11-29
 
| publisher = Cedarcroft Farm Bed & Breakfast - Warrensburg, MO
 
}}</ref>  However, some cultures consider dogs to be [[unclean animals|unclean]].  In some parts of the world, dogs are raised as [[livestock]] to produce [[dog meat]] for human consumption.  In many places, consumption of dog meat is discouraged by social convention or [[Taboo food and drink|cultural taboo]].
 
  
==Terminology==  
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====Sight====
The [[English language|English]] word ''[[wikt:dog|dog]]'', in common usage, refers to the domestic [[pet]] dog, ''Canis lupus familiaris''. The species was originally classified as ''Canis familiaris'' by Linnaeus in 1758. In 1993, dogs were reclassified as a subspecies of the gray wolf, ''Canis lupus'', by the [[Smithsonian Institution]] and the [[American Society of Mammalogists]]. "Dog" is sometimes used to refer collectively to any mammal belonging to the family ''[[Canidae]]'' (as in "the dog family"), such as wolves, foxes, and coyotes. Some members of the family have "dog" in their common names, such as the [[African Wild Dog]]. The constellations [[Canes Venatici]], [[Canis Major]] and [[Canis Minor]] are named from the [[Latin language|Latin]] word for "dog," for their perceived resemblance to dogs.  
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In general, the vision of dogs is not as good as that of humans. They have only limited color vision, equivalent to red-green [[color blindness]] in humans (Alderton 1984).  They also are nearsighted compared to humans and can not see detail as well. The average dog's vision seems to be about 20/50 to 20/100, using the human standard. On the other hand they see better in dim light and may also detect motion better than humans (Davis 1998).
  
The English word ''dog'' might derive from the Old English [[wikt:docga|docga]], a "powerful breed of canine". The French ''dogue'' and Spanish ''dogo'' as in [[dogo Argentino]] are borrowings from English. The English word ''hound'' is a cognate of German ''Hund'', Dutch ''hond'', common Scandinavian ''hund'', Icelandic ''hundur'' which, though referring to a specific breed in English, means "dog" in general in the other Germanic languages. ''Hound'' itself derives from the [[Proto-Indo-European language|Proto-Indo-European]] *kwon-, which is the direct root of the Greek κυων (kuōn) and the indirect root of the Latin [[canis]] through the variant form *kani-.<Ref>
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====Hearing====
{{cite web
 
| url = http://www.bartleby.com/61/roots/IE259.html
 
| title = The American Heritage® Dictionary of the English Language: Fourth Edition.
 
| accessdate = 2006-11-30
 
| publisher = www.bartleby.com
 
}}</Ref>
 
 
 
In breeding circles, a male canine is referred to as a ''dog'', while a female canine is called a ''bitch''. The father of a litter is called the ''sire'', and the mother of a litter is called the ''dam''. Offspring are generally called ''pups'' or ''[[puppy|puppies]]'' until they are about a year old. A group of offspring is a ''litter''. The process of birth is ''whelping''. Many [[Mixed-breed dog#Terms for mixed-breed dogs|terms are used for dogs]] that are not purebred.
 
 
 
==Intelligence==
 
{{main|Dog intelligence}}
 
[[Image:Dog retrieving stick.jpg|thumb|179px|Many dogs can easily be trained to retrieve.]]
 
Among dog lovers, dogs are generally valued for their intelligence, and both [[anecdotal evidence]] and [[Scientific method|scientific research]] suggest that dogs have a reasonably high intelligence. This intelligence is expressed differently with different breeds and individuals, however. For example, [[Border Collie]]s are noted for their ability to learn commands, while other breeds may not be so motivated towards obedience, but instead show their cleverness in devising ways to steal food or escape from a yard. Being highly adaptable animals themselves, dogs have learned to do many jobs as required by humans over the generations. Dogs are employed in various roles across the globe, proving invaluable assets in areas such as search-and-rescue; law enforcement (including attack dogs, sniffer dogs and tracking dogs); guards for livestock, people or property; herding; Arctic exploration sled-pullers; guiding the blind and acting as a pair of ears for the deaf; assisting with hunting, and a great many other roles which they may be trained to assume. 
 
Most dogs rarely have to deal with complex tasks and are unlikely to learn relatively complicated activities (such as opening doors) unaided. Some dogs (such as [[guide dog]]s for the visually impaired) are specially trained to recognize and avoid dangerous situations.
 
 
 
===Evaluation of a dog's intelligence===
 
The meaning of "intelligence" in general, not only in reference to dogs, is hard to define. Some tests measure problem-solving abilities and others test the ability to learn in comparison to others of the same age.  Defining it for dogs is just as difficult.  It is likely that dogs do not have the ability to premeditate an action to solve a problem. Some dogs may, however, have more drive to keep trying various things until they accidentally reach a solution and still others might have more ability to make the association between the "accident" and the result.{{Fact|date=February 2007}}
 
 
 
For example, the ability to learn quickly could be a sign of [[intelligence (trait)|intelligence]]. Conversely it could be interpreted as a sign of blind subservience and a desire to please.  In contrast, some dogs who do not learn very quickly may have other talents. An example is breeds that are not particularly interested in pleasing their owners, such as [[Siberian Husky|Siberian Huskies]]. Huskies are often fascinated with the myriad of possibilities for escaping from yards, catching small animals, and often figuring out on their own numerous inventive ways of doing both.
 
 
 
[[Assistance dog]]s are also required to be obedient at all times. This means they must learn a tremendous number of commands, understand how to act in a large variety of situations, and recognize threats to their human companion, some of which they might never before have encountered.
 
 
 
Many owners of [[livestock guardian dog|livestock guardian]] breeds believe that breeds like the [[Great Pyrenees]] or the [[Kuvasz]] are not easily trained because their stubborn nature prevents them from seeing the point of such commands as “sit” or “down”. [[Hounds]] may also suffer from this type of ranking. In fact several rank in the bottom tier of "The Intelligence of Dogs" list (such as [[Beagle]]s, [[Bloodhound]]s, and [[Basset Hound]]s).{{Fact|date=February 2007}}  These dogs are bred to have more of a "pack" mentality with other dogs and less reliance on a master's direct commands. While they may not have the same kind of intelligence as a Border Collie, they were not bred to learn and obey commands quickly, but to think for themselves while trailing game.
 
 
 
==Physical characteristics==
 
{{main|Dog anatomy}}
 
[[Image:Weimaraner wb.jpg|thumb|right|Some dogs, like this [[Weimaraner]], have their predatory instincts suppressed in order to better assist human hunters.]]
 
{{see also|Dog health}}
 
Modern [[dog breed]]s show more variation in size, appearance, and behavior than any other domestic animal. Within the range of extremes, dogs generally share attributes with their wild ancestors, the [[Gray Wolf|wolves]]. Dogs are [[predator]]s and [[scavenger]]s, possessing sharp teeth and strong jaws for attacking, holding, and tearing their food. Although selective breeding has changed the appearance of many breeds, all dogs retain basic traits from their distant ancestors. Like many other predatory mammals, the dog has powerful muscles, fused wristbones, a [[cardiovascular]] system that supports both sprinting and endurance, and teeth for catching and tearing. Compared to the bone structure of the human foot, dogs technically walk on their toes.
 
 
 
===Sight===
 
[[Image:Amazonian Dog.JPG|left|A|thumb|200px|Amazonian dogs, such as this one from [[Alto Amazonas]], [[Peru]] are critical for [[indigenous peoples]]' hunting strategies, particularly in the [[rain forest]].]]
 
 
 
Like most mammals, dogs are [[dichromat]]s and have color vision equivalent to red-green [[color blindness]] in humans.<ref name="note1">A&E Television Networks (1998). ''Big Dogs, Little Dogs: The companion volume to the A&E special presentation'', A Lookout Book, GT Publishing. ISBN 1-57719-353-9 (hardcover).</ref><ref name="note2">Alderton, David (1984). ''The Dog'', Chartwell Books. ISBN 0-89009-786-0.</ref>
 
 
 
Different breeds of dogs have different eye shapes and dimensions, and they also have different [[retina]] configurations.<ref>{{cite web | last = | first = | authorlink = | coauthors = Jonica Newby, Caroline Penry-Davey | title = Catalyst: Dogs' Eyes | work = | publisher = Australian Broadcasting Corporation | date = [[2003-09-25]] | url = http://www.abc.net.au/catalyst/stories/s953902.htm | format = | doi = | accessdate = 2006-11-26 }}</ref>  Dogs with long noses have a "''visual streak''" which runs across the width of the retina and gives them a very wide field of excellent vision, while those with short noses have an "''area centralis''" &mdash; a central patch with up to three times the density of nerve endings as the ''visual streak'' &mdash; giving them detailed sight much more like a human's.
 
 
 
Some breeds, particularly the best [[sighthound]]s, have a field of vision up to 270° (compared to 180&deg; for humans), although broad-headed breeds with short noses have a much narrower field of vision, as low as 180°.<ref name="note1" /><ref name="note2" />
 
 
 
===Hearing===
 
 
[[Image:PennyAlertPricksEars.jpg|thumb|right|This [[mixed-breed dog]] angles her ears to find the source of a sound.]]
 
[[Image:PennyAlertPricksEars.jpg|thumb|right|This [[mixed-breed dog]] angles her ears to find the source of a sound.]]
Dogs detect sounds as low as the 16 to 20 [[Hertz|Hz]] frequency range (compared to 20 to 70 Hz for humans) and above 45 kHz<ref>[http://www.lsu.edu/deafness/HearingRange.html How well do dogs and other animals hear]</ref> (compared to 13 to 20 kHz for humans),<ref name="note2" /> and in addition have a degree of ear mobility that helps them to rapidly pinpoint the exact location of a sound. Eighteen or more muscles can tilt, rotate and raise or lower a dog's ear. Additionally, a dog can identify a sound's location much faster than a human can, as well as hear sounds up to four times the distance that humans are able to. Those with more natural ear shapes, like those of wild canids like the fox, generally hear better than those with the floppier ears of many domesticated species.
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The hearing of dogs is superior to humans. They can detect sounds that are lower and those that are higher in frequency. They also have a degree of ear mobility that helps them to rapidly pinpoint the exact location of a sound. Eighteen or more muscles can tilt, rotate, and raise or lower a dog's [[ear]]. Additionally, a dog can identify a sound's location much faster than a human can, as well as hear sounds up to four times the distance that humans are able to. Those with more natural ear shapes, like those of wild canids, generally hear better than those with the floppy ears of many domesticated species.
  
===Smell===
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====Smell====
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[[Image:Bloodhund r54.jpg|left|thumb|Scent hounds, especially the Bloodhound, are known for their keen sense of smell]]
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Dogs have nearly 220 million smell-sensitive [[cell (biology)|cells]] over an area about the size of a pocket [[handkerchief]] (compared to 5 million over an area the size of a [[postage stamp]] for humans).  Some breeds have been selectively bred for excellence in detecting scents, even compared to their canine brethren.
  
Dogs have nearly 220 million smell-sensitive cells over an area about the size of a pocket [[handkerchief]] (compared to 5 million over an area the size of a [[postage stamp]] for humans).  Some breeds have been selectively bred for excellence in detecting scents, even compared to their canine brethren. What information a dog actually detects when he is scenting is not perfectly understood; although once a matter of debate, it now seems to be well established that dogs can distinguish two different types of scents when trailing, an air scent from some person or thing that has recently passed by, as well as a ground scent that remains detectable for a much longer period. The characteristics and behavior of these two types of scent trail would seem, after some thought, to be quite different, the air scent being intermittent but perhaps less obscured by competing scents, whereas the ground scent would be relatively permanent with respect to careful and repetitive search by the dog, but would seem to be much more contaminated with other scents. In any event, it is established by those who train tracking dogs that it is impossible to teach the dog how to track any better than it does naturally; the object instead is to motivate it properly, and teach it to maintain focus on a single track and ignore any others that might otherwise seem of greater interest to an untrained dog. An intensive search for a scent, for instance searching a ship for contraband, can actually be very fatiguing for a dog, and the dog must be motivated to continue this hard work for a long period of time.
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What information a dog actually detects when it is scenting is not perfectly understood. Although once a matter of debate, it now seems to be well established that dogs can distinguish two different types of scents when trailing: An air scent from some person or thing that has recently passed by, as well as a ground scent that remains detectable for a much longer period. The characteristics and behavior of these two types of scent trail would seem, after some thought, to be quite different, the air scent being intermittent but perhaps less obscured by competing scents, whereas the ground scent would be relatively permanent with respect to careful and repetitive search by the dog, but would seem to be much more contaminated with other scents. In any event, it is established by those who train tracking dogs that it is impossible to teach the dog how to track any better than it does naturally; the object instead is to motivate it properly, and teach it to maintain focus on a single track and ignore any others that might otherwise seem of greater interest to an untrained dog. An intensive search for a scent, for instance searching a ship for contraband, can actually be very fatiguing for a dog, and the dog must be motivated to continue this hard work for a long period of time.
  
===Coat color===
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===Reproduction===
Domestic dogs often display the remnants of countershading, a common natural camouflage pattern.  The general theory of countershading is that an animal that is lit from above will appear lighter on its upper half and darker on its lower half where it will usually be in its own shade. This is a pattern that predators can learn to watch for.  A countershaded animal will have dark coloring on its upper surfaces and light coloring below.  This reduces the general visibility of the animal. One reminder of this pattern is that many breeds will have the occasional "blaze", stripe, or "star" of white fur on their chest or undersides.
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[[Image:CatahoulaLitter wb.jpg|thumb|right|A Catahoula leopard dog mother nursing her litter of puppies.]]
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In domestic dogs, sexual maturity ([[puberty]]) begins to happen around age 6 to 12 months for both males and females, although this can be delayed until up to two years old for some large breeds. [[Adolescence]] for most domestic dogs is around 12 to 15 months, beyond which they are for the most part more adult than puppy. As with other domesticated species, [[domestication]] has selectively bred for higher [[libido]] and earlier and more frequent breeding cycles in dogs than in their wild ancestors.
  
===Sprint metabolism===
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Dogs bear their litters roughly 9 weeks after [[fertilization]], although the length of gestation can vary from 56 to 72 days. A general rule of thumb is that a [[mammal]] will produce half as many offspring as the number of teats on the mother. This rule is altered in domesticated animals since larger litters are often favored for economic reasons and in dogs, particularly, the great range of sizes and shapes plays a role in how many healthy puppies a female can carry. An average litter consists of about six puppies, though this number may vary widely based on the breed of dog. Small dogs generally produce from one to four puppies in each litter, while much larger breeds may average as many as 12 pups in each litter. The number of puppies also varies with the mother's age and health, the father's [[sperm]] count, the timing of the breeding, and many other factors.
  
Dogs can generate large amounts of energy for a short period of time. This setup is useful for hunting and combat. An example of this is that almost any dog, even one that is usually slow and lazy, can outsprint any human. However, almost any human can catch up with any dog in a marathon run, or other activity where endurance is needed.
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===Lifespan===
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The typical lifespan of dogs varies considerably by breed. For example, many giant dog breeds (such as great danes) average only 7 or 8 years, while some small terrier breeds, or toy breeds, might live to be older than 20. The average lifespan for mixed-breed and midsize dogs is about 13 to 14 years. The longest-lived dog with reliable documentation died at 29 years and 27 days; the breed of the dog was an Australian cattle dog and it lived in Virginia, [[United States]], with the date of death in 1939. Although the lifespans of all living species are mostly uncontrollable, one can significantly extend a dog's life by feeding it the right kinds of foods, giving it regular exercise, treating its [[disease]]s, caring for its special needs, and giving it love and comfort.
  
A dog's heart and lungs are oversized relative to its body and its normal everyday needs.  A dog also has relatively more red blood cells than a human. Most of the time the dog will keep the extra red blood cells stored in its spleen.  When the animal enters into a situation where its full metabolism is required, such as play, catching game, or fighting other dogs, the extra cells are released into the bloodstream. The "oversized" heart and lungs will now be running at full capacity, and the animal will have an enhanced ability to engage in aerobic activity.  This activity will produce internal heating.  Dogs, being covered in fur, are limited in their ability to cool down.  After a short time the animal must either cease its athletic activity or risk harming itself from overheating.  One can easily observe this pattern of intense activity followed by rest periods in puppies.  During the rest phase the spleen collects red blood cells and the animal may pant to cool down.
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===Social behavior===
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Dogs, like almost all canids, are social animals. [[Wolf|Wolves]], and it is assumed the ancestors of dogs, almost always live in pairs, a male and a female with their young, or in larger groups called [[pack]]s. Within the pack, the individual animals communicate with each other by visual signals, such as body and facial expressions and tail wagging, as well as by vocal signals, such as growls and howls. Pack members sometimes hunt individually and sometimes as a group, depending on the situation, including the size of the prey (Kruuk 2002).
  
==Working, utility and assistance dogs==
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The members of the pack cooperate in many activities, such as hunting, caring for the young, and defending their territory from intruders. Within each pack, there is a dominant pair which takes the lead, while the others support them. In most cases, only the dominant pair reproduces. Wolves struggle to establish dominance and sometimes fight with each other, but this seldom leads to serious injury or death since the losing animal will display submissive behavior which discourages further attacks.  Studies of feral dogs have found that they often form packs of 3 to 5 individuals (Clutton-Brook 1999; Nowak 1993; Voelker 1986).
[[Image:Labrador Retriever assistance dog.jpg|thumb|right|200px|[[Labrador Retrievers]] are often used as assistance dogs.]]
 
{{main|Working dog}}
 
  
There are several types of working dogs:
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== Origins ==
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According to recent [[Genetics|genetic]] research, the lineage of dogs separated from that of wolves about 100,000 years ago (Wayne 1997). It is not known when the ancestors of dogs began to associate with humans. The first [[fossil]] evidence of domesticated dogs, which had changed in physical form from their wolf ancestors, dates from around 15,000 years ago. Dog fossils from about this time have been found in [[Asia]], [[Europe]], [[North America]], and [[South America]] (Clutton-Brock 1999).
  
*[[Assistance dog]]s which help the seeing and hearing-impaired. Others are trained to help those with epilepsy and psychiatric disorders, by detecting the onset of the condition so they can seek help. The typical assistance dog is a sociable breed such as a [[Labrador Retriever]].
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Other genetic research seems to indicate that [[China]] was the site of the original domestication of dogs (McGourty 2002). Genetic research has identified 14 ancient dog breeds, with the oldest being the Chow Chow, Shar Pei, Akita Inu, Shiba Inu, and Basenji. Many of the 14 breeds are associated with China and Japan (Savolainen 2002).
*The [[detection dog]], which is a dog trained to and works at using its senses (almost always the sense of smell) to detect substances such as explosives or [[illegal drugs]].
 
*[[Guard dog]]s trained to protect property. These are commonly [[Doberman pinscher|Doberman Pinschers]], [[German shepherd|German Shepherds]] and [[Rottweiler]]s or mixes of these breeds.
 
*[[Herding dogs]] trained to protect livestock in the range against predators. [[sheep dog|Sheepdogs]] are a common breed.
 
*[[Hunting dog]]s that search for and retrieve game. Pointers and retrievers are typical hunting breeds.
 
*[[Police dogs]], typically German Shepherds, are trained to assist law enforcement officers.
 
*[[Rescue dogs]] that search for missing humans. The archetypal breeds are the [[St. Bernard (dog)|St. Bernard]] and the [[Bloodhound]], although any dog with a good sense of smell can be trained to perform this task.
 
*[[Therapy dog]]s, with friendly and gentle temperaments, trained to provide comfort and affection to hospitalized and institutionalized patients.
 
*The [[war dog]], used by the military to detect mines and enemy soldiers.
 
  
==Show and sport (competition) dogs==
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== Human relationships ==
[[Image:English-Springer-Spaniel.jpg|thumb|An [[English Springer Spaniel]] during a show in a country where [[Docking (animals)|docking]] is illegal]]
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Dogs are highly [[social animal]]s. This can account for their trainability, playfulness, and ability to fit into human households and social situations. This similarity has earned dogs a unique position in the realm of interspecies relationships.
{{main|Show dog}}
 
{{seealso|Dog sport}}
 
  
Owners of dogs often enter them in competitions, whether show (breed conformation shows) or sports, including dog racing & dog sledding. The winners garner much prestige and prize money. These dogs are often bred specifically for competition, which may not be entirely beneficial for the breeds due to the magnification of hereditary defects.
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The loyalty and devotion that dogs demonstrate as part of their natural instincts as pack animals closely mimics the human ideas of [[love]] and [[friendship]], leading many dog owners to view their pets as full-fledged family members. Conversely, dogs seem to view their human companions as members of their pack, and make few, if any, distinctions between their owners and fellow dogs.
  
*[[Dog agility]] is a sport in which dogs complete a timed obstacle course.
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Dogs fill a variety of roles in human society and are often trained as working dogs. For dogs that do not have traditional jobs, a wide range of dog sports provide the opportunity to exhibit their natural skills. In many countries, the most common and perhaps most important role of dogs is as companions or pets. In the United States, about 36 percent of households own at least one dog, with the total number of animals about 62 million (AVMA 2002).  
*[[Dogsled racing]] is a winter sport where a team of dogs, usually high performance mixed breed dogs called [[alaskan husky|Alaskan Huskies]] or [[Eurohound]]s, pull a sled and driver (called a [[mushing|musher]]). The [[Iditarod Trail Sled Dog Race]] is perhaps the most famous of these races. [[Dog sled]]ding is an ancient form of transportation and still a very effective way of moving freight across this type of terrain.
 
*[[Dog racing]], almost always [[Greyhounds]], involves dogs racing at betting tracks in a sport not unlike horseracing, reaching speeds of 40 miles per hour. Elsewhere, [[Dachshunds]] are often raced, as a humorous sidelight in charity events.
 
*[[Dog fighting]] and [[Bait (dogs)|dog baiting]] are [[blood sports]] involving dogs. They are illegal in most jurisdictions, but are still occasionally performed underground.
 
*The [[show dog]], purebreds entered in conformation dog shows and evaluated by how closely they match the ideal characteristics of the breed. The [[Westminster Kennel Club Dog Show]] is one of the most famous of this type of competition.
 
  
==Diet==
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Dogs have lived and worked with humans in so many roles that their loyalty has earned them the unique sobriquet, "man's best friend."
  
At present, there is some debate as to whether domestic dogs should be classified as [[omnivores]] or [[carnivores]], by diet.
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==Terminology==
The classification in the Order [[Carnivora]] does not necessarily mean that a dog's diet must be restricted to [[meat]]; unlike an obligate carnivore, such as the [[cat]] family with its shorter [[small intestine]], a dog is dependent on neither meat-specific protein nor a very high level of protein in order to fulfill its basic dietary requirements. Dogs are able to healthily digest a variety of foods including [[vegetable]]s and [[cereal|grains]], and in fact dogs can consume a large proportion of these in their diet. Wild canines not only eat available plants to obtain [[essential amino acid]]s, but also obtain nutrients from vegetable matter from the stomach and intestinal contents of their [[herbivorous]] prey, which they usually consume. Domestic dogs can survive healthily on a reasonable and carefully designed [[vegetarian]] diet, particularly if [[egg (food)|eggs]] and [[milk]] products are included. Some sources suggest that a dog fed on a [[strict vegetarian]] diet may develop [[dilated cardiomyopathy]] since it lacks [[L-carnitine]],<ref>''Small animal internal medicine'', RW Nelson, Couto page 107</ref> however, maintaining a balanced diet is also a factor since [[L-carnitine]] is found naturally in many [[nut (fruit)|nuts]], [[seed]]s, [[bean]]s, [[vegetable]]s, [[fruit]]s and [[whole grain]]s. In the wild, dogs can survive on a vegetarian diet when animal prey is not available. However it has been noted, both by observation of extremely stressful conditions such as the Iditarod Trail Sled Dog Race and by scientific studies of similar conditions, that high-protein (approximately 40%) diets including meat help prevent damage to [[muscle]] tissue. (This research is also true for some other mammals.) This level of protein corresponds to the percentage of protein found in the wild dog's diet when prey is abundant; higher levels of protein seem to confer no added benefit.
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The [[English language|English]] word "dog," in common usage, refers to the domestic [[pet]] dog, ''Canis lupus familiaris.'' The species was originally classified as ''Canis familiaris'' by Linnaeus in 1758. In 1993, dogs were reclassified as a subspecies of the gray wolf, ''Canis lupus,'' by the [[Smithsonian Institution]] and the American Society of Mammalogists.  
  
Dogs frequently eat [[grass]], which is a harmless activity. Explanations abound, but rationales such as that it neutralizes [[acid]], or that eating grass might make the dog [[vomit]], so dogs eat grass to remove unwanted substances from their [[stomach]]s, are at best educated guesses. Indeed, dogs may eat grass when they have an upset stomach in order to cause vomiting. The grass binds with the offending material. Then the grass acts as an irritant and causes the dog to vomit, bringing up the poisonous material at the same time..<ref> [http://www.wonderquest.com/DogsGrass.htm Why dogs eat grass.]</ref> Dogs do vomit more readily than humans, as part of their typical feeding behavior of gulping down food then regurgitating indigestible bones, fur, ''etc.'' This behavior is typical of pack feeding in the wild, where the most important thing is to get as much of the kill as possible before others consume it all. Individual domestic dogs, however, may be very "picky" eaters, in the absence of this evolutionary pressure. <br />
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The English word "dog" might derive from the Old English "''docga,''" a "powerful breed of canine," like the modern mastiffThe English word "hound" is a cognate of German ''Hund,'' Dutch ''hond,'' common Scandinavian ''hund,'' Icelandic ''hundur,'' which, though referring to a specific breed in English, means "dog" in general in the other Germanic languages. "Hound" itself derives from the [[Proto-Indo-European language|Proto-Indo-European]] *kwon-, which is the direct root of the Greek κυων (kuōn) and the indirect root of the Latin [[canis]] through the variant form *kani-.
  
Also, dogs eat grass, because it's a normal part of their diet. Wild canine relatives of dogs — wolves and foxes — eat all of an animal when they catch it. They end up, therefore, eating grasses and plants in the intestines of these animals. Domestic dogs, eating prepared dog food, lack this part of their diet and so they eat grass. Moreover, they just like it.  A veterinarian remembers a beagle that picks raspberries faster than his owner. Finally, dogs may ''appear'' to eat grass, when they are just running the blades through their mouth to gather information. Their sense of smell and taste may act together to detect if other animals have walked through their area or urinated on the grass.<ref> [http://www.wonderquest.com/DogsGrass.htm Why dogs eat grass.]</ref>
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In breeding circles, a male canine is referred to as a "dog," while a female canine is called a "bitch." The father of a litter is called the "sire," and the mother of a litter is called the "dam." Offspring are generally called "pups" or "puppies" until they are about a year old. A group of offspring is a "litter." The process of birth is "whelping." Many terms are used for dogs that are not purebred.
  
===Dangerous substances===
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The [[constellation]]s Canes Venatici, Canis Major, and Canis Minor are named from the [[Latin language|Latin]] word for "dog," for their perceived resemblance to dogs. [[Sirius]], the brightest [[star]] in the sky as seen from earth, is called the "dog star" because it is found in Canis Major. The time when Sirius rises just before sunrise, mainly July and August, are called the "dog days of summer."
  
Some foods commonly enjoyed by humans are dangerous to dogs, including [[chocolate]] ([[Theobromine poisoning]]), [[onion]]s, [[grape]]s and [[raisin]]s,<ref>{{cite web | url = http://www.aspca.org/site/News2?page=NewsArticle&id=16645 | title = ASPCA Animal Poison Control Center Issues Nationwide Update: Raisins and Grapes Can Be Toxic To Dogs | accessmonthday = September 2 | accessyear = 2006 | date = [[2004-07-06]] | work = ASPCA Press Releases | publisher = American Society for the Prevention of Cruelty to Animals }}</ref> some types of [[chewing gum|gum]], certain sweeteners<ref>{{cite web|url=http://today.reuters.co.uk/news/articlenews.aspx?type=topNews&storyID=2006-09-29T225449Z_01_N29420115_RTRUKOC_0_UK-LIFE-DOGS.xml|title=Dog owners warned over sugar-free items|publisher=Reuters}}</ref> and [[Macadamia nut]]s. It is currently believed that the only dangerous substance in chocolate is [[cocoa]], meaning that forms of chocolate without this ingredient, such as [[white chocolate]] may become re-evaluated for their safety in future.
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==Breeds==
  
The [[Grape and raisin toxicity in dogs|acute danger]] from grapes and raisins has been uncovered only since about 2000, and made public slowly since then. At present the cause is not known. Whatever the reason, since only small quantities are necessary to induce acute [[renal failure]], dogs should not be fed grapes or raisins, and [[Sultana (grape)|sultanas]] and [[currants]] should likely be withheld as well.
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Different breeds of dogs emerged over time due to environmental factors and human selection.  In ancient [[Egypt]], dogs looking like the modern greyhound and the mastiff are pictured in art. In ancient [[Roman]] writings, miniature pet dogs like modern toy breeds are mentioned (Clutton-Brook 1999).
  
Cooked [[bone]]s should '''never''' be given to dogs, as the heat changes the chemical and physical properties so that they cannot be chewed properly, splintering into jagged shards, and resist digestion.
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There are said to be over 450 different dog breeds in the world. As of 2006, the American Kennel Club recognized 153 breeds, divided into 7 groups (AKC 2006):
  
Human medications should not be given to a dog as a substitute for their regular medication as some can be especially [[toxic]], especially [[paracetamol]]/acetaminophen (Tylenol).
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* '''Sporting breeds''' were originally bred and trained to assist gun-wielding hunters in finding and recovering [[bird]]s and other small game. They include various pointers, spaniels, and retrievers.
[[Alcoholic beverage]]s pose much of the same hazards to dogs as they do to humans.
 
  
Dogs may also find some poisons attractive, including [[antifreeze]], [[snail]] bait, [[slug]] bait, [[insect]] bait, and [[rodent]] poisons. [[Antifreeze]] may be one of the most insidious of poisons to dogs because of its sweet taste and because a dog may walk upon or lie down upon a spill of it or its residue and then lick it off. Dogs must be kept strictly away from antifreeze and not allowed access to any place that has had a spill of it that has not been completely removed.  
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* '''Hounds''' were bred to track and engage larger hunted animals. They include the bloodhound, the English foxhound, and the beagle.
  
Plants such as [[caladium]], [[dieffenbachia]] and [[philodendron]] will cause [[throat]] irritations that will burn the throat going down as well as coming up. [[Hops]] are particularly dangerous and even small quantities can lead to [[malignant hyperthermia]].<ref>{{cite journal | first = K. L.| last = Duncan | coauthors = W. R. Hare and W. B. Buck | date = [[1997-01-01]] | title = Malignant hyperthermia-like reaction secondary to ingestion of hops in five dogs | journal = Journal of the American Veterinary Medical Association | volume = 210 | issue = 1 | pages = 51-4 | id = {{PMID|8977648}}}}</ref>
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* '''Working breeds''' were bred for various tasks; such as guarding people, livestock, and property; pulling sleds; and rescuing people from water or snow. They include the rotweiler, the saint bernard, and the akita.
  
[[Amaryllis]], [[daffodil]], [[english ivy]], [[Iris (plant)|iris]], and [[tulip]] (especially the bulbs) cause [[gastric]] irritation and sometimes [[central nervous system]] excitement followed by [[coma]], and, in severe cases, even death.
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* '''Terriers''' were originally bred to catch and kill small animals, especially [[rat]]s, without the help of a human hunter. Popular terriers include the airedale terrier and the Parson Russell terrier (also called the Jack Russell terrier).
  
Ingesting [[foxglove]], [[lily of the valley]], [[Ranunculaceae|larkspur]] and [[oleander]] can be life threatening because the [[cardiovascular]] system is affected. Equally life threatening is the [[yew]] which affects the nervous system. If any of these plants are ingested, get the dog to a veterinarian immediately.
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* '''Toy Breeds''' are smaller dogs bred to be pets. They include the pug, the pekingese, and the chihuahua.
  
Many household cleaners such as [[ammonia]], [[bleach]], [[disinfectants]], [[drain cleaner]], [[soap]]s, [[detergent]]s, and other cleaners, [[mothball]]s and [[matches]] are dangerous to dogs, as are cosmetics such as [[deodorant]]s, [[hair coloring]], [[nail polish]] and remover, [[permanent wave|home permanent]] lotion, and [[suntan lotion]].
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* '''Non-sporting breeds''' are medium-sized or large dogs bred to be pets, or in the case of the dalmation, a companion to [[horse]]s. The poodle is one of the most popular non-sporting breeds.
  
[[Zinc]] toxicity, mostly in the form of the ingestion of [[Cent (United States coin)|US pennies]] minted after 1982, is commonly fatal in dogs where it causes a severe [[hemolytic anemia]].<ref>Stowe CM, Nelson R, Werdin R, et al: Zinc phosphide poisoning in dogs. JAVMA 173:270, 1978</ref>
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* '''Herding breeds''' were bred to herd [[sheep]] or [[cattle]]. They include the collie, the German shepherd, and the Old English sheepdog.
  
Dogs will occasionally eat their own feces or the feces of other species if available, such as that of cats and horses. This is known as [[coprophagia]], and may be indicative of boredom or hunger, although there is no definitive reason known. Owners of both cats and dogs may find that dogs treat the catbox as a snack bar. Such behavior should be discouraged, as it could lead to [[Toxoplasmosis]]. Dogs seem to have different preferences in relation to eating feces. Some are attracted to the stools of deer, cows, or horses. Others will eat the stools of other dogs. And a great many dogs are attracted to cat droppings, possibly because cat foods are very high in protein and the dogs are going after undigested nutrients.<ref> [http://www.petlibrary.co.uk/dog-care/why-do-dogs-eact-poop.html Why do dogs eat poop.]</ref>
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Through history different dog breeds have gone in and out of fashion. In England, the King Charles Spaniel fell out of favor in 1689 when the house of Stuart was overthrown in the [[Glorious Revolution]] and was reborn in the twentieth century as the Cavalier King Charles Spaniel. In the United States, the beagle became very popular in the 1960s because of the cartoon character [[Snoopy]]. The most popular breed in the United States today is the Labrador retriever (AKC 2006).
  
==Reproduction==
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==Mixed breed dogs==
{{main|Canine reproduction}}
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[[Image:Working dog edit.jpg|thumb|left|Mixed-breed dogs, such as this Australian Border Collie cross, are often desired as working animals for their robust health and intelligence.]]
===General===
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The majority of dogs in the world are of mixed breed or no definite breed. It is estimated that 60 percent of dogs in the [[United States]] are mixed breeds, and the number it thought to be much higher in poorer countries. Advocates of mixed breed dogs as working dogs and pets say that they are more intelligent and have fewer health problems than purebreds (Kilcommons 1996).
In domestic dogs, sexual maturity ([[puberty]]) begins to happen around age 6 to 12 months for both males and females, although this can be delayed until up to two years old for some large breeds. [[Adolescence]] for most domestic dogs is around 12 to 15 months, beyond which they are for the most part more adult than puppy. As with other domesticated species, [[domestication]] has selectively bred for higher [[libido]] and earlier and more frequent breeding cycles in dogs, than in their wild ancestors.
 
  
===Males===
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==Working, utility, and assistance dogs==
{{sectstub}}
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[[Image:Labrador Retriever assistance dog.jpg|thumb|Labrador retrievers are often used as assistance dogs.]]
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[[Image:Belgian Shepherd Malinois on top of tank.jpg|thumb|Belgian Malinois war dog.]]
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The first ways that dogs were useful to humans were as helpers in hunting and as protectors.  These roles grew out of the dogs' natural instincts. As time went on, dogs were trained to perform other tasks and were specially breed to do so. Some of the modern types of working dogs are:
  
===Females===
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* '''Assistance dogs''' help the seeing and hearing-impaired. Others are trained to help those with [[epilepsy]] and psychiatric disorders, by detecting the onset of the condition so they can seek help. The typical assistance dog is a sociable breed such as a Labrador retriever. In the United States, people with assistance dogs are guaranteed access to public places by law.
Females have a twice-yearly fertile period (technically every 7 months in most breeds), known as 'seasons', during which her body prepares for [[pregnancy]], and at the peak she will come "into heat", her fertile period, during which time she will be mentally and physically receptive to [[copulation]]. A female is able to become pregnant on her first heat; this is not normally considered advisable as she is mentally and physically still young in other ways.  
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* '''Detection dogs'''  are trained to, and work at, using their senses (almost always the sense of smell) to detect substances such as explosives or [[illegal drugs]].
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* '''Guard dogs''' are trained to protect people and property. These are commonly doberman pinschers, German shepherds, and rottweilers, or mixes of these breeds. Guard dogs differ from '''watch dogs''' in that they are expected to engage an intruder, while watch dogs are only expected to bark to alert the property owner.
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* '''Herding dogs''' are trained to control livestock in the range. Sheepdogs and collies are common breeds.
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* '''Hunting dogs''' search for and retrieve game. Pointers and retrievers are typical hunting breeds.
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* '''Police dogs,''' typically German shepherds, are trained to assist law enforcement officers, often apprehending fleeing suspects. ''Schutzhund'' is a special test developed in [[Germany]] in the early 1900s to test if a German shepherd has the right talents and temperment to be successfully trained as a police dog.
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* '''Rescue dogs''' are those that search for missing humans. The archetypal breeds are the Saint Bernard and the bloodhound, although any dog with a good sense of smell can be trained to perform this task. One of the most famous dog rescues in history is that of [[Napoleon Bonaparte]], a poor swimmer, who fell into the ocean one night in 1814, and was rescued by a Newfoundland.
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* '''Show dogs''' do tricks for the entertainment of audiences. Poodles and American Eskimo dogs were very popular as show dogs in [[circus]]es and traveling shows in the past.  Now many types of dog perform in [[Motion picture|movies]] and [[television]].
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* '''Therapy dogs,''' with friendly and gentle temperaments, are trained to provide comfort and affection to hospitalized and institutionalized patients. The modern concept of the therapy dog was begun in 1976 by Elaine Smith, an American [[nurse]].
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* '''War dogs''' are used by the military to detect mines and enemy soldiers. War dogs, looking much like modern mastiffs, are pictured in ancient [[Assyria|Assyrian]] art. Dogs continued to be used in war through Roman and Medieval times. In 1560, Queen [[Elizabeth I]] of England sent 8,000 war dogs, equipped with spiked collars and coats of mail, with her army sent to [[Ireland]] to suppress a rebellion (Kilcommons 1996). In [[World War II]], the Soviet army trained dogs to carry explosives and run under enemy tanks, blowing them up.
  
As with most domesticated species, one of the first and strongest effects seen from selective breeding is selection for cooperation with the breeding process as directed by humans. In domestic dogs, one of the behaviors that is noted is the abolition of the pair bond seen in wild canines. The ability of female domestic dog to come into [[estrus]] at any time of the year and usually twice a year is also valued. The amount of time between cycles varies greatly among different dogs, but a particular dog's cycle tends to be consistent through her life. This is also called ''in season'' or ''in heat''. Conversely, undomesticated canine species experience estrus once a year, typically in late winter.
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==Show and sport (competition) dogs==
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[[Image:English-Springer-Spaniel.jpg|thumb|An English Springer spaniel in a dog show]]
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[[Image:BC_eye.jpg|thumb|The border collie uses a direct stare at sheep, known as "the eye," to intimidate while herding at a trial.]]
  
Most female dogs come into season for the first time between 6 and 12 months, although some larger breeds delay until as late as 2 years. Like most mammals, the age that a bitch first comes into season is mostly a function of her current body weight as a proportion of her body weight when fully mature. The different rates of maturation are responsible for the [[menarche]], not the chronological age. They then experience fertile seasons biannually until old age. Female dogs do not experience menopause, although their cycles will become irregular and fertility becomes unpredictable as they become older. Dogs over around 7 or 8 years are usually considered no longer appropriate for breeding, but can still remain fertile.
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Owners of dogs often enter them in competitions, whether show (breed conformation shows) or sports, including dog racing and dog sledding. The winners garner both prestige and prize money. These dogs are often bred specifically for competition, which may not be entirely beneficial for the breeds, due to the magnification of hereditary defects.
  
===Copulation===
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* '''Dog agility''' is a sport in which dogs complete a timed obstacle course.  It is a fairly new sport, started in 1978 in the [[United Kingdom]].
{{Unreferencedsect|date=January 2007}}
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* '''Tracking trials'''  encourage dogs to make use of their strongest facility by emulating the finding of a lost person or article in a situation where the performance of the dog can be fairly assessed. Because of this, the tracks laid are not the wanderings, which may characterize a lost person, nor do they include deliberate attempts by the tracklayer to deceive the following dog.
When [[copulating]], a male canine initially mounts the female from behind, as with most [[tetrapod]]s, a position known informally as [[doggy style]]. The female will hold her tail to the side and allow this if receptive. If unreceptive she may sit or lie down, snap, move away, or otherwise be uncooperative or not allow mating. The male will often move about as he tries to get a good purchase upon her, and whilst attempting [[sexual penetration|penetration]] of his [[penis]] to the female's [[vulva]]. At the time of penetration, the penis is not truly [[erection|erect]].  Rather, it is held rigid by a small bone, known as the [[baculum]].  
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*'''Dogsled racing''' is a winter sport where a team of dogs pull a sled and driver (called a musher). The Iditarod Trail Sled Dog Race is perhaps the most famous of these races. Dog sledding is an ancient form of transportation and still a very effective way of moving freight across this type of terrain. Most modern dog sledders use mixed breed dogs.
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* '''Dog racing,''' almost always emphasizing greyhounds, involves dogs racing at betting tracks in a sport not unlike horse racing, reaching speeds of 40 miles per hour. Elsewhere, dachshunds are often raced, as a humorous sidelight in charity events.
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*'''Dog fighting''' and '''dog baiting''' are blood sports involving dogs. In the past, they were very popular but now they are illegal in most jurisdictions, but are still occasionally performed underground. Some of the breeds which have been used in dog fighting are the akita, the Irish terrier, and the American pit bull terrier (often called the pit bull). 
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* '''Disc dog''' is the more generic name for what is commonly called  '''Frisbee dog.'''  In disc dog competitions, dogs and their human disc throwers compete in events such as distance catching and somewhat choreographed freestyle catching. The sport celebrates the bond between handler and dog, by allowing them to work together. The term "disc" is preferred because "Frisbee" is a trademark (held by Wham-O) for a brand of flying disc.
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* '''Flyball''' is an event in which teams of dogs race against each other from a start/finish line, over a line of hurdles, to a box that releases a tennis ball to be caught when the dog presses the spring loaded pad, then back to their handlers while carrying the ball. Flyball has become popular because it is a sport in which all sizes and breeds of dog can compete.  
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* '''Sheepdog trials''' (or simply '''dog trials''') are competitive events in which herding dog breeds move [[sheep]] around a field, fences, gates, or enclosures as directed by their handlers. Such events are particularly associated with hill farming areas, where sheep range widely on largely [[fence|unfenced]] land. These trials are popular in the [[United Kingdom]], [[Ireland]], [[South Africa]], [[Canada]], the [[United States]], [[Australia]], [[New Zealand]], and other farming nationsSome venues allow only dogs of known herding breeds to compete; others allow any dog that has been trained to herd.
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* '''Earthdog trials''' test dachshunds and small terriers for their working abilities and instinct. Earthdog den trials involve underground tunnels that dogs must negotiate, while scenting a rat, the "quarry." The dog must follow the scent to the quarry and then "work" the quarry. Depending on the sanctioning organization, “working” means barking, scratching, staring, pawing or such at it, although the quarry is protected at all times.
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* '''Field trials''' are highly competitive events at which hunting dogs compete against one another. Field trials are organized by kennel clubs or other gun dog organizations.
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* '''Obedience trials''' are events in which a dog must perfectly execute a predefined set of tasks when directed to do so by its handler. Obedience competition provides an opportunity for a person and a dog to work as a highly tuned team.
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* '''Dog shows''' involve purebreds entered in conformation dog show and evaluated by how closely they match the ideal characteristics of the breed. The Westminster Kennel Club Dog Show is one of the most famous of this type of competition.
  
When the male achieves penetration, he will often hold tighter and thrust faster. At this point the male's penis expands. Canine reproduction is different from human [[sexual intercourse]], because human males acquire an erection prior to penetration. Canine males enter first, then swell and become erect.  
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==Dogs in research==
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Dogs have long been used as subjects of [[scientific research]]. In the 1890s, the Russian scientist [[Ivan Pavlov]] used dogs to study psychological conditioning. Much of the research which led to the discovery of [[insulin]] and to the successful treatment of [[diabetes]] in humans was carried out using dogs as experimental subjects. On November 3, 1957, a Russian dog, Laika, became the first animal to orbit the earth.
  
The male dog has a [[bulbus glandis]], a spherical area of [[erectile tissue]] at the base of the [[penis]], which traps the penis inside the female's [[vagina]] during copulation as it becomes engorged with blood. [http://arbl.cvmbs.colostate.edu/hbooks/pathphys/reprod/semeneval/dog.html] 
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Dogs presently make up only a small percentage of animals used in research; [[rat]]s and [[Mouse|mice]] far outnumber them. However, they are especially important in medical studies, for instance on the effects of drugs.  Among the reasons for this are their fairly large size and their ease in being taken care of and controlled.
  
Once the penis is locked into the vagina by the bulbus glandis, the male will usually lift a leg and swing it over the female's back while turning around. The two stand with their hind ends touching and the penis locked inside the vagina while [[ejaculation]] occurs, decreasing leakage of [[semen]] from the vagina. After some time, typically 5 - 20 minutes (but sometimes longer), the bulbus glandis disengorges, allowing the mates to separate. [[Virginity|Virgin]] dogs can become quite distressed at finding themselves unable to separate during their first copulation, and may try to pull away or run. Dog breeders' often suggest it is appropriate to help calm and settle the mating dogs once this stage is reached, if they show anxiety.
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Beagles are especially popular for research because of their calm and friendly nature. Most research beagles are especially bred for this purpose. Other research dogs are obtained from various sources including pound seizure, newspaper ads, strays, and lost or stolen pets. Over the years there have been protests against the treatment of research dogs and laws have been passed to protect them, and in some cases to prevent former pet dogs from being used in research (HSUS 2007).
  
Note that similar canine mounting behavior (sometimes with pelvic thrusting) is also used by [[dominance (biology)|dominant]] canines of both sexes. Dominance mounting, with or without thrusting, should not be confused with copulatory mounting, in which the thrusting is short term until a "tie" is achieved.
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==Dog health==
 
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Dogs are susceptible to various [[disease]]s, ailments, and poisons, some of which effect humans in the same way, others of which are unique to dogs.  
===Gestation and litters===
 
[[Image:CatahoulaLitter wb.jpg|thumb|right|A [[Catahoula Leopard Dog|Catahoula Leopard]] mother nursing her litter of puppies.]]
 
Dogs bear their litters roughly 9 weeks after [[fertilization]], although the length of gestation can vary from 56 to 72 days. A general rule of thumb is that a mammal will produce half as many offspring as the number of teats on the mother. This rule is altered in domesticated animals since larger litters are often favored for economic reasons and in dogs, particularly, the great range of sizes and shapes plays a role in how many healthy puppies a female can carry. An average litter consists of about six '''[[puppy|puppies]]''', though this number may vary widely based on the breed of dog. [[Toy dog]]s generally produce from one to four puppies in each litter, while much larger breeds may average as many as 12 pups in each litter. The number of puppies also varies with the mother's age and health, the father's sperm count, the timing of the breeding, and many other factors.
 
  
Some breeds have been developed to emphasize certain physical traits beyond the point at which they can safely bear litters on their own. For example, the [[Bulldog]] often requires [[artificial insemination]] and almost always requires [[cesarean section]] for giving birth. Since a mother can provide nutrients and care to only a limited number of offspring, humans must assist in the care and feeding when the litter exceeds approximately eight puppies.
+
===Diet===
 +
At present, there is some debate as to whether domestic dogs should be classified as [[omnivore]]s or [[carnivore]]s, by diet.
  
==Spaying and neutering==
+
The classification in the Order [[Carnivora]] does not necessarily mean that a dog's diet must be restricted to [[meat]]; unlike an obligate carnivore, such as the [[cat]] family with its shorter [[small intestine]], a dog is dependent on neither meat-specific [[protein]] nor a very high level of protein in order to fulfill its basic dietary requirements. Dogs are able to healthily digest a variety of foods including [[vegetable]]s and [[cereal|grains]], and in fact dogs can consume a large proportion of these in their diet. Wild canines not only eat available plants to obtain [[amino acid#Essential amino acids|essential amino acid]]s, but also obtain nutrients from vegetable matter from the stomach and intestinal contents of their herbivorous prey, which they usually consume.
{{detail|Spaying and neutering}}
 
Spaying (females only) and neutering (both genders but more usually males) refers to the [[sterilization]] of animals, usually by removal of the male's [[testicle]]s or the female's [[ovary|ovaries]] and [[uterus]], in order to eliminate the ability to procreate, and reduce sex drive. Neutering has also been known to reduce aggression in male dogs, but can occasionally increase aggression in female dogs.<ref>{{cite journal |author=Heidenberger E, Unshelm J |title=[Changes in the behavior of dogs after castration] |language=German |journal=Tierärztliche Praxis |volume=18 |issue=1 |pages=69-75 |year=1990 |pmid=2326799 |doi=}}</ref>
 
  
Animal control agencies in the United States and the [[ASPCA]] advise that dogs not intended for further breeding should be spayed or neutered so that they do not have undesired puppies.<ref>{{cite web
+
Domestic dogs can survive healthily on a reasonable and carefully designed [[vegetarian]] diet, particularly if [[egg (food)|eggs]] and [[milk]] products are included. Some sources suggest that a dog fed on a strict vegetarian diet may develop dilated cardiomyopathy since it lacks L-[[carnitine]] (Nelson 2003), however, maintaining a balanced diet is also a factor since L-carnitine is found naturally in many nuts, [[seed]]s, [[bean]]s, [[vegetable]]s, [[fruit]]s, and [[whole grain]]s. In the wild, dogs can survive on a vegetarian diet when animal prey is not available. However it has been noted, both by observation of extremely stressful conditions such as the Iditarod Trail Sled Dog Race and by scientific studies of similar conditions, that high-protein (approximately 40 percent) diets including meat help prevent damage to [[muscle]] tissue. (This research is also true for some other mammals.) This level of protein corresponds to the percentage of protein found in the wild dog's diet when prey is abundant; higher levels of protein seem to confer no added benefit.
|url=http://www.aspca.org/site/PageServer?pagename=adopt_spayneuter
 
|title=Top 10 reasons to spay/neuter your pet
 
|publisher=American Society for Prevention of Cruelty to Animals
 
|accessdate=2007-05-16}}</ref> In some countries of Europe, spaying or neutering of dogs is less common, and owners are usually instead advised to keep their dogs on leash and under supervision to avoid unwanted matings; drugs are used to prevent pregnancy and abort unwanted litters. {{Fact|date=January 2007}}
 
  
Because of the overpopulation of dogs in some countries, puppies born to strays or as the result of accidental breedings often end up being killed in animal shelters. Spaying and neutering can also decrease the risk of hormone-driven diseases such as mammary cancer, as well as undesired hormone-driven behaviors.  However, certain medical problems are more likely after neutering, such as [[urinary incontinence]] in females<ref>{{cite journal |author=Arnold S |title=[Urinary incontinence in castrated bitches. Part 1: Significance, clinical aspects and etiopathogenesis] |language=German |journal=Schweiz. Arch. Tierheilkd. |volume=139 |issue=6 |pages=271-6 |year=1997 |pmid=9411733 |doi=}}</ref> and [[prostate cancer]] in males.<ref>{{cite journal |author=Johnston SD, Kamolpatana K, Root-Kustritz MV, Johnston GR |title=Prostatic disorders in the dog |journal=Anim. Reprod. Sci. |volume=60-61 |issue= |pages=405-15 |year=2000 |pmid=10844211 |doi=}}</ref>  The hormonal changes involved with sterilization are likely to somewhat change the animal's personality, however, and some object to spaying and neutering as the sterilization could be carried out without the excision of organs.
+
===Spaying and neutering===
 
+
In the United States, most pet dogs are spayed or neutered. Spaying is the surgical removal of the female's [[ovaries]] and [[uterus]]. Neutering is the removal of the male's [[testicles]]. Spayed and neutered dogs avoid some diseases such as breast and testicular [[cancer]].  Behaviors such as urinating in the house, running away, and aggression are also reduced; and unwanted litters of puppies are avoided, which could contribute to the problem of stray and feral dogs (ASPCA 2014).
It is not essential for a female dog to either experience a heat cycle or have puppies before spaying, and likewise, a male dog does not need the experience of mating before neutering. However, there is mounting evidence {{Fact|date=June 2007}} that spaying female dogs before they experience at least one heat cycle and neutering males before they reach sexual maturity predisposes them to earlier allergies, more severe immune-related disorders, increased reactions to vaccinations, and increased incidence of certain cancers (e.g. bone cancer). Behavioral problems now being attributed {{Fact|date=June 2007}} to early sterilization of both male and female dogs are perpetual puppy syndrome (e.g. submissive urination), inhibited social adjustment, increase in cognitive disorders, and increased aggression in females.
 
 
 
Female cats and dogs are seven times more likely to develop [[mammary tumor]]s if they are not spayed before their first [[estrus cycle|heat cycle]]. <ref>{{cite book|author=Morrison, Wallace B.|title=Cancer in Dogs and Cats (1st ed.)|publisher=Williams and Wilkins|year=1998|id=ISBN 0-683-06105-4}}</ref>  The high dietary estrogen content of the average commercial pet food as well as the estrogenic activity of topical pesticides {{Fact|date=June 2007}} may be contributing factors in the development of mammary cancer, especially when these exogenous sources are added to those normal estrogens produced by the body. Dog food containing [[soybean]]s or soybean fractions have been found to contain [[phytoestrogens]] in levels that could have biological effects when ingested longterm.<ref>{{cite journal |author=Cerundolo R, Court MH, Hao Q, Michel KE |title=Identification and concentration of soy phytoestrogens in commercial dog foods |journal=Am. J. Vet. Res. |volume=65 |issue=5 |pages=592-6 |year=2004 |pmid=15141878 |doi=}}</ref>
 
 
 
A female dog can become pregnant on her first heat cycle (which can take place as early as six months), and should be kept away from intact male dogs, including littermates, over the age of 4 months {{Fact|date=February 2007}}.
 
 
 
Gender-preservative surgeries such as vasectomy and tubal ligation are possible, but do not appear to be popular due to the continuation of gender-specific behaviors and disease risks.
 
 
 
==Overpopulation==
 
{{globalize|USA}}
 
According to the Humane Society of the United States, 3-4 million dogs and cats are [[euthanasia|euthanized]] each year in the United States and many more are confined to cages in shelters because there are many more animals than there are homes. Spaying or neutering dogs helps keep overpopulation down.<Ref>
 
{{cite journal
 
  | quotes = Although the cause of pet overpopulation is multifaceted, failure of owners to spay and castrate their animals is a major contributing factor.
 
| last = Mahlow
 
| first = Jane C.
 
| year = 1999
 
| title = Estimation of the proportions of dogs and cats that are surgically sterilized
 
| journal = Journal of the American Veterinary Medical Association (excerpt quoted by spayusa.org)
 
| volume = 215
 
| pages = 640-643
 
| url = http://www.spayusa.org/main_directory/02-facts_and_education/stats_surveys/javma_articles/02dogs-cats-sterilized.asp
 
| accessdate = 2006-11-30
 
}}</Ref>
 
Local humane societies, SPCA's and other animal protection organizations urge people to spay or neuter their pets and to adopt animals from shelters instead of purchasing them. Several notable public figures have spoken out against animal over population, including [[Bob Barker#Animal rights|Bob Barker]]. On his [[game show]], [[The Price is Right]], Barker stresses the issue at the end of every episode, saying: "Help control the pet population. Have your pets spayed or neutered."
 
 
 
==Dog health==
 
{{main|Dog health}}
 
{{further|Category:Dog health}}
 
 
 
Dogs are susceptible to various diseases, ailments, and poisons, some of which affect humans in the same way, others of which are unique to dogs. Dogs, like all mammals, are also susceptible to heat exhaustion when dealing with high levels of humidity and/or extreme temperatures.<ref>{{cite web|url=http://www2.dasnr.okstate.edu/index.php?option=com_content&task=view&id=257&Itemid=103 |title=Summer heat can be tough on pets |accessdate=2006-08-21 | first = Trisha | last = Gedon | date = [[2006-05-25]] | work = Division of Agricultural Sciences and Natural Resources | publisher = Oklahoma State University }}</ref>
 
  
 
===Diseases===  
 
===Diseases===  
Infectious diseases commonly associated with dogs include [[rabies]] (hydrophobia), [[canine parvovirus]], and [[canine distemper]]. Congenital diseases of dogs can include a wide range from [[hip dysplasia]] and [[luxating patella|medial patellar luxation]] to [[Epilepsy in animals|epilepsy]] and [[pulmonic stenosis]]. Canines can get just about anything a human can get (excluding many infections which are species specific) like hypothyroidism, cancer, dental disease, heart disease, etc.  
+
Infectious diseases commonly associated with dogs include [[rabies]] (hydrophobia), canine parvovirus, and canine distemper. Congenital diseases of dogs can include a wide range from hip dysplasia and medial patellar luxation to [[epilepsy]] and [[pulmonic stenosis]]. Canines can get just about anything a human can get (excluding many infections, which are species specific) like hypothyroidism, cancer, dental disease, heart disease, [[Addison's disease]], and so forth.  
  
Two serious medical conditions affecting dogs are [[pyometra]], affecting [[spay and neuter|unspayed]] females of all types and ages, and [[bloat]], which affects the larger breeds or deep chested dogs. Both of these are [[Acute (medical)|acute]] conditions, and can kill rapidly; owners of dogs which may be at risk should learn about such conditions as part of good animal care.
+
Two serious medical conditions affecting dogs are [[pyometra]], affecting unspayed females of all types and ages, and [[bloat]], which affects the larger breeds or deep chested dogs. Both of these are acute conditions, and can kill rapidly; owners of dogs which may be at risk should learn about such conditions as part of good animal care.
 
 
{{see also|Vaccination of dogs}}
 
  
 
===Parasites===  
 
===Parasites===  
Common external parasites are various species of [[flea]]s, [[tick]]s, and [[mites]]. Internal parasites include [[hookworm]]s, [[tapeworm]]s, [[roundworm]]s, and [[heartworms]]. See also [[CVBD]] (Canine Vector-Borne Diseases).
+
Common external parasites are various species of [[flea]]s, [[tick]]s, and [[mite]]s. Internal parasites include [[hookworm]]s, [[tapeworm]]s, [[roundworm]]s, and [[heartworms]].  
  
 
===Common physical disorders===  
 
===Common physical disorders===  
Some breeds of dogs are also prone to certain genetic ailments, such as [[hip dysplasia]], [[luxating patella]]s, [[cleft palate]], [[blindness]], or [[hearing impairment|deafness]]. Dogs are also susceptible to the same ailments that humans are, including [[Diabetes in cats and dogs|diabetes]], [[epilepsy]], [[cancer]], and [[arthritis]]. [[Gastric torsion]] and [[bloat]] is a dangerous problem in some large-chested breeds.
+
Some breeds of dogs are also prone to certain genetic ailments, such as [[hip dysplasia]], [[luxating patella]]s, [[cleft palate]], [[blindness]], or [[hearing impairment|deafness]]. As noted, dogs are also susceptible to the same ailments that humans are, including [[diabetes]], [[epilepsy]], [[cancer]], and [[arthritis]].
 
 
===Lifespan and old age===
 
{{Main articles|Dog years|Aging in dogs}}
 
The typical lifespan of dogs varies considerably by breed. For example, many giant dog breeds (such as [[Great Danes]]) average only 7 or 8 years, while some small terrier breeds, or [[toy dog|toy breeds]] might live as long as 20 or above. The average lifespan for mixed-breed and midsize dogs is about 13 to 14 years. The longest-lived dog with reliable documentation died at 29 years and 27 days, the breed of the dog was an [[Australian cattle dog]] and it lived in [[Virginia]], [[USA]], the date of death was in 1939. The oldest currently living dog is from [[Canberra, Australia]] and is now 27 years old. Although the lifespans of all living species are mostly uncontrollable, one can significantly extend a dog's life by feeding it the right kinds of foods, giving it regular exercise, treating its diseases, caring for its special needs and giving it love and comfort.
 
 
 
==Behavior==
 
{{see|:Category:Dog training and behavior}}
 
 
 
Dogs are very social animals, but their personality and behavior vary with breed as well as how they are treated by their owners and others who come in contact with them. Physical abuse and starvation can produce very neurotic, dangerous dogs, and even simply failing to socialize them properly may entail maladaptive behaviors. {{Fact|date=February 2007}} It is not uncommon for dogs to [[dog attack|attack]] humans and other animals; however, this is usually because of lack of care or improper upbringing by its owner.
 
 
 
===Laughter in dogs===
 
Laughter might not be confined or unique to humans, despite Aristotle's observation that "only the human animal laughs". The differences between chimpanzee and human laughter may be the result of adaptations that have evolved to enable human speech. However, some behavioral psychologists argue that self-awareness of one's situation, or the ability to identify with somebody else's predicament, are prerequisites for laughter, so animals are not really laughing in the same way that humans do.
 
[[Image:NICO in letterbox.jpg|left|A|thumb|250px|A playful dog]]
 
The dog-laugh sounds similar to a normal pant. However by analyzing the pant using a spectrograph, this pant varies with bursts of frequencies, resulting in a laugh. When this recorded dog-laugh vocalization is played to dogs in a shelter setting, it can initiate play, promote pro-social behavior, and decrease stress levels. In a study by Simonet, Versteeg, and Storie, one hundred and twenty subject dogs residing in a mid-size county animal shelter were observed. Dogs ranging from 4 months to 10 years of age were compared with and without exposure to a dog-laugh recording. The stress behaviors measured included panting, growling, salivating, pacing, barking, cowering, lunging, play-bows, sitting, orienting and lying down. The study resulted in positive findings. Exposure to the dog laughing recording resulted in the following: significantly reduced stress behaviors, increased tail wagging and the display of a play-face when playing was initiated, and more frequent pro-social behavior such as approaching and lip licking. This research suggests exposure to dog-laugh vocalizations can calm and possibly increase shelter adoptions. ([http://www.petalk.org/LaughingDog.pdf Simonet, Versteeg, & Storie 2005]) A dog laughter sample: [http://www.petalk.org/DogLaughSpect.html  Simonet 2005].
 
 
 
==Dog communication==
 
{{main|Dog communication}}
 
 
 
==Ancestry and history of domestication==
 
{{Mergeto|Origin of the domestic dog|date=January 2007}}
 
{{main|Origin of the domestic dog}}
 
[[Image:DogMosaic wb.jpg|thumb|This ancient mosaic, likely Roman, shows a large dog with a collar hunting a lion.]]
 
[[Molecular systematics]] indicate that the domestic dog (''Canis lupus familiaris'') descends from one or more populations of wild wolves (''Canis lupus''). As reflected in the [[Binomial nomenclature|nomenclature]], dogs are descended from the wolf and are able to interbreed with wolves.
 
 
 
The relationship between human and canine has deep roots. [[Wolf]] remains have been found in association with [[Hominidae|hominid]] remains dating from 400,000 years ago. Converging archaeological and genetic evidence indicate a time of [[domestication]] in the late [[Upper Paleolithic]] close to the [[Pleistocene]]/[[Holocene]] boundary, between 17,000 and 14,000 years ago. [[Fossil]] bone morphologies and genetic analysis of current and ancient dog and wolf populations have not yet been able to conclusively determine whether all dogs descend from a single domestication event, or whether dogs were domesticated independently in more than one location. Domesticated dogs may have interbred with local populations of wild wolves on several occasions (so-called introgression).
 
 
 
The earliest dog fossils, two [[skull|crania]] from [[Russia]] and a [[mandible]] from [[Germany]], date from 13,000 to 17,000 years ago. Their likely ancestor is the large northern [[Holarctic]] wolf, ''Canis lupus lupus''. Remains of smaller dogs from [[Mesolithic]] ([[Natufian]]) cave deposits in the [[Middle East]], dated to around 12,000 years ago, have been interpreted as descendants of a lighter Southwest Asian wolf, ''Canis lupus arabs''. [[Rock art]] and skeletal remains indicate that by 14,000 years ago, dogs were present from [[North Africa]] across [[Eurasia]] to [[North America]]. Dog burials at the [[Mesolithic]] cemetery of [[Svaerdborg]] in Denmark suggest that in ancient Europe dogs were valued companions.
 
 
 
Genetic analyses have so far yielded divergent results. Vilà, Savolainen, and colleagues (1997) concluded that dogs split off from wolves between 75,000 and 135,000 years ago, while a subsequent analysis by Savolainen et al. (2002) indicated a "common origin from a single gene pool for all dog populations" between 40,000 and 15,000 years ago in [[East Asia]]. Verginelli et al. (2005), however, suggest both sets of dates must be reevaluated in light of recent findings showing that poorly calibrated molecular clocks have systematically overestimated the age of geologically recent events. On balance, and in agreement with the archaeological evidence, 15,000 years ago is the most likely time for the wolf-dog divergence.
 
 
 
Verginelli examined ancient [[DNA]] evidence from five prehistoric Italian [[canids]] carbon-dated to between 15,000 and 3,000 years old, 341 wolves from several populations worldwide, and 547 purebred dogs. Their results indicate multiple independent origins of dogs and/or of frequent interbreeding between early proto-dogs and wolves throughout a vast geographic range. The detailed history remains unexplored and until further evidence is available, the following section on wolf ancestors must be considered purely speculative.
 
 
 
===Wolf ancestors===
 
Although all wolves belong to the species ''[[Gray Wolf|Canis lupus]]'', there are (or were) many subspecies that had developed a distinctive appearance, social structure, and other traits. For example, the [[Japanese Wolf]] and the [[Eastern Timber Wolf]] possess different distinctive coloration, hunting and social structures. The origin of the dog is so ancient and so worldwide that many varieties of wolf played a part in it. It is wrong to say that dogs descended from modern wolves. They descended from ancestral wolves, and this difference must always be kept in mind. Ancestral wolves of many varieties existed all over the world. Humans are of a tropical origin, and it was there that the domestication of dogs from wolves first took place {{Fact|date=April 2007}}. It follows then, that the first wolves to be domesticated were the warm-climate, short-haired varieties that gave rise to many of our dog breeds, with the long-haired, northern wolves giving rise to our northern breeds when humans reached these latitudes.{{Fact|date=April 2007}}
 
 
 
The [[Indian Wolf]] is thought to have contributed to the development of more breeds of dogs than other subspecies. Many of today's wild dogs, such as the [[dingo]] and the [[pariah dog]]s, are descended from this wolf.  The Indian Wolf is also thought to have bred with descendants of the European wolf to create the [[Mastiff]]s and eventually leading to the development of such diverse breeds as the [[Pug]], the [[St. Bernard (dog)|Saint Bernard]], and the [[Bloodhound]]. The [[Tibetan Mastiff]] is an example of an ancient breed.
 
 
 
The [[Gray Wolf|European wolf]], in turn, may have contributed many of its attributes to the [[Spitz]] dog types, most [[terrier]]s, and many of today's [[sheepdog]]s. The [[Chinese wolf]] is probably ancestor to the [[Pekingese]] and toy [[spaniel]]s, although it is also probable that descendants of the Chinese and European wolves encountered each other over the millennia, contributing to many of the oriental [[toy dog|toy]] breeds.
 
 
 
The [[Eastern Timber Wolf]] is a direct ancestor to most, if not all, of the North American northern [[sled dog]] types. This interbreeding still occurs with dogs living in the [[Arctic]] region, where the attributes of the wolf that enable survival in a hostile environment are valued by humans. Additionally, unintentional crossbreeding occurs simply because dogs and wolves live in the same environment. The general [[reproductive isolation]] which is required to define dogs and wolves as separate species is purely a result of lack of opportunity, stemming from a general mutual unfamiliarity, suspicion, mistrust, and fear.
 
 
 
The [[phenotype|phenotypic]] characteristics that distinguish a wolf from a dog are tenuous. Wolves typically have a "brush tail" and erect ears. While some dog breeds possess one of these characteristics, they rarely possess both.
 
 
 
===Speed of domestication===
 
Current research indicates that domestication, or the attributes of a domesticated animal, can occur much more quickly,<ref>{{cite journal
 
| last = Trut
 
| first = Lyudmila N.
 
| year = 1999
 
| month = [[March]]-[[April]]
 
| title =  Early Canid Domestication: The Farm-Fox Experiment
 
| journal = American Scientist
 
| volume = 87
 
| issue = 2
 
| pages = 160
 
| doi = 10.1511/1999.2.160
 
| url = http://www.amsci.org/amsci/articles/99articles/Trut.html#26879
 
}}</ref> {{Failed verification|date=February 2007}} <!--Cited item does not support the statement to which it is attached—> than previously believed. Domestication of a wild dog may occur within one or two human generations with deliberate [[selective breeding]]. It is also now generally believed that initial domestication was through mutual desire. Wild canines who scavenged around human habitations received more food than their more skittish or fearful counterparts. Canines who attacked people or their children were likely killed or driven away, while those more friendly animals survived. Canines would have been beneficial by chasing away other [[vermin]] or [[scavengers]]. With their sharp senses, they would also be valuable as an alarm against marauding predators. The relationship is theorized to have developed in this way.
 
 
 
A 2007 [[Animal Planet]] documentary discussed a 1900-1920 domestication experiment in Russia that led to rapid unusual changes in the coats of foxes.
 
 
 
==Dog meat for human consumption== 
 
{{Main article|Dog meat}}
 
 
 
In some countries, certain dog breeds, apart from being kept as pets, are raised on farms and slaughtered for consumption. In countries where dogs are particularly popular as household pets, consumption of dogs is generally taboo and considered an abhorrent cultural practice<!-- This seems a tautology. Can anyone tell me the difference between "taboo" and something "considered an abhorrent cultural practice"?—>, but there are exceptions, such as [[Vietnam]], where dogs are popular as both pets and meat.<ref>{{cite web
 
  | last = Arthurs
 
  | first = Clare
 
  | title = Vietnam's dog meat tradition
 
  | publisher = BBC
 
  | date = [[2001-12-31]]
 
  | url = http://news.bbc.co.uk/1/hi/world/asia-pacific/1735647.stm
 
  | accessdate = 2006-10-10 }}</ref>
 
 
 
Although the [[China|Chinese]] were the first to domesticate the dog and keep them as pets, dog meat has been a source of food in China from at least the time of [[Confucius]], and possibly even before. Ancient writings from the [[Zhou Dynasty]] referred to the 'three beasts' (which were bred for food), including pig, goat, and dog. [[Mencius]], the philosopher, recommended dog as the tastiest of all meats.
 
 
 
==Dog abuse==
 
{{main|Cruelty to animals}}
 
Cruelty to dogs refers to treatment that causes unacceptable suffering or [[harm]]. What qualifies as unacceptable suffering varies among countries and cultures. Cruelty can be passive, typified by simple neglect, or active, with malicious intent.<Ref>
 
{{cite web
 
| url = http://www.pet-abuse.com/pages/animal_cruelty.php
 
| title = Animal Cruelty
 
| accessdate = 2006-11-30
 
| publisher = pet-abuse.com
 
}}</Ref>
 
 
 
===Signs of dog abuse===
 
Signs of dog abuse include:
 
* Unusually frightened, fearful or subdued
 
* Fractures
 
* [[Bruising]]
 
* Eye injuries
 
* [[Scald]]s and [[Burn (injury)|burns]]
 
* Signs of [[malnutrition]]
 
* Significant matting or other poor [[grooming]] indicators
 
* Ignored health problems
 
* Injury history incompatible with injury or owner refusing to comment on how injury occurred
 
* Owner showing lack of concern for animal’s injuries
 
* Owner exhibiting [[Fabricated or Induced Illness|Munchausen’s Syndrome by Proxy]] (MSP)
 
 
 
==Famous dogs & dog sayings==
 
{{Main article|List of famous dogs}}
 
 
 
In the long shared history of dogs and humans, there have been many [[List of famous dogs|famous]], and notorious, dogs, and many [[list of fictional dogs|fictional dogs]] have been featured in literature, cinema, and other media.
 
 
 
"I've seen a look in dogs' eyes, a quickly vanishing look of amazed contempt, and I am convinced that basically dogs think humans are nuts." - John Steinbeck<ref>
 
{{cite web
 
| url = http://www.dogquotes.com/dogquotes2.htm
 
| title = Dog Quotes
 
| accessdate = 2007-4-4
 
}}</ref>
 
  
"When a dog wags her tail and barks at the same time, how do you know which end to believe?" - Robin Williams in "Man of the Year"
+
==Environmental and health concerns==
 +
Despite the many benefits which dogs provide for humans, they also do harm.  Dogs sometimes attack and bite humans. In the United States, there are about 7,000 dog attacks per year (Voelker 1986).
  
==Dog breeds==
+
An even more serious problem is [[rabies]], a [[Virus|viral]] [[disease]] that is spread by animal bites. Rabies is always fatal unless treated. The [[World Health Organization]] estimates that about 55,000 people die of rabies each year; mostly in the rural areas of Africa and Asia, and mostly because of dog bites.  In Europe and North America, where most dogs are [[Immunization|immunized]] against rabies, wild animal bites are the most common cause. About 10 million people each year are treated for possible exposure to rabies (WHO 2006). Rabies is an especially serious problem in [[India]], which has about 25 million feral dogs (Sandeep 2002).
[[Image:Roodog2k-ivan-and-boxer.JPG|thumb|The dog has developed into hundreds of breeds with a great degree of variation, such as this [[Miniature Pinscher]] and [[Boxer (dog)|Boxer]].]]
 
There are numerous [[dog breed]]s, with [[List of dog breeds|over 800]] being recognized by various [[kennel club]]s worldwide. As all dog breeds have been derived from mixed-breed dog populations, the term "purebred" has meaning only with respect to a certain number of generations. Many dogs, especially outside the United States of America and Western Europe, belong to no recognized breed.
 
[[Image:Papagility004.jpg|left|thumbnail|200px|The [[Papillon (dog)|Papillon]] is a highly athletic breed of dog]]
 
A few basic [[:Category:Dog types|breed types]] have evolved gradually during the domesticated dog's relationship with humans over the last 10,000 or more years, but most modern breeds are of relatively recent derivation. Many of these are the product of a deliberate process of [[artificial selection]]. Because of this, some breeds are highly specialized, and there is extraordinary morphological diversity across different breeds. Despite these differences, dogs are able to distinguish dogs from other kinds of animal.
 
  
The definition of a dog breed is a matter of some controversy. Depending on the size of the original founding population, closed gene pool breeds can have problems with inbreeding, specifically due to [[founder effect]]. Dog breeders are increasingly aware of the importance of population genetics and of maintaining diverse gene pools. Health testing and new DNA tests can help avoid problems, by providing a replacement for natural selection. Without selection, inbreeding and closed gene pools can increase the risk of severe health or behavioral problems. Some organizations define a breed more loosely, such that an individual may be considered of one breed as long as 75% of its parentage is of that breed. These considerations affect both pets and the show dogs entered in [[conformation show|dog shows]]. Even prize-winning [[purebred]] dogs sometimes possess crippling [[Genetic disorder|genetic defects]] due to founder effect or [[inbreeding]].<ref>{{cite book
+
Dogs can also have a negative effect on the natural environment. In [[New Zealand]], pet dogs allowed to run wild were found to be a serious threat to [[kiwi]]s and other ground nesting birds (ISSG 2007). A study in the city of [[Baltimore]] in the United States done in the 1970s found that about half the dogs in the city were at least partly free ranging and that they also feed on ground nesting birds, although their main food source was garbage (Nowak 1983). In Africa, diseases spread by domestic dogs have contributed to the decline of endangered wild predators, such as the [[lion]], the [[leopard]], and the [[African hunting dog]] (Kruuk 2002).
  | last = Shook
 
  | first = Larry
 
  | title = The Puppy Report: How to Select a Healthy, Happy Dog
 
  | publisher = Ballantine
 
  | date = 1995
 
  | location = New York
 
  | pages = 57-72
 
  | id = ISBN 0-345-38439-3 }}</ref>
 
These problems are not limited to [[purebred]] dogs and can affect mixed-breed populations.<ref>{{cite book
 
  | last = Shook
 
  | first = Larry
 
  | title = The Puppy Report: How to Select a Healthy, Happy Dog
 
  | publisher = Ballantine
 
  | date = 1995
 
  | location = New York
 
  | pages = 13-34
 
  | id = ISBN 0-345-38439-3 }}</ref>  The behavior and appearance of a dog of a particular breed can be predicted fairly accurately, while mixed-breed dogs show a broader range of innovative appearance and behavior.
 
  
In February 2004, the Canine Studies Institute in [[Aurora, Ohio]], arranged recognized breeds of dogs into ten categories.  
+
==Dogs in culture and religion==
 +
[[Image:Hercules capturing Cerberus.jpg|thumb|''Hercules capturing Cerberus'', Engraving by Sebald Beham 1540]]
 +
[[Image:St Rochus.jpg|thumb|right|Statue of St Roch with his dog, in Prague, [[Czech Republic]]]]
 +
Dogs have often appeared in art since the time of ancient [[Egypt]], where they were often included in hunting scenes painted within the tombs of important people.  In [[Renaissance]] Europe, portraits often included a favorite lap dog.
  
[[Mixed-breed dog]]s or [[Mongrel]]s are dogs that do not belong to specific breeds, being mixtures of two or more in variant percentages. Mixed breeds, or dogs with no purebred ancestry, are not inherently "better" or "worse" than purebred dogs as companions, [[pet]]s, [[working dog]]s, or competitors in [[dog sports]]. Sometimes mixed-breed dogs are deliberately bred, for example, the Cockapoo, a mixture of Cocker Spaniel and Miniature [[Poodle]]. Such deliberate crosses may display [[hybrid vigor]] and other desirable traits, but can also lack one or more of the desired traits of their parents, such as temperament or a particular color or coat. However, without genetic testing of the parents, the crosses can sometimes end up inheriting genetic defects that occur in both parental breeds. Deliberately crossing two or more breeds is also a manner of establishing new breeds.
+
Compared to some other animals, dogs have not played such an important role in [[religion]]. In [[Greek mythology]], the underworld was guarded by a three-headed dog named [[Cerberus]], who was later captured by [[Hercules]]. In [[Judaism]], the dog is considered an unclean animal and when they are mentioned in the Bible they are almost always portrayed in an unfavorable light. [[Islam]] also looks on dogs unfavorably. [[Mohammad]] expressed that the company of dogs voids a portion of a Muslim’s good deeds (Malik Ibn Anas 2:969).  
  
===Neoteny in the rapid evolution of diverse dog breeds===
+
In [[Christianity]], [[Jesus]] told the story of the poor man [[Lazarus]], whose sores were licked by street dogs. This has traditionally been seen as showing Lazarus's wretched situation. However, some modern commentators have pointed out that the dogs' saliva could have beneficial effects on the sores and that this could be seen as an example of the dogs' unconditional love (Kilcommons 1996).
{{Mergeto|Origin of the domestic dog|date=January 2007}}
 
This rapid evolution of dogs from wolves is an example of [[neoteny]] or [[pedomorphosis|paedomorphism]]. As with many species, the young wolves are more social and less [[dominant]] than adults; therefore, the selection for these characteristics, whether deliberate or inadvertent, is more likely to result in a simple retention of juvenile characteristics into adulthood than to generate a complex of independent new changes in behavior. (This is true of many domesticated animals, including humans themselves, who have many characteristics similar to young [[bonobo]]s.) This paedomorphic selection naturally results in a retention of juvenile physical characteristics as well. Compared to wolves, many adult dog breeds retain such juvenile characteristics as soft fuzzy fur, round torsos, large heads and eyes, ears that hang down rather than stand erect, etc.; characteristics which are shared by most juvenile [[mammal]]s, and therefore generally elicit some degree of protective and nurturing behavior cross-species from most adult mammals, including humans, who term such characteristics "cute" or "appealing".
 
  
The example of canine neoteny goes even further, in that the various breeds are differently neotenized according to the type of behavior that was selected.[http://books.google.com/books?id=_kOoVw0SIhUC&pg=PA394&lpg=PA394&vq=neotenous+behavior&dq=%22Gould%22+%22Eight+Little+Piggies:+Reflections+in+Natural+History%22+&sig=Q1iF36A6ieMKgh3eRAtSeDKTydQ Stephen Jay Gould; Eight Little Piggies: Reflections in Natural History; W. W. Norton & Company, 1993; pp. 394]
+
The [[Catholic Church]] recognizes Saint Roch (also called Saint Rocco), who lived in the early 1300s in France, as the patron saint of dogs.  It is said that he caught the black plague while doing charitable work and went into the forest expecting to die.  There he was befriended by a dog which licked his sores and brought him food and he was able to recover. August 16 is celebrated as the feast day of Saint Roch (Cultural Catholic 2007).
  
*[[Livestock guardian dog]]s exhibit the controlled characteristics of hunting dogs. Members of this group, such as [[Border Collie]]s, [[Belgian Shepherd Dog (Malinois)|Belgian Malinois]] and [[German Shepherd]]s use tactics of hunter and prey to intimidate and keep control of herds and flocks. Their natural instinct to bring down an animal under their charge is muted by training. Other members of the group, including [[Welsh Corgi]]s, [[Canaan dog]]s, and [[Cattle dog]]s herd with a more aggressive demeanor and make use of body design to elude the defences of their charges.
+
Dogs have often been mentioned favorably in literature. In [[Homer]]'s ''Odyssey'' (written around 800 to 600 B.C.E.) the hero's dog Argos is the only one to recognize him on his return home in disguise. One of the greatest tributes to a dog was by the English poet [[Lord Byron|Byron]] (1788-1824), who wrote of the Newfoundland, Boatswain (Byron 1808):
* [[Gun dog]] breeds used in hunting&mdash;that is, [[pointer (dog)|pointers]], [[setter]]s, [[spaniel]]s, and [[retriever]]s&mdash;have an intermediate degree of paedomorphism; they are at the point where they share in the pack's hunting behavior, but are still in a junior role, not participating in the actual attack. They identify potential prey and freeze into immobility, for instance, but refrain from then stalking the prey as an adult predator would do next; this results in the "pointing" behavior for which such dogs are bred. Similarly, they seize dead or wounded prey and bring it back to the "pack", even though they did not attack it themselves, that is, "retrieving" behavior. Their physical characteristics are closer to that of the mature wild canine than the sheepdog breeds, but they typically do not have erect ears, etc.
+
:"…Beauty without Vanity,
* [[Scent hound|Scenthound]]s maintain an intermediate body type and behavior pattern that causes them to actually pursue prey by tracking their scent, but tend to refrain from actual individual attacks in favor of vocally summoning the pack leaders (in this case, humans) to do the job. They often have a characteristic vocalization called a bay. Some examples are the Beagle, Bloodhound, Basset Hound, Coonhound, Dachshund, Fox Hound, Otter Hound, and Harrier.
+
:Strength without Insolence,
* [[Sighthound]]s, who pursue and attack perceived prey on sight, maintain the mature canine size and some features, such as narrow chest and lean bodies, but have largely lost the erect ears of the wolf and thick double layered coats.  Some examples are the Afghan, Borzoi, Saluki, Sloughi, Pharaoh Hound, Azawakh, Whippet, and Greyhound.
+
:Courage without Ferocity,
* [[Mastiff]]-types are large dogs, both tall and massive with barrel-like chests, large bones, and thick skulls.  They have traditionally been bred for war, protection, and guardian work.
+
:and all the Virtues of Man,
* [[Bulldog]]-types are medium sized dogs bred for combat against both wild and domesticated animals. These dogs have a massive, square skull and large bones with an extremely muscular build and broad shoulders.
+
:without his Vices."
* [[Terrier]]s similarly have adult aggressive behavior, famously coupled with a lack of juvenile submission, and display correspondingly adult physical features such as erect ears, although many breeds have also been selected for size and sometimes [[achondroplasia|dwarfed legs]] to enable them to pursue prey in their burrows.
 
 
 
The least paedomorphic behavior pattern may be that of the [[basenji]], bred in [[Africa]] to hunt alongside humans almost on a peer basis; this breed is often described as highly independent, neither needing nor appreciating a great deal of human attention or nurturing, often described as "catlike" in its behavior. It too has the body plan of an adult canine predator.
 
Of course, dogs in general possess a significant ability to modify their behavior according to experience, including adapting to the behavior of their "pack leaders"&mdash;again, humans. This allows them to be trained to behave in a way that is not specifically the most natural to their breed; nevertheless, the accumulated experience of thousands of years shows that some combinations of nature and nurture are quite daunting, for instance, training [[whippet]]s to guard flocks of sheep.
 
 
 
===Breed popularity===
 
[[Image:01fcirhrpup.jpg|200px|thumb|right]]
 
Breed popularity varies widely over time<Ref>
 
{{cite web
 
| url = http://www.slate.com/id/2122298/
 
| title = Why Americans Love Labrador retrievers
 
| accessdate = 2006-11-30
 
| last = Koerner
 
| first = Brendan I.
 
| authorlink =
 
| coauthors =
 
| date = [[2005-01-08]]
 
| publisher = Slate Magazine Online
 
}}</Ref>
 
and in different parts of the world and different segments of the population. Counting by [[American Kennel Club]] (AKC) registration (not by [[dog licence|licensing registration]] or by [[United Kennel Club]] (UKC) registration, which could present different statistics), the [[Labrador Retriever]] has been the United States's most commonly registered breed of dog since 1991. <Ref>
 
{{cite web
 
| url = http://www.akc.org/news/index.cfm?article_id=2389
 
| title = Labrador Retriever Tops According to AKC's 2004 Registration Statistics
 
| accessdate = 2006-11-30
 
| date = [[2005-01-12]]
 
| publisher = American Kennel Club
 
}}</Ref>
 
However, even within parts of the United States, popularity varies; for example, in 2005 the most-registered breed in New York City was the [[Poodle]] while the [[Yorkshire Terrier]] was the second-most-registered breed in [[Houston]]. <Ref>
 
{{cite web
 
| url = http://www.akc.org/reg/topdogsbycity.cfm
 
| title = Top Breeds By City
 
| accessdate = 2006-11-30
 
| publisher = American Kennel Club
 
}}</Ref>
 
However, animal shelters in many parts of the United States report that the most-commonly available dog for adoption is the [[American Pit Bull Terrier]] or pit bull-type mixes, making up as much as 20% of dogs available for adoption, none of which would be registered with the AKC.<ref>{{cite web
 
|url=http://www.aspca.org/site/PageServer?pagename=cruelty_pitbull
 
|title=Pit Bull Cruelty
 
|publisher=American Society for the Prevention of cruelty to Animals
 
|accessdate=2007-05-11}}</ref> Two decades ago, in 1983, the AKC's top two registered breeds were the [[American Cocker Spaniel]] and the [[Poodle]].<ref>{{cite book|title=World Almanac and Book of Facts|year=1985|publisher=Newspaper Enterprise Association (Doubleday)}}</ref>
 
 
 
In 2006 a British dog journal published an exhaustive research paper which pooled together quantifiable dog breed facts such as average lifespan, cost of maintenance, insurance claims made, known behavioural faults and various other criteria in order to arrive at the conclusion that the Staffordshire Bull Terrier is the 'highest scoring' dog breed in terms of desirable and established pedigree dog breed traits. <Ref>
 
{{cite web
 
| url = http://www.k9magazine.com
 
| title = British Dog Magazine
 
| accessdate = 2007-05-05
 
| publisher = K9 Media
 
}}</Ref>
 
 
 
In the [[United Kingdom]], [[The Kennel Club]] reports that the most-registered breed from at least 1999 to 2005 was the Labrador Retriever. It rounds out the top three for 1999 to 2005 with the [[German Shepherd Dog]], also popular in the States, and the [[English Cocker Spaniel]]<Ref>
 
{{cite web
 
| url = http://www.the-kennel-club.org.uk/pressoffice/press_top20.html
 
| title = The Kennel Club's top twenty of registered breeds
 
| accessdate = 2006-11-30
 
| publisher = The Kennel Club
 
}}</Ref>
 
, which is no longer even in the top ten in the States. In the United Kindom a national dog adoption and rescue service indicates that the most common breed appearing in shelters is the Greyhound followed by the Staffordshire Bull Terrier. <Ref>
 
{{cite web
 
| url = http://www.dogsblog.com
 
| title = UK dog adoption and rescue service
 
| accessdate = 2007-05-05
 
| publisher = Dogs Blog
 
}}</Ref>
 
 
 
==In religion==
 
===Islam===
 
 
 
:''See also [[Islam and animals#Views regarding particular animals|Islam and animals]]''
 
 
 
[[Muhammad]], the prophet of [[Islam]] expressed that the company of dogs voids a portion of a Muslim’s good deeds.<ref> Malik Ibn Anas, al-Muwatta’ (Egypt: al-Babi al-Halabi, n.d.), 2:969. Reported in El Fadl</ref>
 
 
 
Islamic law considers dogs to be unclean but this is not derived from the Qur'an.<ref name="EoQ_Dog"/> Muhammad is said to have given an order to kill all the dogs in Medina because the angel Gabriel does not enter a house in which there is a dog. This hadith is however the "occasion for revelation" of verse {{Quran-usc|5|4}} which indicates the importance of the hunting dog.<ref name="EoQ_Dog"> Bruce Fudge, [[Encyclopedia of the Quran]], ''Dog'' </ref> David Gordon White states that Islam, like Judaism and Christianity, usually cast the dog in a negative light by emphasizing its impurity and often identify dogs as demons or minions of the Devil. The story of the Seven sleepers of Ephesus in the Qur'an (and also role of the dog in early Christianity) are the striking exceptions.<ref> David Gordon White, Encyclopedia of religion, ''Dog'', p.2393 </ref>. Muhammad didn't like dogs (and most Muslims do not have dogs as pets). According to one tradition attributed to Muhammad, black dogs are evil, or even devils, in animal form.<ref>{{cite web
 
|url=http://www.scholarofthehouse.org/dinistrandna.html
 
|title=Dogs in the Islamic Tradition and Nature
 
|publisher=Dogs in the Islamic Tradition and Nature, quoting Encyclopedia of Religion and Nature
 
|author=Dr. Khaled Abou El Fadl
 
|year=[[2004]]
 
|accessdate=2007-05-11}}</ref>
 
 
 
Muhammad commanded Muslims not trade or deal in dogs.<ref>Ahmad Ibn Shu‘ayb al-Nisa’i, Sunan al-Nisa’i (Beirut: Dar al-Kitab al-‘Arabi, n.d.), 7: 309 (The commentaries by al-Suyuti and al-Sanadi are in the margins). Ibn Hajar al-‘Asqalani, Fath al-Bari, 4:426. All reported in El Fadl.</ref>
 
 
 
Muhammad said that dogs and [[donkey]]s - if they pass in front of men in prayer - they will void or nullify that prayer.<ref>Al-Nawawi, Sahih Muslim, 3-4:450-1; Ahmad Ibn Hanbal, Musnad, 5:194, 197, 202, 208; Abu Bakr Ibn al-‘Arabi, ‘Aridat al-Ahwadhi bi Sharh Sahih al-Tirmidhi (Beirut: Dar al-Kutub al-‘Ilmiyya, n.d.), 1:133. All reported in El-Fadl.</ref>
 
 
 
Dr. [[Khaled Abou El Fadl]], a professor of law at the [[UCLA School of Law]] where he teaches [[Islamic law]], has written on the subject.<ref>{{cite web
 
|url=http://www.scholarofthehouse.org/dinistrandna.html
 
|title=Chapter 80 of The Search for Beauty in Islam The Lord of the Essence: A Fatwa on Dogs
 
|author=Dr. Khaled Abou El Fadl
 
|publisher=www.scholarofthehouse.org
 
|accessdate=2007-05-14}}</ref><ref>{{cite web
 
|url=http://scholarofthehouse.stores.yahoo.net/tloofesfaond.html
 
|title=Dogs in the Islamic Tradition and Nature
 
|author=Dr. Khaled Abou El Fadl
 
|publisher=www.scholarofthehouse.org
 
|accessdate=2007-05-14}}</ref>
 
 
 
==See also==
 
*[[Origin of the domestic dog]]
 
*[[Ability to swim]]
 
* [[Attack dog]]
 
* [[Bark (dog)]]
 
* [[Dog attack]]
 
* [[Dog communication]]
 
* [[Dog licence]]
 
* [[Dog odor]]
 
* [[Fear of dogs]]
 
* [[List of dog breeds]]
 
* [[Puppy]]
 
* The [[Dog king]]
 
* [[List of most popular dog breeds]]
 
* [[Therapy dog|Therapy Dog]]
 
* [[Guard dog]]
 
* [[War Dog]]
 
* [[Family Dog]]
 
* [[Wolf-dog hybrid]]
 
  
 
==References==
 
==References==
 
+
* Alderton, D. 1984. ''The Dog''. London: Chartwell Books. ISBN 0890097860
{{reflist|2}}
+
* American Kennel Club (AKC). 2006. ''The Complete Dog Book''. New York: Ballantine Books. ISBN 0345476263
 
+
* American Society for the Prevention of Cruelty to Animals (ASPCA). 2014. [https://www.aspca.org/pet-care/top-10-reasons-spay-or-neuter-your-pet Top 10 reasons to spay or neuter your pet]. ''ASPCA''. Retrieved December 18, 2014.  
==Further reading==
+
* American Veterinary Medical Association (AVMA). 2002. [http://www.avma.org/reference/marketstats/ownership.asp Market research statistics: Cat and dog ownership]. ''AVMA''. Retrieved August 6, 2007.
* Abrantes, Roger (1999). ''Dogs Home Alone''. Wakan Tanka, 46 pages. ISBN 0-9660484-2-3 (paperback).
+
* Byron, G. G. 1808. [http://www.poetryloverspage.com/poets/byron/byron_ind.html Epitaph to a dog]. ''Poetry Lovers Page''. Retrieved August 2, 2007.
* A&E Television Networks (1998). ''Big Dogs, Little Dogs: The companion volume to the A&E special presentation'', A Lookout Book, GT Publishing. ISBN 1-57719-353-9 (hardcover).
+
* Clutton-Brook, J. 1999. ''A Natural History of Domesticated Mammals.'' Cambridge, UK: Cambridge University Press. ISBN 0521634954
* Alderton, David (1984). ''The Dog'', Chartwell Books. ISBN 0-89009-786-0.
+
* Cultural Catholic LLC. 2007. [http://www.culturalcatholic.com/SaintRoch.htm Saint Roch]. ''Cultural Catholic''. Retrieved August 8, 2007.
* Brewer, Douglas J. (2002) ''Dogs in Antiquity: Anubis to Cerberus: The Origins of the Domestic Dog'', Aris & Phillips ISBN 0-85668-704-9
+
* Davis, J. 1998. [http://www.uwsp.edu/psych/dog/LA/davis2.htm Color and acuity differences between dogs and humans]. University of Wisconsin, Stevens Point. Retrieved August 2, 2007.
* Coppinger, Raymond and Lorna Coppinger (2002). ''Dogs: A New Understanding of Canine Origin, Behavior and Evolution'', University of Chicago Press ISBN 0-226-11563-1
+
* Humane Society of the United States (HSUS). 2007. [http://www.hsus.org/animals_in_research/species_used_in_research/dog.html Animals used in research: Dog profile]. ''Human Society of the United States''. Retrieved August 6, 2007.
*Cunliffe, Juliette (2004). ''The Encyclopedia of Dog Breeds''. Parragon Publishing. ISBN 0-7525-8276-3.
+
* Invasive Species Specialist Group (ISSG). 2007. [http://www.issg.org/database/species/ecology.asp?si=146&fr=1&sts=sss ''Canis lupus'' (mammal)]. ''Invasive Species Specialist Group''. Retrieved August 8, 2007.
* Derr, Mark (2004). ''Dog's Best Friend: Annals of the Dog-Human Relationship''. University of Chicago Press. ISBN 0-226-14280-9
+
* Killcommons, B., and M. Capuzzo. 1996. ''Mutts: America's Dogs''. New York: Time Warner. ISBN 0446519499
* Donaldson, Jean (1997). ''The Culture Clash''. James & Kenneth Publishers. ISBN 1-888047-05-4 (paperback).
+
* Kruuk, H. 2002. ''Hunter and Hunted: Relationships Between Carnivores and People.'' Cambridge, UK: Cambridge University Press. ISBN 0521814103
*Fogle, Bruce, DVM (2000). ''The New Encyclopedia of the Dog''. Doring Kindersley (DK). ISBN 0-7894-6130-7.
+
* Malik Ibn Anas, al-Muwatta’ (Egypt: al-Babi al-Halabi, n.d.), 2:969. Reported in El Fadl
*Grenier, Roger (2000). ''The Difficulty of Being a Dog''. University of Chicago Press. ISBN 0-226-30828-6
+
* McGourty, C. 2002. [http://news.bbc.co.uk/2/hi/science/nature/2498669.stm Origin of dogs traced]. ''BBC News''. Retrieved July 23, 2007.
* Milani, Myrna M. (1986). ''The Body Language and Emotion of Dogs: A practical guide to the Physical and Behavioral Displays Owners and Dogs Exchange and How to Use Them to Create a Lasting Bond'', William Morrow, 283 pages. ISBN 0-688-12841-6 (trade paperback).
+
*Nelson, R. W., and C. G. Couto. 2003. ''Small Animal Internal Medicine, Third Edition''. Philadelphia: Mosby. ISBN 032301724X
* Pfaffenberger, Clare (1971). ''New Knowledge of Dog Behavior''. Wiley, ISBN 0-87605-704-0 (hardcover); Dogwise Publications, 2001, 208 pages, ISBN 1-929242-04-2 (paperback).
+
* Nowak, R. M., and J. L. Paradiso. 1983. ''Walker's Mammals of the World''. Baltimore: The Johns Hopkins University Press. ISBN 0801825253
* Savolainen, P. et al. (2002). Genetic Evidence for an East Asian Origin of Domestic Dogs. ''Science'' '''298'''. 5598: 1610&ndash;1613.
+
* Sandeep, K. 2002. Man's worst enemy. ''The Hindu''.
* Shook, Larry (1995). "Breeders Can Hazardous to Health", ''The Puppy Report: How to Select a Healthy, Happy Dog'', Chapter Two, pp. 13&ndash;34. Ballantine, 130 pages, ISBN 0-345-38439-3 (mass market paperback); Globe Pequot, 1992, ISBN 1-55821-140-3 (hardcover; this is much cheaper should you buy).
+
* Savolainen, P.,  Y. Zhang, et al. [http://adsabs.harvard.edu/abs/2002Sci...298.1610S Genetic evidence for an East Asian origin of domestic dogs]. ''Science''. Retrieved July 24, 2007.
* Shook, Larry (1995). ''The Puppy Report: How to Select a Healthy, Happy Dog'', Chapter Four, "Hereditary Problems in Purebred Dogs", pp. 57&ndash;72. Ballantine, 130 pages, ISBN 0-345-38439-3 (mass market paperback); Globe Pequot, 1992, ISBN 1-55821-140-3 (hardcover; this is much cheaper should you buy).
+
* Voelker, W. 1986. ''The Natural History of Living Mammals''. Medford, New Jersey: Plexus Publishing. ISBN 0937548081
* Thomas, Elizabeth Marshall (1993). ''The Hidden Life of Dogs'' (hardcover), A Peter Davison Book, Houghton Mifflin. ISBN 0-395-66958-8.
+
* Wayne, R. K., C. Vila, et al. 1997. [http://www.mnh.si.edu/GeneticsLab/StaffPage/MaldonadoJ/PublicationsCV/Science_Dog_Paper.pdf Multiple and ancient origins of the domestic dog]. ''Science''. June 13, 1997. Retrieved July 24, 2007.
* Verginelli, F. et al. (2005). Mitochondrial DNA from Prehistoric Canids Highlights Relationships Between Dogs and South-East European Wolves. ''Mol. Biol. Evol.'' '''22''': 2541&ndash;2551.
+
* World Health Organization (WHO). 2006. [http://www.who.int/mediacentre/factsheets/fs099/en/ Rabies]. ''World Health Organization''. Retrieved August 8, 2007.
*''Small animal internal medicine'', RW Nelson, Couto page 107
 
 
 
==External links==
 
{{sisterlinks}}
 
{{Wikispecies}}
 
 
 
*[http://www.akc.org/ American Kennel Club]
 
*[http://www.ankc.aust.com/breed_list.html Australian National Kennel Club]
 
*[http://www.ckc.ca/ Canadian Kennel Club]
 
*[http://www.fci.be Fédération Cynologique Internationale (FCI) - World Canine Organisation]
 
*[http://www.dogdomain.com/fci-1.htm FCI International breed standards]
 
*[http://www.the-kennel-club.org.uk The Kennel Club (UK)]
 
*[http://www.nzkc.org.nz/dogselect.html New Zealand Kennel Club]
 
*[http://www.doggeek.com Dog information and resources]
 
  
  
{{credit|Dog|138982039}}
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{{Credit8|Dog|138982039|Disc_dog|147147444|Sheepdog_trial|122079811|Earthdog_trial|145340673|Field_trial|135006059|Obedience_trial|149131153|Tracking_trial|139330271|Flyball|141312739}}
 
[[Category:Life sciences]]
 
[[Category:Life sciences]]
 +
[[Category:Animals]]
 +
[[Category:Mammals]]
 +
[[Category:Carnivores]]

Latest revision as of 11:04, 2 August 2023

Domestic dog
Fossil range: Late Pleistocene - Recent
other images of dogs
Conservation status
Conservation status: Domesticated
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Canidae
Genus: Canis
Species: C. lupus
Subspecies: C. l. familiaris
Trinomial name
Canis lupus familiaris

The dog (Canis lupus familiaris or Canis familiaris) is usually considered to be humankind's first, and perhaps most important, domestic animal. It generally is classified as a subspecies of the gray wolf (Canis lupus), a mammal of the Canidae family of the order Carnivora.

The word "dog" also is used to refer to some other members of the Canidae family, for instance the bush dog (Speothos venaticus) of South America and the raccoon dog (Nyctereutes procyonoides) of Asia. The family Canidae itself often is called the "dog family" and includes jackals and coyotes, which are closely related to dogs and wolves, as well as foxes, which are less closely related. Members of the Canidae family are called "canids."

The trainability, devotion, playfulness, and ability to fit into human households have earned dogs a unique position in the realm of interspecies relationships. Dogs have lived and worked with humans in so many roles—hunting, herding and protecting sheep and cattle, guarding people and property, rescuing people, companionship, and so forth—that their loyalty has earned them the unique sobriquet, "man's best friend." Their actions mimic the human idea of love and friendship, leading many dog owners to view their pets as family members.

In a process generally thought to span at least 14,000 years, humans have domesticated the dog, produced innumerable breeds, and bred and trained them to perform a great variety of useful services and sometimes dubious actions, including as guard dogs trained to instill fear in people and as sports dogs taught to fight other dogs to the death.

Lateral view of a dog's bone structure.

Characteristics

Dogs, like wolves and other canids, are specialized for a life of hunting and eating other animals. Their bodies are compact and muscular and their legs are long, which gives them speed and endurance for chasing prey. Their jaws are large and their teeth are well suited for grasping, biting, and tearing flesh. Their digestive systems, however, are suited for some plant foods as well as flesh.

Most dogs are covered with a thick coat of fur, which protects them from the elements. Alaskan malamutes and Siberian huskies, which originated in the Arctic, are able to live outside without shelter in temperatures far below freezing. Dogs' coats come in a wide range of colors, including black, white, brown, yellow, red, and gray. They come in a range of lengths and textures from very short to long, either straight or curly (AKC 2006).

A dog's heart and lungs are oversized relative to its body and its normal everyday needs. A dog also has relatively more red blood cells than a human. Most of the time the dog will keep the extra red blood cells stored in its spleen. When the animal enters into a situation where its full metabolism is required, such as play, catching game, or fighting other dogs, the extra cells are released into the bloodstream. The "oversized" heart and lungs would then be running at full capacity, and the animal would have an enhanced ability to engage in aerobic activity. This activity produces internal heat. Dogs, being covered in fur, are limited in their ability to cool down. After a short time the animal must either cease its athletic activity or risk harming itself from overheating. One can easily observe this pattern of intense activity followed by rest periods in puppies. During the rest phase, the spleen collects red blood cells and the animal may pant to cool down.

Dogs, like most carnivores, are naturally active only 4 to 6 hours each day and prefer to sleep the rest of the time (Kruuk 2002).

Dogs, like other canids, are considered to be among the most intelligent of all animals, as shown by their ability to adapt to changing conditions in human society.

The dog's ancestor, the gray wolf, is the largest canid; with adults weighing between 18 and 80 kg (40 to 175 lbs). Modern dogs can weigh anywhere from 1 to over 100 kg (2.2 to 220 lbs). Dingos, wild dogs of Australia which are descended from domestic dogs, average 10 to 20 kg (22 to 44 lbs) in weight (Nowak 1983).

Senses

Sight

In general, the vision of dogs is not as good as that of humans. They have only limited color vision, equivalent to red-green color blindness in humans (Alderton 1984). They also are nearsighted compared to humans and can not see detail as well. The average dog's vision seems to be about 20/50 to 20/100, using the human standard. On the other hand they see better in dim light and may also detect motion better than humans (Davis 1998).

Hearing

This mixed-breed dog angles her ears to find the source of a sound.

The hearing of dogs is superior to humans. They can detect sounds that are lower and those that are higher in frequency. They also have a degree of ear mobility that helps them to rapidly pinpoint the exact location of a sound. Eighteen or more muscles can tilt, rotate, and raise or lower a dog's ear. Additionally, a dog can identify a sound's location much faster than a human can, as well as hear sounds up to four times the distance that humans are able to. Those with more natural ear shapes, like those of wild canids, generally hear better than those with the floppy ears of many domesticated species.

Smell

Scent hounds, especially the Bloodhound, are known for their keen sense of smell

Dogs have nearly 220 million smell-sensitive cells over an area about the size of a pocket handkerchief (compared to 5 million over an area the size of a postage stamp for humans). Some breeds have been selectively bred for excellence in detecting scents, even compared to their canine brethren.

What information a dog actually detects when it is scenting is not perfectly understood. Although once a matter of debate, it now seems to be well established that dogs can distinguish two different types of scents when trailing: An air scent from some person or thing that has recently passed by, as well as a ground scent that remains detectable for a much longer period. The characteristics and behavior of these two types of scent trail would seem, after some thought, to be quite different, the air scent being intermittent but perhaps less obscured by competing scents, whereas the ground scent would be relatively permanent with respect to careful and repetitive search by the dog, but would seem to be much more contaminated with other scents. In any event, it is established by those who train tracking dogs that it is impossible to teach the dog how to track any better than it does naturally; the object instead is to motivate it properly, and teach it to maintain focus on a single track and ignore any others that might otherwise seem of greater interest to an untrained dog. An intensive search for a scent, for instance searching a ship for contraband, can actually be very fatiguing for a dog, and the dog must be motivated to continue this hard work for a long period of time.

Reproduction

A Catahoula leopard dog mother nursing her litter of puppies.

In domestic dogs, sexual maturity (puberty) begins to happen around age 6 to 12 months for both males and females, although this can be delayed until up to two years old for some large breeds. Adolescence for most domestic dogs is around 12 to 15 months, beyond which they are for the most part more adult than puppy. As with other domesticated species, domestication has selectively bred for higher libido and earlier and more frequent breeding cycles in dogs than in their wild ancestors.

Dogs bear their litters roughly 9 weeks after fertilization, although the length of gestation can vary from 56 to 72 days. A general rule of thumb is that a mammal will produce half as many offspring as the number of teats on the mother. This rule is altered in domesticated animals since larger litters are often favored for economic reasons and in dogs, particularly, the great range of sizes and shapes plays a role in how many healthy puppies a female can carry. An average litter consists of about six puppies, though this number may vary widely based on the breed of dog. Small dogs generally produce from one to four puppies in each litter, while much larger breeds may average as many as 12 pups in each litter. The number of puppies also varies with the mother's age and health, the father's sperm count, the timing of the breeding, and many other factors.

Lifespan

The typical lifespan of dogs varies considerably by breed. For example, many giant dog breeds (such as great danes) average only 7 or 8 years, while some small terrier breeds, or toy breeds, might live to be older than 20. The average lifespan for mixed-breed and midsize dogs is about 13 to 14 years. The longest-lived dog with reliable documentation died at 29 years and 27 days; the breed of the dog was an Australian cattle dog and it lived in Virginia, United States, with the date of death in 1939. Although the lifespans of all living species are mostly uncontrollable, one can significantly extend a dog's life by feeding it the right kinds of foods, giving it regular exercise, treating its diseases, caring for its special needs, and giving it love and comfort.

Social behavior

Dogs, like almost all canids, are social animals. Wolves, and it is assumed the ancestors of dogs, almost always live in pairs, a male and a female with their young, or in larger groups called packs. Within the pack, the individual animals communicate with each other by visual signals, such as body and facial expressions and tail wagging, as well as by vocal signals, such as growls and howls. Pack members sometimes hunt individually and sometimes as a group, depending on the situation, including the size of the prey (Kruuk 2002).

The members of the pack cooperate in many activities, such as hunting, caring for the young, and defending their territory from intruders. Within each pack, there is a dominant pair which takes the lead, while the others support them. In most cases, only the dominant pair reproduces. Wolves struggle to establish dominance and sometimes fight with each other, but this seldom leads to serious injury or death since the losing animal will display submissive behavior which discourages further attacks. Studies of feral dogs have found that they often form packs of 3 to 5 individuals (Clutton-Brook 1999; Nowak 1993; Voelker 1986).

Origins

According to recent genetic research, the lineage of dogs separated from that of wolves about 100,000 years ago (Wayne 1997). It is not known when the ancestors of dogs began to associate with humans. The first fossil evidence of domesticated dogs, which had changed in physical form from their wolf ancestors, dates from around 15,000 years ago. Dog fossils from about this time have been found in Asia, Europe, North America, and South America (Clutton-Brock 1999).

Other genetic research seems to indicate that China was the site of the original domestication of dogs (McGourty 2002). Genetic research has identified 14 ancient dog breeds, with the oldest being the Chow Chow, Shar Pei, Akita Inu, Shiba Inu, and Basenji. Many of the 14 breeds are associated with China and Japan (Savolainen 2002).

Human relationships

Dogs are highly social animals. This can account for their trainability, playfulness, and ability to fit into human households and social situations. This similarity has earned dogs a unique position in the realm of interspecies relationships.

The loyalty and devotion that dogs demonstrate as part of their natural instincts as pack animals closely mimics the human ideas of love and friendship, leading many dog owners to view their pets as full-fledged family members. Conversely, dogs seem to view their human companions as members of their pack, and make few, if any, distinctions between their owners and fellow dogs.

Dogs fill a variety of roles in human society and are often trained as working dogs. For dogs that do not have traditional jobs, a wide range of dog sports provide the opportunity to exhibit their natural skills. In many countries, the most common and perhaps most important role of dogs is as companions or pets. In the United States, about 36 percent of households own at least one dog, with the total number of animals about 62 million (AVMA 2002).

Dogs have lived and worked with humans in so many roles that their loyalty has earned them the unique sobriquet, "man's best friend."

Terminology

The English word "dog," in common usage, refers to the domestic pet dog, Canis lupus familiaris. The species was originally classified as Canis familiaris by Linnaeus in 1758. In 1993, dogs were reclassified as a subspecies of the gray wolf, Canis lupus, by the Smithsonian Institution and the American Society of Mammalogists.

The English word "dog" might derive from the Old English "docga," a "powerful breed of canine," like the modern mastiff. The English word "hound" is a cognate of German Hund, Dutch hond, common Scandinavian hund, Icelandic hundur, which, though referring to a specific breed in English, means "dog" in general in the other Germanic languages. "Hound" itself derives from the Proto-Indo-European *kwon-, which is the direct root of the Greek κυων (kuōn) and the indirect root of the Latin canis through the variant form *kani-.

In breeding circles, a male canine is referred to as a "dog," while a female canine is called a "bitch." The father of a litter is called the "sire," and the mother of a litter is called the "dam." Offspring are generally called "pups" or "puppies" until they are about a year old. A group of offspring is a "litter." The process of birth is "whelping." Many terms are used for dogs that are not purebred.

The constellations Canes Venatici, Canis Major, and Canis Minor are named from the Latin word for "dog," for their perceived resemblance to dogs. Sirius, the brightest star in the sky as seen from earth, is called the "dog star" because it is found in Canis Major. The time when Sirius rises just before sunrise, mainly July and August, are called the "dog days of summer."

Breeds

Different breeds of dogs emerged over time due to environmental factors and human selection. In ancient Egypt, dogs looking like the modern greyhound and the mastiff are pictured in art. In ancient Roman writings, miniature pet dogs like modern toy breeds are mentioned (Clutton-Brook 1999).

There are said to be over 450 different dog breeds in the world. As of 2006, the American Kennel Club recognized 153 breeds, divided into 7 groups (AKC 2006):

  • Sporting breeds were originally bred and trained to assist gun-wielding hunters in finding and recovering birds and other small game. They include various pointers, spaniels, and retrievers.
  • Hounds were bred to track and engage larger hunted animals. They include the bloodhound, the English foxhound, and the beagle.
  • Working breeds were bred for various tasks; such as guarding people, livestock, and property; pulling sleds; and rescuing people from water or snow. They include the rotweiler, the saint bernard, and the akita.
  • Terriers were originally bred to catch and kill small animals, especially rats, without the help of a human hunter. Popular terriers include the airedale terrier and the Parson Russell terrier (also called the Jack Russell terrier).
  • Toy Breeds are smaller dogs bred to be pets. They include the pug, the pekingese, and the chihuahua.
  • Non-sporting breeds are medium-sized or large dogs bred to be pets, or in the case of the dalmation, a companion to horses. The poodle is one of the most popular non-sporting breeds.
  • Herding breeds were bred to herd sheep or cattle. They include the collie, the German shepherd, and the Old English sheepdog.

Through history different dog breeds have gone in and out of fashion. In England, the King Charles Spaniel fell out of favor in 1689 when the house of Stuart was overthrown in the Glorious Revolution and was reborn in the twentieth century as the Cavalier King Charles Spaniel. In the United States, the beagle became very popular in the 1960s because of the cartoon character Snoopy. The most popular breed in the United States today is the Labrador retriever (AKC 2006).

Mixed breed dogs

Mixed-breed dogs, such as this Australian Border Collie cross, are often desired as working animals for their robust health and intelligence.

The majority of dogs in the world are of mixed breed or no definite breed. It is estimated that 60 percent of dogs in the United States are mixed breeds, and the number it thought to be much higher in poorer countries. Advocates of mixed breed dogs as working dogs and pets say that they are more intelligent and have fewer health problems than purebreds (Kilcommons 1996).

Working, utility, and assistance dogs

Labrador retrievers are often used as assistance dogs.
Belgian Malinois war dog.

The first ways that dogs were useful to humans were as helpers in hunting and as protectors. These roles grew out of the dogs' natural instincts. As time went on, dogs were trained to perform other tasks and were specially breed to do so. Some of the modern types of working dogs are:

  • Assistance dogs help the seeing and hearing-impaired. Others are trained to help those with epilepsy and psychiatric disorders, by detecting the onset of the condition so they can seek help. The typical assistance dog is a sociable breed such as a Labrador retriever. In the United States, people with assistance dogs are guaranteed access to public places by law.
  • Detection dogs are trained to, and work at, using their senses (almost always the sense of smell) to detect substances such as explosives or illegal drugs.
  • Guard dogs are trained to protect people and property. These are commonly doberman pinschers, German shepherds, and rottweilers, or mixes of these breeds. Guard dogs differ from watch dogs in that they are expected to engage an intruder, while watch dogs are only expected to bark to alert the property owner.
  • Herding dogs are trained to control livestock in the range. Sheepdogs and collies are common breeds.
  • Hunting dogs search for and retrieve game. Pointers and retrievers are typical hunting breeds.
  • Police dogs, typically German shepherds, are trained to assist law enforcement officers, often apprehending fleeing suspects. Schutzhund is a special test developed in Germany in the early 1900s to test if a German shepherd has the right talents and temperment to be successfully trained as a police dog.
  • Rescue dogs are those that search for missing humans. The archetypal breeds are the Saint Bernard and the bloodhound, although any dog with a good sense of smell can be trained to perform this task. One of the most famous dog rescues in history is that of Napoleon Bonaparte, a poor swimmer, who fell into the ocean one night in 1814, and was rescued by a Newfoundland.
  • Show dogs do tricks for the entertainment of audiences. Poodles and American Eskimo dogs were very popular as show dogs in circuses and traveling shows in the past. Now many types of dog perform in movies and television.
  • Therapy dogs, with friendly and gentle temperaments, are trained to provide comfort and affection to hospitalized and institutionalized patients. The modern concept of the therapy dog was begun in 1976 by Elaine Smith, an American nurse.
  • War dogs are used by the military to detect mines and enemy soldiers. War dogs, looking much like modern mastiffs, are pictured in ancient Assyrian art. Dogs continued to be used in war through Roman and Medieval times. In 1560, Queen Elizabeth I of England sent 8,000 war dogs, equipped with spiked collars and coats of mail, with her army sent to Ireland to suppress a rebellion (Kilcommons 1996). In World War II, the Soviet army trained dogs to carry explosives and run under enemy tanks, blowing them up.

Show and sport (competition) dogs

An English Springer spaniel in a dog show
The border collie uses a direct stare at sheep, known as "the eye," to intimidate while herding at a trial.

Owners of dogs often enter them in competitions, whether show (breed conformation shows) or sports, including dog racing and dog sledding. The winners garner both prestige and prize money. These dogs are often bred specifically for competition, which may not be entirely beneficial for the breeds, due to the magnification of hereditary defects.

  • Dog agility is a sport in which dogs complete a timed obstacle course. It is a fairly new sport, started in 1978 in the United Kingdom.
  • Tracking trials encourage dogs to make use of their strongest facility by emulating the finding of a lost person or article in a situation where the performance of the dog can be fairly assessed. Because of this, the tracks laid are not the wanderings, which may characterize a lost person, nor do they include deliberate attempts by the tracklayer to deceive the following dog.
  • Dogsled racing is a winter sport where a team of dogs pull a sled and driver (called a musher). The Iditarod Trail Sled Dog Race is perhaps the most famous of these races. Dog sledding is an ancient form of transportation and still a very effective way of moving freight across this type of terrain. Most modern dog sledders use mixed breed dogs.
  • Dog racing, almost always emphasizing greyhounds, involves dogs racing at betting tracks in a sport not unlike horse racing, reaching speeds of 40 miles per hour. Elsewhere, dachshunds are often raced, as a humorous sidelight in charity events.
  • Dog fighting and dog baiting are blood sports involving dogs. In the past, they were very popular but now they are illegal in most jurisdictions, but are still occasionally performed underground. Some of the breeds which have been used in dog fighting are the akita, the Irish terrier, and the American pit bull terrier (often called the pit bull).
  • Disc dog is the more generic name for what is commonly called Frisbee dog. In disc dog competitions, dogs and their human disc throwers compete in events such as distance catching and somewhat choreographed freestyle catching. The sport celebrates the bond between handler and dog, by allowing them to work together. The term "disc" is preferred because "Frisbee" is a trademark (held by Wham-O) for a brand of flying disc.
  • Flyball is an event in which teams of dogs race against each other from a start/finish line, over a line of hurdles, to a box that releases a tennis ball to be caught when the dog presses the spring loaded pad, then back to their handlers while carrying the ball. Flyball has become popular because it is a sport in which all sizes and breeds of dog can compete.
  • Sheepdog trials (or simply dog trials) are competitive events in which herding dog breeds move sheep around a field, fences, gates, or enclosures as directed by their handlers. Such events are particularly associated with hill farming areas, where sheep range widely on largely unfenced land. These trials are popular in the United Kingdom, Ireland, South Africa, Canada, the United States, Australia, New Zealand, and other farming nations. Some venues allow only dogs of known herding breeds to compete; others allow any dog that has been trained to herd.
  • Earthdog trials test dachshunds and small terriers for their working abilities and instinct. Earthdog den trials involve underground tunnels that dogs must negotiate, while scenting a rat, the "quarry." The dog must follow the scent to the quarry and then "work" the quarry. Depending on the sanctioning organization, “working” means barking, scratching, staring, pawing or such at it, although the quarry is protected at all times.
  • Field trials are highly competitive events at which hunting dogs compete against one another. Field trials are organized by kennel clubs or other gun dog organizations.
  • Obedience trials are events in which a dog must perfectly execute a predefined set of tasks when directed to do so by its handler. Obedience competition provides an opportunity for a person and a dog to work as a highly tuned team.
  • Dog shows involve purebreds entered in conformation dog show and evaluated by how closely they match the ideal characteristics of the breed. The Westminster Kennel Club Dog Show is one of the most famous of this type of competition.

Dogs in research

Dogs have long been used as subjects of scientific research. In the 1890s, the Russian scientist Ivan Pavlov used dogs to study psychological conditioning. Much of the research which led to the discovery of insulin and to the successful treatment of diabetes in humans was carried out using dogs as experimental subjects. On November 3, 1957, a Russian dog, Laika, became the first animal to orbit the earth.

Dogs presently make up only a small percentage of animals used in research; rats and mice far outnumber them. However, they are especially important in medical studies, for instance on the effects of drugs. Among the reasons for this are their fairly large size and their ease in being taken care of and controlled.

Beagles are especially popular for research because of their calm and friendly nature. Most research beagles are especially bred for this purpose. Other research dogs are obtained from various sources including pound seizure, newspaper ads, strays, and lost or stolen pets. Over the years there have been protests against the treatment of research dogs and laws have been passed to protect them, and in some cases to prevent former pet dogs from being used in research (HSUS 2007).

Dog health

Dogs are susceptible to various diseases, ailments, and poisons, some of which effect humans in the same way, others of which are unique to dogs.

Diet

At present, there is some debate as to whether domestic dogs should be classified as omnivores or carnivores, by diet.

The classification in the Order Carnivora does not necessarily mean that a dog's diet must be restricted to meat; unlike an obligate carnivore, such as the cat family with its shorter small intestine, a dog is dependent on neither meat-specific protein nor a very high level of protein in order to fulfill its basic dietary requirements. Dogs are able to healthily digest a variety of foods including vegetables and grains, and in fact dogs can consume a large proportion of these in their diet. Wild canines not only eat available plants to obtain essential amino acids, but also obtain nutrients from vegetable matter from the stomach and intestinal contents of their herbivorous prey, which they usually consume.

Domestic dogs can survive healthily on a reasonable and carefully designed vegetarian diet, particularly if eggs and milk products are included. Some sources suggest that a dog fed on a strict vegetarian diet may develop dilated cardiomyopathy since it lacks L-carnitine (Nelson 2003), however, maintaining a balanced diet is also a factor since L-carnitine is found naturally in many nuts, seeds, beans, vegetables, fruits, and whole grains. In the wild, dogs can survive on a vegetarian diet when animal prey is not available. However it has been noted, both by observation of extremely stressful conditions such as the Iditarod Trail Sled Dog Race and by scientific studies of similar conditions, that high-protein (approximately 40 percent) diets including meat help prevent damage to muscle tissue. (This research is also true for some other mammals.) This level of protein corresponds to the percentage of protein found in the wild dog's diet when prey is abundant; higher levels of protein seem to confer no added benefit.

Spaying and neutering

In the United States, most pet dogs are spayed or neutered. Spaying is the surgical removal of the female's ovaries and uterus. Neutering is the removal of the male's testicles. Spayed and neutered dogs avoid some diseases such as breast and testicular cancer. Behaviors such as urinating in the house, running away, and aggression are also reduced; and unwanted litters of puppies are avoided, which could contribute to the problem of stray and feral dogs (ASPCA 2014).

Diseases

Infectious diseases commonly associated with dogs include rabies (hydrophobia), canine parvovirus, and canine distemper. Congenital diseases of dogs can include a wide range from hip dysplasia and medial patellar luxation to epilepsy and pulmonic stenosis. Canines can get just about anything a human can get (excluding many infections, which are species specific) like hypothyroidism, cancer, dental disease, heart disease, Addison's disease, and so forth.

Two serious medical conditions affecting dogs are pyometra, affecting unspayed females of all types and ages, and bloat, which affects the larger breeds or deep chested dogs. Both of these are acute conditions, and can kill rapidly; owners of dogs which may be at risk should learn about such conditions as part of good animal care.

Parasites

Common external parasites are various species of fleas, ticks, and mites. Internal parasites include hookworms, tapeworms, roundworms, and heartworms.

Common physical disorders

Some breeds of dogs are also prone to certain genetic ailments, such as hip dysplasia, luxating patellas, cleft palate, blindness, or deafness. As noted, dogs are also susceptible to the same ailments that humans are, including diabetes, epilepsy, cancer, and arthritis.

Environmental and health concerns

Despite the many benefits which dogs provide for humans, they also do harm. Dogs sometimes attack and bite humans. In the United States, there are about 7,000 dog attacks per year (Voelker 1986).

An even more serious problem is rabies, a viral disease that is spread by animal bites. Rabies is always fatal unless treated. The World Health Organization estimates that about 55,000 people die of rabies each year; mostly in the rural areas of Africa and Asia, and mostly because of dog bites. In Europe and North America, where most dogs are immunized against rabies, wild animal bites are the most common cause. About 10 million people each year are treated for possible exposure to rabies (WHO 2006). Rabies is an especially serious problem in India, which has about 25 million feral dogs (Sandeep 2002).

Dogs can also have a negative effect on the natural environment. In New Zealand, pet dogs allowed to run wild were found to be a serious threat to kiwis and other ground nesting birds (ISSG 2007). A study in the city of Baltimore in the United States done in the 1970s found that about half the dogs in the city were at least partly free ranging and that they also feed on ground nesting birds, although their main food source was garbage (Nowak 1983). In Africa, diseases spread by domestic dogs have contributed to the decline of endangered wild predators, such as the lion, the leopard, and the African hunting dog (Kruuk 2002).

Dogs in culture and religion

Hercules capturing Cerberus, Engraving by Sebald Beham 1540
Statue of St Roch with his dog, in Prague, Czech Republic

Dogs have often appeared in art since the time of ancient Egypt, where they were often included in hunting scenes painted within the tombs of important people. In Renaissance Europe, portraits often included a favorite lap dog.

Compared to some other animals, dogs have not played such an important role in religion. In Greek mythology, the underworld was guarded by a three-headed dog named Cerberus, who was later captured by Hercules. In Judaism, the dog is considered an unclean animal and when they are mentioned in the Bible they are almost always portrayed in an unfavorable light. Islam also looks on dogs unfavorably. Mohammad expressed that the company of dogs voids a portion of a Muslim’s good deeds (Malik Ibn Anas 2:969).

In Christianity, Jesus told the story of the poor man Lazarus, whose sores were licked by street dogs. This has traditionally been seen as showing Lazarus's wretched situation. However, some modern commentators have pointed out that the dogs' saliva could have beneficial effects on the sores and that this could be seen as an example of the dogs' unconditional love (Kilcommons 1996).

The Catholic Church recognizes Saint Roch (also called Saint Rocco), who lived in the early 1300s in France, as the patron saint of dogs. It is said that he caught the black plague while doing charitable work and went into the forest expecting to die. There he was befriended by a dog which licked his sores and brought him food and he was able to recover. August 16 is celebrated as the feast day of Saint Roch (Cultural Catholic 2007).

Dogs have often been mentioned favorably in literature. In Homer's Odyssey (written around 800 to 600 B.C.E.) the hero's dog Argos is the only one to recognize him on his return home in disguise. One of the greatest tributes to a dog was by the English poet Byron (1788-1824), who wrote of the Newfoundland, Boatswain (Byron 1808):

"…Beauty without Vanity,
Strength without Insolence,
Courage without Ferocity,
and all the Virtues of Man,
without his Vices."

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