Difference between revisions of "Primate" - New World Encyclopedia

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{{Taxobox_ordo_entry | taxon = '''Primates'''}}<br>{{Taxobox_authority | author = [[Carolus Linnaeus|Linnaeus]] | date = [[1758]]}}
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{{Taxobox_ordo_entry | taxon = '''Primates'''}}<br>{{Taxobox_authority | author = [[Carolus Linnaeus|Linnaeus]] | date = 1758}}
 
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{{Taxobox_end}}
 
{{Taxobox_end}}
  
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A '''primate''' (L. ''prima'', first) is any member of the biological '''Primates''', the group that contains all the species commonly related to the [[lemur]]s, [[monkey]]s, and [[ape]]s, with the latter category including [[human]]s.   
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A '''primate''' (L. ''prima,'' first) is any [[mammal]] of the biological order '''Primates,''' the group that contains all the species commonly related to the [[lemur]]s, [[monkey]]s, and [[ape]]s, with the latter category including [[human]]s.  Primates are characterized by being anatomically unspecialized, with limbs capable of performing a variety of functions, refined five-digit hands and feet adapted for grasping (including opposable thumbs), comparatively flattened snouts, and prolonged pre- and postnatal development, among other features.
  
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The Primates order is divided informally into three main groupings: ''prosimians'', ''monkeys of the New World'', and ''monkeys and apes of the Old World''. The prosimians are species whose bodies most closely resemble that of the early proto-primates. The most well known of the prosimians, the [[lemur]]s, are located on the island of [[Madagascar]] and to a lesser extent on the Comoros Islands, isolated from the rest of the world. The New World monkeys include the familiar [[capuchin monkey|capuchin]], [[howler monkey|howler]], and [[squirrel monkey]]s. They live exclusively in the Americas. Discounting humans, the rest of the [[simian]]s (monkeys and apes), the Old World monkeys and the apes, inhabit Africa and southern and central Asia, although fossil evidence shows many species existed in [[Europe]] as well.
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The Primates order is divided informally into three main groupings: [[Prosimian]]s, monkeys of the New World, and monkeys and apes of the Old World. The prosimians are species whose bodies most closely resemble that of the early proto-primates. The most well known of the prosimians, the [[lemur]]s, are located on the island of [[Madagascar]] and, to a lesser extent, on the Comoros Islands, isolated from the rest of the world. The New World monkeys include the familiar [[capuchin monkey|capuchin]], [[howler monkey|howler]], and [[squirrel monkey]]s. They live exclusively in the Americas. Discounting humans, the rest of the simians (monkeys and apes), the Old World monkeys, and the apes, inhabit Africa and southern and central Asia, although fossil evidence shows many species once existed in [[Europe]] as well.
  
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Primates are found all over the world. Non-human primates occur mostly in [[Central America|Central]] and [[South America]], [[Africa]], and southern [[Asia]]. A few species exist as far north in the Americas as southern [[Mexico]], and as far north in Asia as northern [[Japan]].
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Based on purely biological aspects ([[DNA]], proteins, morphology, etc.), it is clear the humans are primates. Indeed, humans and chimpanzees share more than 98 percent identity by various molecular comparisons ([[protein]] sequences, allele differences, DNA [[nucleotide]] sequences) (Wood 2006). However, humans are also defined in cultural, spiritual, psychological, and behavioral terms that make them qualitatively very different from other primates. Despite this, effort is sometimes made to consider humans "just primates," to the extent that efforts are actually being made to legally define other primates as "persons."
  
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The English singular ''primate'' is a back-formation from the [[Latin]] name ''Primates'', which itself was the plural of the Latin ''primas'' ("one of the first, excellent, noble").
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Primates are found all over the world. Non-human primates occur mostly in [[Central America|Central]] and [[South America]], [[Africa]], and southern [[Asia]]. A few species exist as far north in the Americas as southern [[Mexico]], and as far north in Asia as northern [[Japan]]. The earliest [[fossil]]s of primates date to the late [[Cretaceous]] period (Mayr 2001).
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The English singular, ''primate,'' is a back-formation from the [[Latin]] name ''Primates,'' which itself was the plural of the Latin ''primas'' ("one of the first, excellent, noble").
  
 
==Physical description==
 
==Physical description==
  
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All primates have five fingers (pentadactyly), a generalized dental pattern, and a primitive (unspecialized) body plan. Another distinguishing feature of primates is fingernails. Opposing thumbs are also a characteristic primate feature, but are not limited to this order; [[opossum]]s, for example, also have opposing thumbs. In primates, the combination of opposing thumbs, short fingernails (rather than claws), and long, inward-closingfingers is considered a relic of the ancestral practice of brachiating through trees. Forward-facing color binocular vision was also useful for the brachiating ancestors, particularly for finding and collecting food, although recent studies suggest it was more useful in courtship. All primates, even those that lack the features typical of other primates (like lorises), share eye orbit characteristics, such as a postorbital bar, that distinguish them from other taxonomic orders.
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All primates have five fingers (pentadactyl), a generalized dental pattern, and an unspecialized body plan. Another distinguishing feature of primates is fingernails. Opposing thumbs are also a characteristic primate feature, but are not limited to this order; [[opossum]]s, for example, also have opposing thumbs. In primates, the combination of opposing thumbs, short fingernails (rather than claws), and long, inward-closing fingers is considered a relic of the ancestral practice of brachiating through trees. Forward-facing color binocular [[eye|vision]] would also have been useful for the brachiating ancestors, particularly for finding and collecting food, although some studies suggest it was more useful in courtship. All primates, even those that lack the features typical of other primates (like lorises), share eye orbit characteristics, such as a postorbital bar, that distinguish them from other taxonomic orders.
  
 
Old World species tend to have significant sexual dimorphism. This is characterized most in size difference, with males being up to a bit more than twice as heavy as females. New World species generally form pair bonds and these species (including [[tamarin]]s and [[marmoset]]s) generally do not show a significant size difference between the sexes.
 
Old World species tend to have significant sexual dimorphism. This is characterized most in size difference, with males being up to a bit more than twice as heavy as females. New World species generally form pair bonds and these species (including [[tamarin]]s and [[marmoset]]s) generally do not show a significant size difference between the sexes.
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==Habitat==
 
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Many modern species of primates live mostly in trees and hardly ever come to the ground. Other species are partially terrestrial, such as [[baboon]]s and the Patas Monkey. Only a few species are fully terrestrial, such as the Gelada and Gorilla. 
 
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Primates live in a diverse number of forested habitats, including [[rainforest]]s, mangrove forests, and mountain forests to altitudes of over 3000 m. Although most species are generally shy of water, a few are fine swimmers and are comfortable in swamps and watery areas, including the Proboscis Monkey, De Brazza's Monkey, and Allen's Swamp Monkey, which even has small webbing between its fingers. Some primates, such the Rhesus Macaque and the Hanuman Langur, are hemerophile species and cities and villages have become their typical habitat.<ref name="German">http://de.wikipedia.org/wiki/Primaten</ref>
 
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As the table below illustrates, in many primate species, the males are larger than the females. However this picture is incomplete. All but one of these are [[Catarrhini|Old World]] species, and in this group the [[mating system]] is usually [[polygyny|polygynous]]; [[sexual dimorphism]] is expected with this kind of social structure. As the table shows, sexual dimorphism is much less in the marmosets ([[New World monkey|New World]]) than in the other species listed, and this is characteristic of marmosets and tamarins in comparison with the Old World monkeys and [[ape]]s. This is because marmosets and tamarins generally form [[monogamy|pair bond]]s.
 
  
 
{| border="1" cellpadding="3" style="border-collapse:collapse; margin:0 auto;"
 
{| border="1" cellpadding="3" style="border-collapse:collapse; margin:0 auto;"
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|78.4 kg (172 lb)
 
|78.4 kg (172 lb)
 
|-
 
|-
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|[[Patas Monkey]]
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|Patas Monkey
 
|5.5 kg (12 lb)
 
|5.5 kg (12 lb)
 
|10 kg (22 lb)
 
|10 kg (22 lb)
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|19 kg (42 lb)
 
|19 kg (42 lb)
 
|-
 
|-
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|[[Pygmy Marmoset]]
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|Pygmy Marmoset
 
|120 g (4.2 oz)
 
|120 g (4.2 oz)
 
|140 g (5 oz)
 
|140 g (5 oz)
 
|}
 
|}
  
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==Prosimians==
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==Habitat==
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'''Prosimians''' are generally considered the most primitive extant primates, representing forms that were ancestral to [[monkey]]s and [[ape]]s. With the exception of the tarsiers, all of the prosimians are in the suborder Strepsirrhini. These include the lemurs, Aye-aye, and lorises. The tasiers are placed in the suborder Haplorrhini (with the monkeys and apes). Due to this reason the classification is not considered valid in terms of phylogeny, as they do not share a unique last common ancestor, and anatomical traits.
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Many modern species of primates live mostly in trees and hardly ever come to the ground. Other species are partially terrestrial, such as [[baboon]]s and the Patas Monkey. Only a few species are fully terrestrial, such as the Gelada and [[Gorilla]].
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==New World monkeys==
 
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The '''New World monkeys''' are the four families of primates that are found in Central and South America: the ''Cebidae'' (marmosets, tamarins, capuchins, and squirrel monkeys), ''Aotidae'' (night or owl monkeys), ''Pitheciidae'' (titis, sakis, and uakaris), and ''Atelidae'' (howler, spider, and woolly monkeys). The four families are ranked together as the '''Platyrrhini''' parvorder, placing them in a different grouping from the Old World monkeys and the apes.
 
  
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All New World monkeys differ slightly from Old World monkeys in many aspects, but the most prominent of which is the nose. This is the feature used most commonly to distinguish between the two groups. The scientific name for New world monkey, Platyrrhini, means "flat nosed," therefore the noses are flatter, with side facing nostrils, compared to the narrow noses of the Old World monkey. Most New world monkeys have long, often prehensile tails. Many are small, arboreal, and nocturnal, so our knowledge of them is less comprehensive than that of the more easily observed Old World monkeys. Unlike most Old World monkeys, many New World monkeys form monogamous pair bonds, and show substantial paternal care of young.
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Primates live in a diverse number of forested habitats, including [[rainforest]]s, mangrove forests, and mountain forests to altitudes of over 3000 m. Although most species are generally shy of water, a few are fine swimmers and are comfortable in swamps and watery areas, including the [[Proboscis Monkey]], [[De Brazza's Monkey]], and [[Allen's Swamp Monkey]], which even has small webbing between its fingers. Some primates, such the Rhesus Macaque and the Hanuman Langur, are common in cities and villages.
  
 +
==Types of primates==
  
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==Old World monkeys and apes==
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The three basic groups of primates are prosimians, New World monkeys, and Old World monkeys and apes. The [[monkey]]s (New World monkeys and Old World monkeys) and [[ape]]s are also referred to as simians.
  
 +
 +
===Prosimians===
 +
[[Image:Ring tailed lemur and twins.jpg|thumb|right|200px|The Ring-tailed Lemur (''Lemur catta'') is a prosimian of the family Lemuridae.]]
 +
'''Prosimians''' are generally considered the most primitive extant (living) primates, representing forms that were ancestral to [[monkey]]s and [[ape]]s. With the exception of the tarsiers, all of the prosimians are in the suborder ''Strepsirrhini.'' These include the lemurs, Aye-aye, and lorises. The tasiers are placed in the suborder ''Haplorrhini'' (with the monkeys and apes). Due to this reason, the classification is not considered valid in terms of phylogeny, as they do not share a unique last common ancestor or anatomical traits.
  
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===Old World monkeys===
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===New World monkeys===
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The '''Old World monkeys''' or '''Cercopithecidae''' are a group of [[primate]]s, falling in the [[superfamily]] '''Cercopithecoidea''' in the [[clade]] [[Catarrhini]].  From the point of view of superficial appearance, they are unlike [[ape]]s in that most have tails (the family name means "tailed ape"), and unlike the [[New World monkeys]] in that their tails are never [[prehensile tail|prehensile]]. Technically, the distinction of catarrhines from [[platyrrhines]] depends on the structure of the nose, and the distinction of Old World monkeys from apes depends on [[dentition]].
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The '''New World [[monkey]]s''' are the four families of primates that are found in Central and South America: The ''Cebidae'' (marmosets, tamarins, capuchins, and squirrel monkeys), ''Aotidae'' (night or owl monkeys), ''Pitheciidae'' (titis, sakis, and uakaris), and ''Atelidae'' (howler, spider, and woolly monkeys). [[Image:AtelesGeoffroyi.jpg|thumb|right|175px|Spider monkey ''A. geoffroyi'']] The four families are ranked together as the '''Platyrrhini''' parvorder, placing them in a different grouping from the Old World monkeys and the apes.
  
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Several Old World monkeys have anatomical oddities. The [[colobus]] monkeys have a stub for a thumb; the [[Proboscis Monkey]] has an extraordinary nose while the [[snub-nosed monkey]]s have almost no nose at all; the penis of the male [[Mandrill]] is colored red and the scrotum has a lilac color, while the face also has bright coloration like the genitalia and this develops in only the dominant male of a multi-male group.
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All New World monkeys differ slightly from Old World monkeys in many aspects, but the most prominent difference is the nose, which is the feature most commonly used to distinguish between the two groups. The scientific name for New world monkey, Platyrrhini, means "flat nosed," therefore the noses are flatter, with side facing nostrils, compared to the narrow noses of the Old World monkey. Most New World monkeys have long, often prehensile tails. Many are small, arboreal, and nocturnal, so knowledge of them is less comprehensive than that of the more easily observed Old World monkeys. Unlike most Old World monkeys, many New World monkeys form monogamous pair bonds and show substantial paternal care of young.
  
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The Old World monkeys are native to Africa and Asia today, but are also known from Europe in the fossil record. They include many of the most familiar species of non-human primates.
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===Old World monkeys and apes===
  
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Two [[subfamily|subfamilies]] are recognised, the [[Cercopithecinae]], which are mainly [[Africa]]n but include the diverse [[genus]] of [[macaque]]s which are [[Asia]]n and North [[Africa]]n, and the [[Colobinae]], which includes most of the Asian genera but also the African colobus monkeys.
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====Old World monkeys====
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[[Image:Monkey_batu.jpg|thumb|left|100px|Old World monkey: Crab-eating Macaque from Malaysia]]
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The '''Old World [[monkey]]s,''' or ''Cercopithecidae'' family, are placed in the superfamily ''Cercopithecoidea'' in the clade ''Catarrhini.''  From the point of view of superficial appearance, they are unlike apes in that most have tails (the family name means "tailed ape"), and unlike the New World monkeys in that their tails are never prehensile (adapted to be able to grasp and hold objects).  Technically, the distinction of catarrhines from platyrrhines (New World monkeys) depends on the structure of the nose, and the distinction of Old World monkeys from apes depends on dentition.
  
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* '''Superfamily Cercopithecoidea'''
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The Old World monkeys are native to [[Africa]] and [[Asia]] today, but are also known from [[Europe]] in the fossil record. They include many of the most familiar species of non-human primates, such as baboons, macaques, and colobus monkeys. There are about 135 known species of Old World monkeys.  
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** '''Family Cercopithecidae''': Old World monkeys  
 
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*** Subfamily [[Cercopithecinae]]
 
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**** Tribe [[Cercopithecini]]
 
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***** Genus ''[[Allen's Swamp Monkey|Allenopithecus]]''
 
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****** [[Allen's Swamp Monkey]], ''Allenopithecus nigroviridis''
 
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***** Genus ''[[Miopithecus]]''
 
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****** [[Angolan Talapoin]], ''Miopithecus talapoin''
 
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****** [[Gabon Talapoin]], ''Miopithecus ogouensis''
 
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***** Genus ''[[Patas Monkey|Erythrocebus]]''
 
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****** [[Patas Monkey]], ''Erythrocebus patas''
 
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***** Genus ''[[Chlorocebus]]''
 
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****** [[Green Monkey]], ''Chlorocebus sabaceus''
 
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****** [[Grivet]], ''Chlorocebus aethiops''
 
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****** [[Bale Mountains Vervet]], ''Chlorocebus djamdjamensis''
 
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****** [[Tantalus Monkey]], ''Chlorocebus tantalus''
 
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****** [[Vervet Monkey]], ''Chlorocebus pygerythrus''
 
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****** [[Malbrouck]], ''Chlorocebus cynosuros''
 
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***** Genus ''[[Cercopithecus]]''
 
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****** [[Dryas Monkey]] or Salongo Monkey, ''Cercopithecus dryas''
 
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****** [[Diana Monkey]], ''Cercopithecus diana''
 
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****** [[Roloway Monkey]], ''Cercopithecus roloway''
 
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****** [[Greater Spot-nosed Monkey]], ''Cercopithecus nictitans''
 
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****** [[Blue Monkey]], ''Cercopithecus mitis''
 
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****** [[Silver Monkey]], ''Cercopithecus doggetti''
 
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****** [[Golden Monkey]], ''Cercopithecus kandti''
 
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****** [[Sykes's Monkey]], ''Cercopithecus albogularis''
 
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****** [[Mona Monkey]], ''Cercopithecus mona''
 
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****** [[Campbell's Mona Monkey]], ''Cercopithecus campbelli''
 
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****** [[Lowe's Mona Monkey]], ''Cercopithecus lowei''
 
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****** [[Crested Mona Monkey]], ''Cercopithecus pogonias''
 
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****** [[Wolf's Mona Monkey]], ''Cercopithecus wolfi''
 
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****** [[Dent's Mona Monkey]], ''Cercopithecus denti''
 
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****** [[Lesser Spot-nosed Monkey]], ''Cercopithecus petaurista''
 
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****** [[White-throated Guenon]], ''Cercopithecus erythrogaster''
 
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****** [[Sclater's Guenon]], ''Cercopithecus sclateri''
 
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****** [[Red-eared Guenon]], ''Cercopithecus erythrotis''
 
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****** [[Moustached Guenon]], ''Cercopithecus cephus''
 
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****** [[Red-tailed Monkey]], ''Cercopithecus ascanius''
 
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****** [[L'Hoest's Monkey]], ''Cercopithecus lhoesti''
 
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****** [[Preuss's Monkey]], ''Cercopithecus preussi''
 
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****** [[Sun-tailed Monkey]], ''Cercopithecus solatus''
 
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****** [[Hamlyn's Monkey]] or Owl-faced Monkey, ''Cercopithecus hamlyni''
 
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****** [[De Brazza's Monkey]], ''Cercopithecus neglectus''
 
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**** Tribe [[Papionini]]
 
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***** Genus ''[[Macaca]]''
 
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****** [[Barbary Macaque]], ''Macaca sylvanus''
 
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****** [[Lion-tailed Macaque]], ''Macaca silenus''
 
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****** [[Southern Pig-tailed Macaque]] or Beruk, ''Macaca nemestrina''
 
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****** [[Northern Pig-tailed Macaque]], ''Macaca leonina''
 
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****** [[Pagai Island Macaque]] or Bokkoi, ''Macaca pagensis''
 
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****** [[Siberut Macaque]], ''Macaca siberu''
 
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****** [[Moor Macaque]], ''Macaca maura''
 
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****** [[Booted Macaque]], ''Macaca ochreata''
 
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****** [[Tonkean Macaque]], ''Macaca tonkeana''
 
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****** [[Heck's Macaque]], ''Macaca hecki''
 
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****** [[Gorontalo Macaque]], ''Macaca nigriscens''
 
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****** [[Celebes Crested Macaque]] or Black "Ape", ''Macaca nigra''
 
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****** [[Crab-eating Macaque]] or Long-tailed Macaque or Kera, ''Macaca fascicularis''
 
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****** [[Stump-tailed Macaque]] or Bear Macaque, ''Macaca arctoides''
 
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****** [[Rhesus Macaque]], ''Macaca mulatta''
 
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****** [[Formosan Rock Macaque]], ''Macaca cyclopis''
 
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****** [[Japanese Macaque]], ''Macaca fuscata''
 
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****** [[Toque Macaque]], ''Macaca sinica''
 
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****** [[Bonnet Macaque]], ''Macaca radiata''
 
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****** [[Assam Macaque]], ''Macaca assamensis''
 
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****** [[Tibetan Macaque]] or Milne-Edwards' Macaque, ''Macaca thibetana''
 
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****** [[Arunachal Macaque]] or Munzala, ''Macaca munzala''
 
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***** Genus ''[[Lophocebus]]''
 
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****** [[Grey-cheeked Mangabey|Gray-cheeked Mangabey]], ''Lophocebus albigena''
 
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******* ''Lophocebus albigena albigena''
 
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******* ''Lophocebus albigena osmani''
 
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******* ''Lophocebus albigena johnstoni''
 
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****** [[Black Crested Mangabey]], ''Lophocebus aterrimus''
 
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****** [[Opdenbosch's Mangabey]], ''Lophocebus opdenboschi''
 
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***** Genus ''[[Rungwecebus]]''
 
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****** [[Kipunji]], ''Rungwecebus kipunji''
 
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***** Genus ''[[Papio]]''
 
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****** [[Hamadryas Baboon]], ''Papio hamadryas''
 
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****** [[Guinea Baboon]], ''Papio papio''
 
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****** [[Olive Baboon]], ''Papio anubis''
 
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****** [[Yellow Baboon]], ''Papio cynocephalus''
 
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****** [[Chacma Baboon]], ''Papio ursinus''
 
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***** Genus ''[[Theropithecus]]''
 
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****** [[Gelada]], ''Theropithecus gelada''
 
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***** Genus ''[[Cercocebus]]''
 
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****** [[Sooty Mangabey]], ''Cercocebus atys''
 
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******* ''Cercocebus atys atys''
 
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******* ''Cercocebus atys lunulatus''
 
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****** [[Collared Mangabey]], ''Cercocebus torquatus''
 
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****** [[Agile Mangabey]], ''Cercocebus agilis''
 
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****** [[Golden-bellied Mangabey]], ''Cercocebus chrysogaster''
 
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****** [[Tana River Mangabey]], ''Cercocebus galeritus''
 
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****** [[Sanje Mangabey]], ''Cercocebus sanjei''
 
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***** Genus ''[[Mandrillus]]''
 
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****** [[Mandrill]], ''Mandrillus sphinx''
 
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****** [[Drill (mammal)|Drill]], ''Mandrillus leucophaeus''
 
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*** Subfamily [[Colobinae]]
 
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**** African group
 
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***** Genus ''[[Colobus]]''
 
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****** [[Black Colobus]], ''Colobus satanas''
 
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****** [[Angola Colobus]], ''Colobus angolensis''
 
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****** [[King Colobus]], ''Colobus polykomos''
 
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****** [[Ursine Colobus]], ''Colobus vellerosus''
 
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****** [[Mantled Guereza]], ''Colobus guereza''
 
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***** Genus ''[[Piliocolobus]]''
 
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****** [[Western Red Colobus]], ''Piliocolobus badius''
 
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******* [[Bay Red Colobus]], ''Piliocolobus badius badius''
 
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******* [[Temminck's Red Colobus]], ''Piliocolobus badius temminckii''
 
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******* [[Miss Waldrone's Red Colobus]], ''Piliocolobus badius waldronae''
 
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****** [[Pennant's Colobus]], ''Piliocolobus pennantii''
 
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******* [[Bioko Red Colobus]], ''Piliocolobus pennantii pennantii''
 
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******* [[Niger Delta Red Colobus]], ''Piliocolobus pennantii epieni''
 
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******* [[Bouvier's Red Colobus]], ''Piliocolobus pennantii bouvieri''
 
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****** [[Preuss's Red Colobus]], ''Piliocolobus preussi''
 
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****** [[Thollon's Red Colobus]], ''Piliocolobus tholloni''
 
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****** [[Central African Red Colobus]], ''Piliocolobus foai''
 
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******* [[Foa Red Colobus]], ''Piliocolobus foai foai''
 
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******* [[Elliot's Red Colobus]], ''Piliocolobus foai ellioti''
 
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******* ''Piliocolobus foai oustaleti''
 
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******* ''Piliocolobus foai semlikiensis''
 
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******* [[Lomami Red Colobus]], ''Piliocolobus foai parmentierorum''
 
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****** [[Ugandan Red Colobus]], ''Piliocolobus tephrosceles''
 
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****** [[Uzyngwa Red Colobus]], ''Piliocolobus gordonorum''
 
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****** [[Zanzibar Red Colobus]], ''Piliocolobus kirkii''
 
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****** [[Tana River Red Colobus]], ''Piliocolobus rufomitratus''
 
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***** Genus ''[[Procolobus]]''
 
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****** [[Olive Colobus]], ''Procolobus verus''
 
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**** Langur (leaf monkey) group
 
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***** Genus ''[[Gray langur|Semnopithecus]]''
 
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****** [[Nepal Gray Langur]], ''Semnopithecus schistaceus''
 
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****** [[Kashmir Gray Langur]], ''Semnopithecus ajax''
 
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****** [[Tarai Gray Langur]], ''Semnopithecus hector''
 
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****** [[Northern Plains Gray Langur]], ''Semnopithecus entellus''
 
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****** [[Black-footed Gray Langur]], ''Semnopithecus hypoleucos''
 
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****** [[Southern Plains Gray Langur]], ''Semnopithecus dussumieri''
 
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****** [[Tufted Gray Langur]], ''Semnopithecus priam''
 
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***** Genus ''[[Trachypithecus]]''
 
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****** ''T. vetulus'' group
 
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******* [[Purple-faced Langur]], ''Trachypithecus vetulus''
 
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******* [[Nilgiri Langur]], ''Trachypithecus johnii''
 
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****** ''T. cristatus'' group
 
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******* [[Javan Lutung]], ''Trachypithecus auratus''
 
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******* [[Silvery Lutung]] or Silvered Leaf Monkey, ''Trachypithecus cristatus''
 
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******* [[Indochinese Lutung]], ''Trachypithecus germaini''
 
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******* [[Tenasserim Lutung]], ''Trachypithecus barbei''
 
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****** ''T. obscurus'' group
 
−
******* [[Dusky Leaf Monkey]] or Spectacled Leaf Monkey, ''Trachypithecus obscurus''
 
−
******* [[Phayre's Leaf Monkey]], ''Trachypithecus phayrei''
 
−
****** ''T. pileatus'' group
 
−
******* [[Capped Langur]], ''Trachypithecus pileatus''
 
−
******* [[Shortridge's Langur]], ''Trachypithecus shortridgei''
 
−
******* [[Gee's Golden Langur]], ''Trachypithecus geei''
 
−
****** ''T. francoisi'' group
 
−
******* [[Francois' Langur]], ''Trachypithecus francoisi''
 
−
******* [[Hatinh Langur]], ''Trachypithecus hatinhensis''
 
−
******* [[White-headed Langur]], ''Trachypithecus poliocephalus''
 
−
******* [[Laotian Langur]], ''Trachypithecus laotum''
 
−
******* [[Delacour's Langur]], ''Trachypithecus delacouri''
 
−
******* [[Indochinese Black Langur]], ''Trachypithecus ebenus''
 
−
***** Genus ''[[Presbytis]]''
 
−
****** [[Sumatran Surili]], ''Presbytis melalophos''
 
−
****** [[Banded Surili]], ''Presbytis femoralis''
 
−
****** [[Sarawak Surili]], ''Presbytis chrysomelas''
 
−
****** [[White-thighed Surili]], ''Presbytis siamensis''
 
−
****** [[White-fronted Surili]], ''Presbytis frontata''
 
−
****** [[Javan Surili]], ''Presbytis comata''
 
−
****** [[Thomas's Langur]], ''Presbytis thomasi''
 
−
****** [[Hose's Langur]], ''Presbytis hosei''
 
−
****** [[Maroon Leaf Monkey]], ''Presbytis rubicunda''
 
−
****** [[Mentawai Langur]] or Joja, ''Presbytis potenziani''
 
−
****** [[Natuna Island Surili]], ''Presbytis natunae''
 
−
**** Odd-Nosed group
 
−
***** Genus ''[[Pygathrix]]''
 
−
****** [[Red-shanked Douc]], ''Pygathrix nemaeus''
 
−
****** [[Black-shanked Douc]], ''Pygathrix nigripes''
 
−
****** [[Gray-shanked Douc]], ''Pygathrix cinerea''
 
−
***** Genus ''[[Rhinopithecus]]''
 
−
****** [[Golden Snub-nosed Monkey]], ''Rhinopithecus roxellana''
 
−
****** [[Black Snub-nosed Monkey]], ''Rhinopithecus bieti''
 
−
****** [[Gray Snub-nosed Monkey]], ''Rhinopithecus brelichi''
 
−
****** [[Tonkin Snub-nosed Langur]], ''Rhinopithecus avunculus''
 
−
***** Genus ''[[Proboscis Monkey|Nasalis]]''
 
−
****** [[Proboscis Monkey]], ''Nasalis larvatus''
 
−
***** Genus ''[[Pig-tailed Langur|Simias]]''
 
−
****** [[Pig-tailed Langur]], ''Simias concolor''
 
  
 +
Two subfamilies of Old World monkeys are recognized, the ''Cercopithecinae,'' which are mainly African, but include the diverse genus of [[macaque]]s, which are Asian and North African, and the ''Colobinae,'' which includes most of the Asian genera, as well as the African [[colobus]] monkeys.
  
 +
====Apes====
 +
[[Image:Weisshandgibbon tierpark berlin.jpg|thumb|right|150px|Ape: Lar Gibbon (Hylobatidae family)]]
  
 +
'''[[Ape]]s''' are the members of the ''Hominoidea'' superfamily of primates. Under the [[taxonomy|taxonomic]] scheme that is currently common, there are two families of '''hominoids:'''
  
−
===Apes===
+
*The family ''Hylobatidae'' consists of 4 genera and 12 species of [[gibbon]]s, collectively known as the "lesser apes";
−
'''Apes''' are the members of the '''Hominoidea''' [[superfamily]] of [[primates]], which includes [[human]]s. Under current classification, there are two [[family (biology)|families]] of '''hominoids''':
+
*The family ''Hominidae,'' consisting of [[gorilla]]s, [[chimpanzee]]s, [[orangutan]]s, and [[human]]s, is collectively known as the "great apes."
  
−
*the family [[Hylobatidae]] consists of 4 genera and 12 species of [[gibbon]]s, including the [[Lar Gibbon]] and the [[Siamang]], collectively known as the "lesser apes"
+
Awkwardly, the term "hominid" is still used with the specific meaning of extinct animals more closely related to humans than the other great apes (for example, [[Australopithecus|Australopithecines]]).  
−
*the family [[Hominidae]] consisting of [[gorilla]]s, [[chimpanzee]]s, [[orangutan]]s, and humans, collectively known as the "great apes".  
 
  
−
A few other primates have the word "ape" in their common names, but they are not regarded as true apes.  
+
In another classification, historically common, members of the Hominoidea are divided into three families, the Hylobatidae (gibbons), the ''Pongidae'' (orangutans, gorillas, chimpanzees, and bonobos), and the Hominidae (humans). In this case, the term hominid accurately applies to all human and humanlike forms, while great apes applies only to gorillas, chimpanzees, and orangutans.
  
−
Except for gorillas and humans, all true apes are agile climbers of trees. They are best described as [[omnivorous]], their diet consisting of fruit, grass seeds, and in most cases some quantities of meat and invertebrates&mdash;either hunted or scavenged&mdash;along with anything else available and easily digested. They are native to [[Africa]] and [[Asia]], although humans have spread to all parts of the world.
+
Some researchers place gorillas and chimpanzees (and the related bonobos) into the ''Panidae'' family, while orangutans remain in the Pongidae family, and humans in the Hominidae. The bonobos are sometimes considered a variety of chimpanzees and are referred to as pygmy chimpanzees.
  
−
Most ape species are rare or [[endangered species|endangered]]. The chief threat to most of the endangered species is loss of tropical rainforest habitat, though some populations are further imperiled by hunting for [[bushmeat]].
+
The eight extant (living) genera of hominoids are the four great ape genera (''Homo'' (humans), ''[[chimpanzee|Pan]]'' (chimpanzees), ''[[Gorilla]]'', and ''[[Pongo]]'' (orangutans)), and the four genera of [[gibbon]]s (''Hylobates,'' ''Hoolock,'' ''Nomascus,'' and ''Symphalangus''). The genus for the hoolock gibbons was changed from ''Bunopithecus'' to ''Hoolock'' (Mootnick and Groves 2005).
  
−
==Historical and modern terminology==
+
A few other primates have the word "ape" in their common names, but they are not regarded as true apes.  
−
"Ape" (Old Eng. ''apa''; Dutch ''aap''; Old Ger. ''affo''; Welsh ''epa''; Old Czech ''op'') is a word of uncertain origin and is possibly an [[Onomatopoeia|onomatopoetic]] imitation of animal chatter. The term has a history of rather imprecise usage. Its earliest meaning was a tailless (and therefore exceptionally human-like) non-human primate, but as [[zoology|zoological]] knowledge developed it became clear that taillessness occurred in a number of different and otherwise unrelated [[species]].  
 
  
−
The original usage of "ape" in [[English language|English]] may have referred to the [[baboon]], an [[Africa]]n [[Old World monkey|monkey]]. Two tailless species of [[macaque]] are commonly named as apes, the [[Barbary Ape]] of North Africa (introduced into [[Gibraltar]]), ''Macaca sylvanus'', and the Sulawesi Black Ape or [[Celebes Crested Macaque]], ''M. nigra''.
+
Both great apes and lesser apes fall within parvorder ''Catarrhini,'' which also includes the Old World monkeys of [[Africa]] and [[Eurasia]]. Within this group, both families of apes can be distinguished from these [[monkey]]s by the number of cusps on their molars (apes have five&mdash;the "Y-5" molar pattern, Old World monkeys have only four in a "bilophodont" pattern). Apes have more mobile shoulder joints and arms, ribcages that are flatter front-to-back, and a shorter, less mobile spine compared to Old World monkeys. These are all anatomical adaptations to vertical hanging and swinging locomotion (brachiation) in the apes. All living members of the Hylobatidae and Hominidae are tailless, and thus humans have been referred to as tailless, bipedal, primates. However there are also primates in other families that lack tails, and at least one (the Pig-Tailed Langur) that has been known to walk significant distances bipedally.
  
−
Until a handful of decades ago, humans were thought to be distinctly set apart from the other apes (even from the other great apes), so much so that many people still don't think of the term "apes" to include humans at all. However, it is not considered accurate by many biologists to think of apes in a biological sense without considering humans to be included. The terms "non-human apes" or "non-human great apes" is used with increasing frequency to show the relationship of humans to the other apes while yet talking only about the non-human species.
+
Except for gorillas and humans, all true apes are agile climbers of trees. They are best described as omnivorous, their diet consisting of fruit, grass seeds, and in most cases some quantities of meat and invertebrates&mdash;either hunted or scavenged&mdash;along with anything else available and easily digested. The apes are native to [[Africa]] and [[Asia]], although humans have spread to all parts of the world.
  
−
A group of apes may be referred to as a ''troop'' or a ''shrewdness''.
+
Most ape species are rare or [[endangered species|endangered]]. The chief threat to most of the endangered species is loss of tropical [[rainforest]] habitat, though some populations are further imperiled by hunting.
  
−
==Biology==
+
Many people do not consider the term "apes" to include humans, although [[biology|biologists]] generally do consider humans as apes. The terms "non-human apes" or "non-human great apes" is used with increasing frequency to both show the relationship of humans to the other species classified in the families ''Hylobatidae'' and ''Hominidae,'' while maintaining the distinction of humans.
  
−
<!--[[Image:ChimpHands.jpg|250px|thumb|Shot of a chimpanzee's forelimbs.]]—>
+
== Classification==
  
−
The gibbon family, Hylobatidae, is composed of thirteen medium-sized species. Their major distinction are their long arms which they use to [[brachiation|brachiate]] through the trees. As an evolutionary adaption to this [[arboreal]] lifestyle, their wrists are ball and socket joints. The largest of the gibbons, the [[Siamang]], weighs up to 23 kg (50 lb). In comparison, the smallest great ape is the [[Common Chimpanzee]] at a modest 40 to 65 kg (88 to 143 lb).
+
The Primate order lies in a tight clustering of related orders (the Euarchontoglires) within the Eutheria, a subclass of [[Mammal|Mammalia]]. Recent molecular genetic research on primates, flying [[lemur]]s, and treeshrews has shown that the two species of flying lemur (Dermoptera) are more closely related to the primates than the treeshrews of the order Scandentia, even though the treeshrews were at one time considered primates. These three orders make up the Euarchonta clade. This clade combines with the Glires clade (made up of the Rodentia, rodents, and Lagomorpha, rabbits) to form the Euarchontoglires clade. Variously, both Euarchonta and Euarchontoglires are ranked as superorders. Also, some scientists consider Dermoptera a suborder of Primates and call the "true" primates the suborder Euprimates.
  
−
The great ape family was previously referred to as [[Pongidae]], and humans (and fossil hominids) were omitted from it, but on grounds of relatedness there is no argument for doing this. Chimpanzees, gorillas, humans and orangutans are all more closely related to one another than any of these four genera are to the gibbons. Awkwardly, however, the term "hominid" is still used with the specific meaning of extinct animals more closely related to humans than the other great apes (for example, [[australopithecine]]s). It is now usual to use even finer divisions, such as [[subfamily (biology)|subfamilies]] and [[tribe (biology)|tribe]]s to distinguish which hominoids are being discussed. Current evidence implies that humans share a common, extinct, ancestor with the chimpanzee line, from which we separated more recently than the gorilla line.
+
  Euarchontoglires
−
 
+
    ├─Glires
−
Both great apes and lesser apes fall within [[Catarrhini]], which also includes the [[Old World monkeys]] of [[Africa]] and [[Eurasia]]. Within this group, both families of apes can be distinguished from these [[monkey]]s by the number of cusps on their [[molar (tooth)|molar]]s (apes have five&mdash;the "Y-5" molar pattern, Old World monkeys have only four in a "bilophodont" pattern). Apes have more mobile shoulder joints and arms, ribcages that are flatter front-to-back, and a shorter, less mobile spine compared to Old World monkeys. These are all anatomical adaptations to vertical hanging and swinging locomotion (brachiation) in the apes. All living members of the Hylobatidae and Hominidae are tailless, and humans can therefore accurately be referred to as bipedal apes. However there are also primates in other families that lack tails, and at least one (the [[Pig-Tailed Langur]]) that has been known to walk significant distances bipedally.
+
    │    ├─rodents (Rodentia)
−
 
+
    │    └─rabbits, hares, pikas (Lagomorpha)
−
Although the hominoid fossil record is far from complete, and the evidence is often fragmentary, there is enough to give a good outline of the evolutionary history of humans. The time of the split between humans and living apes used to be thought to have occurred 15 to 20 million years ago, or even up to 30 or 40 million years ago. Some apes occurring within that time period, such as ''[[Ramapithecus]]'', used to be considered as [[hominin]]s, and possible ancestors of humans. Later fossil finds indicated that ''Ramapithecus'' was more closely related to the orangutan, and new biochemical evidence indicated that the last common ancestor of humans and other hominins occurred between 5 and 10 million years ago, and probably in the lower end of that range.
+
    └─Euarchonta
−
 
+
          ├─treeshrews (Scandentia)
−
==Cultural aspects of non-human apes==
+
          └─N.N.
−
The intelligence and humanoid appearance of non-human apes are responsible for legends which attribute human qualities; for example, they are sometimes said to be able to speak but refuse to do so in order to avoid work. They are also said to be the result of a curse&mdash;a [[Jewish]] [[folktale]] claims that one of the races who built the [[Tower of Babel]] became non-human apes as punishment, while [[Muslim]] lore says that the Jews of [[Elath]] became non-human apes as punishment for [[fishing]] on the [[Sabbath]]. [[Christianity|Christian]] folklore claims that non-human apes are a symbol of [[lust]] and were created by [[Satan]] in response to [[God]]'s creation of humans. It is uncertain whether any of these references is to any specific non-human apes, since all date from a period when the distinction between non-human apes and monkeys was not widely understood, or not understood at all.
+
              ├─flying lemurs (Dermoptera
−
 
+
              └─N.N.
−
Humans and the other apes share many similarities, including the ability to properly use tools and imitate others. Recent studies at [[Yale]] test some of these similarities. A professor and his/her students gave a challenge to baby humans and baby chimps. Both groups were shown a way that might solve the challenge. However what both the baby humans and chimps did not know was that the way that was shown was incorrect. Both times the baby humans tried they imitated what they were shown and failed at the attempt. For the chimps, the first time they tried, they followed the same path and failed. However on a second time around they succeeded in finding a new path and actually completed the objective. The professor interpreted that baby chimps learned from experience while baby humans just imitated what they were shown. This gave scientists key information in understanding the cultural aspects of ape life and evolutionary similarities between humans and apes.
+
                    ├─Plesiadapiformes (extinct)
−
 
+
                    └─primates (Primates)
−
There have also been recent breakthroughs in evidence of ape culture that go beyond what was explained above. This was further explored by scientists at the convention in St. Louis.<ref>{{cite web | url = http://livescience.com/animalworld/060228_ape_culture.html | publisher = Livescience.com | year = [[2006-02-28]] | title = Case Closed: Apes Got Culture | author = Corey Binns}}</ref>
 
−
 
 
−
==New subspecies?==
 
−
In 2002, a new giant ape troop was discovered in the [[Democratic Republic of Congo]]. These apes share many features of both chimpanzees and gorillas. According to a report from [[BBC News Online]],<ref>{{cite web | url = http://news.bbc.co.uk/1/hi/sci/tech/3730574.stm | title = 'New' giant ape found in DR Congo | publisher = [[BBC News Online]] | year = [[2004-10-10]]}}</ref>, the apes have large black faces, are two meters (6.6 feet) tall and make nests on the ground, all like gorillas. However, they live hundreds of kilometers from any other gorilla troops, and their diet is high in fruits, similar to the chimpanzee diet and are said by the locals to be active hunters, even overpowering local lions. They were given the common name of [[Bili Ape]].
 
−
 
 
−
Subsequent molecular investigation of pelt samples showed them to be [[Common Chimpanzee]]s who had individually adapted to local conditions.  This would indicate that they may be a subspecies of the common chimp, although this only definitively points to a chimp maternal ancestor, as mtDNA (mitochondreal DNA) is transferred from mother to child. This analysis would not rule out the possibility of a gorilla/chimp hybrid where the ancestral father is a gorilla.
 
−
 
 
−
==History of hominoid taxonomy==
 
−
The history of hominoid taxonomy is somewhat confusing and complex. The names of subgroups have changed their meaning over time as new evidence from fossil discoveries, anatomy comparisons and DNA sequences, has changed understanding of the relationships between hominoids. The story of the hominoid taxonomy is one of gradual demotion of humans from a special position in the taxonomy to being one branch among many. It also illustrates the growing influence of [[cladistics]] (the science of classifying living things by strict descent) on taxonomy.
 
−
 
 
−
[[As of 2006]], there are eight extant [[genus|genera]] of hominoids. They are the four great ape genera (''[[Homo (genus)|Homo]]'' (humans), ''[[chimpanzee|Pan]]'' (chimpanzees), ''[[Gorilla]]'', and ''[[Pongo]]'' (orangutans)), and the four genera of [[gibbon]]s (''[[Hylobates]]'', ''[[Hoolock]]'', ''[[Nomascus]]'', and ''[[Symphalangus]]'').<ref name=MSW3/> (The genus for the [[hoolock gibbon]]s was recently changed from ''[[Bunopithecus]]'' to ''Hoolock''.<ref name=Mootnick>{{cite journal | last = Mootnick | first = A. | coauthors = Groves, C. P. | year = 2005 | title = A new generic name for the hoolock gibbon (Hylobatidae) | journal = International Journal of Primatology | issue = 26 | pages = 971-976}}</ref>)
 
−
 
 
−
In 1758, [[Carolus Linnaeus]], relying on second- or third-hand accounts, placed a second species in ''Homo'' along with ''H. sapiens'': ''Homo troglodytes'' ("cave-dwelling man"). It is not clear to which animal this name refers, as Linnaeus had no specimen to refer to, hence no precise description. Linnaeus named the orangutan ''Simia satyrus'' ("satyr monkey"). He placed the three genera ''Homo'', ''[[Simia]]'' and ''Lemur'' in the family of Primates.
 
−
 
 
−
The ''troglodytes'' name was used for the chimpanzee by [[Johann Friedrich Blumenbach|Blumenbach]] in 1775 but moved to the genus ''Simia''. The orangutan was moved to the genus ''[[Pongo]]'' in 1799 by [[Bernard Germain Étienne de la Ville, Comte de Lacépède|Lacépède]].
 
−
 
 
−
Linnaeus's inclusion of humans in the primates with monkeys and apes was troubling for people who denied a close relationship between humans and the rest of the animal kingdom. Linneaus's Lutheran [[Archbishop]] had accused him of "impiety."  In a letter to [[Johann Georg Gmelin]] dated [[February 25]], [[1747]], Linnaeus wrote:
 
−
 
 
−
:It is not pleasing to me that I must place [[humans]] among the [[primate]]s, but man is intimately familiar with himself. Let's not quibble over words. It will be the same to me whatever name is applied. But I desperately seek from you and from the whole world a general difference between men and [[simian]]s from the principles of [[Natural History]]. I certainly know of none. If only someone might tell me one! If I called man a [[Simia|simian]] or vice versa I would bring together all the [[theologians]] against me.  Perhaps I ought to, in accordance with the law of Natural History.<ref>{{cite web | url = http://linnaeus.c18.net/Letters/display_txt.php?id_letter=L0783 | title = Letter,  Carl Linnaeus to Johann Georg Gmelin. Uppsala, Sweden, 25 February 1747 | publisher = Swedish Linnaean Society}}</ref>
 
−
 
 
−
Accordingly, [[Johann Friedrich Blumenbach]] in the first edition of his ''Manual of Natural History'' (1779), proposed that the primates be divided into the [[Quadrumana]] (four-handed, i.e. apes and monkeys) and [[Bimana]] (two-handed, ie. humans). This distinction was taken up by other naturalists, most notably [[Georges Cuvier]]. Some elevated the distinction to the level of [[order (biology)|order]].
 
−
 
 
−
However, the many affinities between humans and other primates — and especially the great apes — made it clear that the distinction made no scientific sense. [[Charles Darwin]] wrote, in ''[[The Descent of Man, and Selection in Relation to Sex|The Descent of Man]]'':
 
−
 
 
−
:The greater number of naturalists who have taken into consideration the whole structure of man, including his mental faculties, have followed [[Johann Friedrich Blumenbach|Blumenbach]] and [[Georges Cuvier|Cuvier]], and have placed man in a separate Order, under the title of the Bimana, and therefore on an equality with the orders of the Quadrumana, [[Carnivora]], etc. Recently many of our best naturalists have recurred to the view first propounded by [[Carolus Linnaeus|Linnaeus]], so remarkable for his sagacity, and have placed man in the same Order with the Quadrumana, under the title of the Primates. The justice of this conclusion will be admitted: for in the first place, we must bear in mind the comparative insignificance for classification of the great development of the brain in man, and that the strongly-marked differences between the skulls of man and the Quadrumana (lately insisted upon by [[Theodor Ludwig Wilhelm Bischoff|Bischoff]], [[Christoph Theodor Aeby|Aeby]], and others) apparently follow from their differently developed brains. In the second place, we must remember that nearly all the other and more important differences between man and the Quadrumana are manifestly adaptive in their nature, and relate chiefly to the erect position of man; such as the structure of his hand, foot, and pelvis, the curvature of his spine, and the position of his head.<ref>{{cite book | author = [[Charles Darwin]] | title = [[The Descent of Man]] | year = 1871}}</ref>
 
−
 
 
−
<!-- ...history from 1870 to 1960 goes here... —>
 
−
 
 
−
[[Image:Hominoid taxonomy 1.png|right|300px]]
 
−
Until about 1960, the hominoids were usually divided into two families: humans and their extinct relatives in [[Hominidae]], the other apes in [[Pongidae]].<ref>{{cite journal | author = [[George Gaylord Simpson|G. G. Simpson]] | year = 1945 | title = The principles of classification and a classification of mammals | journal = Bull. Am. Mus. Nat. Hist. | volume = 85 | pages = 1–350}}</ref>
 
−
 
 
−
<br style="clear:both"/>
 
−
[[Image:Hominoid taxonomy 2.png|right|300px]]
 
−
The 1960s saw the application of techniques from [[molecular biology]] to primate taxonomy. Goodman used his 1963 immunological study of serum proteins to propose a division of the hominoids into three families, with the non-human great apes in Pongidae and the lesser apes (gibbons) in [[Hylobatidae]].<ref>{{cite book | author = M. Goodman | year = 1963 | chapter = Man’s place in the phylogeny of the primates as reflected in serum proteins | editor = S. L. Washburn | title = Classification and human evolution | publisher = Aldine, Chicago | pages = 204–234}}</ref> The [[trichotomy]] of hominoid families, however, prompted scientists to ask which family [[speciation|speciated]] first from the common hominoid ancestor.
 
−
 
 
−
<br style="clear:both"/>
 
−
[[Image:Hominoid taxonomy 3.png|right|300px]]
 
−
Within the superfamily Hominoidea, gibbons are the [[outgroup]]: this means that the rest of the hominoids are more closely related to each other than any of them are to gibbons. This led to the placing of the other great apes into the family Hominidae along with humans, by demoting the Pongidae to a subfamily; the Hominidae family now contained the subfamilies [[Homininae]] and [[Ponginae]]. Again, the three-way split in Ponginae led scientists to ask which of the three genera is least related to the others.
 
−
 
 
−
<br style="clear:both"/>
 
−
[[Image:Hominoid taxonomy 4.png|right|300px]]
 
−
Investigation showed [[orangutan]]s to be the outgroup, but comparing humans to all three other hominid genera showed that African apes (chimpanzees and gorillas) and humans are more closely related to each other than any of them are to orangutans. This led to the placing of the African apes in the subfamily Homininae, forming another three-way split. This classification was first proposed by M. Goodman in 1974.<ref>{{cite journal | journal = Annual Review of Anthropology | volume = 3 | pages = 203–228 | year = 1974 | url = http://arjournals.annualreviews.org/doi/abs/10.1146/annurev.an.03.100174.001223 | title = Biochemical Evidence on Hominid Phylogeny | author = M. Goodman}}</ref>
 
−
 
 
−
<br style="clear:both"/>
 
−
[[Image:Hominoid taxonomy 5.png|right|300px]]
 
−
To try to resolve the hominine trichotomy, some authors proposed the division of the subfamily Homininae into the tribes Gorillini (African apes) and Hominini (humans).
 
−
 
 
−
<br style="clear:both"/>
 
−
[[Image:Hominoid taxonomy 6.png|right|300px]]
 
−
However, DNA comparisons provide convincing evidence that within the subfamily Homininae, gorillas are the outgroup. This suggests that chimpanzees should be in Hominini along with humans. This classification was first proposed (though one rank lower) by M. Goodman et. al. in 1990.<ref>{{cite journal | journal = Journal of Molecular Evolution | year = 1990 | volume = 30 | pages = 260–266 | url = http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=Retrieve&db=PubMed&list_uids=2109087&dopt=Abstract | title = Primate evolution at the DNA level and a classification of hominoids | author = M. Goodman, D. A. Tagle, D. H. Fitch, W. Bailey, J. Czelusniak, B. F. Koop, P. Benson, J. L. Slightom}}</ref> See [[Human evolutionary genetics]] for more information on the speciation of humans and great apes.
 
−
 
 
−
<br style="clear:both"/>
 
−
[[Image:Hominoid taxonomy 7.png|right|300px]]
 
−
Later DNA comparisons split the gibbon genus ''Hylobates'' into four genera: ''[[Hylobates]]'', ''[[Hoolock]]'', ''[[Nomascus]]'', and ''[[Symphalangus]]''.<ref name=MSW3/><ref name=Mootnick/>
 
−
 
 
−
<br clear=all>
 
−
 
 
−
==Classification and evolution==
 
−
As discussed above, hominoid taxonomy has undergone several changes. Current understanding is that the apes diverged from the [[Old World monkey]]s about 25 million years ago. The lesser and greater apes split about 18 mya, and the hominid splits happen 14 mya (''Pongo''), 7 mya (''Gorilla''), and 3-5 mya (''Homo'' & ''Pan'')
 
−
 
 
−
* '''Superfamily Hominoidea'''<ref name=MSW3/>
 
−
** Family [[Hylobatidae]]: gibbons
 
−
*** Genus ''[[Hylobates]]''
 
−
*** Genus ''[[Hoolock]]''
 
−
*** Genus ''[[Symphalangus]]''
 
−
*** Genus ''[[Nomascus]]''
 
−
** Family [[Hominidae]]: great apes
 
−
*** Genus ''[[Pongo]]'': orangutans
 
−
*** Genus ''[[Gorilla]]'': gorillas
 
−
*** Genus ''[[Homo (genus)|Homo]]'': humans
 
−
*** Genus ''[[Chimpanzee|Pan]]'': chimpanzees
 
−
 
 
−
==Legal status== 
 
−
 
 
−
Humans are the only apes recognized as [[person]]s and protected in law by the [[United Nations]] [[Universal Declaration of Human Rights]]<ref>{{cite web | url = http://www.un.org/Overview/rights.html | title = Universal Declaration of Human Rights | year = [[1948-12-10]] | publisher = United Nations}}</ref> and by all governments, though to varying degrees. The non-human apes are not classified as persons, which means that where their interests intersect with that of humans they have no legal status.
 
−
 
 
−
Many argue that the other apes' cognitive capacity in itself, as well as their close genetic relationship to human beings, dictates an acknowledgement of personhood. The [[Great Ape Project]], founded by Australian philosopher [[Peter Singer]], is campaigning to have the United Nations endorse its [[Declaration on Great Apes]], which would extend to all species of [[chimpanzee]]s, [[gorilla]]s, and [[orangutan]]s the protection of three basic interests: the right to life, the protection of individual [[liberty]], and the prohibition of [[torture]].
 
−
 
 
−
== See also ==
 
−
* [[List of fictional apes]]
 
  
−
==References==
 
−
<references/>
 
  
 +
[[Image:Ring_tailed_lemurs.jpg|200px|right|thumb|Ring-tailed Lemur, a strepsirrhine primate]]
 +
In older classifications, the Primates were divided into two superfamilies: Prosimii and Anthropoidea. The Prosimii included all of the [[prosimian]]s: All of Strepsirrhini plus the tarsiers. The Anthropoidea contained all of the [[simian]]s.
  
−
== Classification and evolution==
+
In modern cladistic reckonings, the Primate order is also a true clade. The suborder Strepsirrhini, the "wet-nosed" primates, split off from the primitive primate line about 63 million years ago (mya). The seven strepsirhine families are the four related [[lemur]] families and the three remaining families that include the lorises, the Aye-aye, the galagos, and the pottos. Some classification schemes wrap the Lepilemuridae into the Lemuridae and the Galagidae into the Lorisidae, yielding a three-two family split instead of the four-three split as presented here.  
−
===Close relations===
 
−
The Primate [[order (biology)|order]] lies in a tight clustering of related orders (the [[Euarchontoglires]]) within the [[Eutheria]], a subclass of [[Mammalia]]. Recent molecular genetic research on primates, [[flying lemur]]s, and [[treeshrew]]s has shown that the two species of flying lemur ([[Dermoptera]]) are more closely related to the primates than the treeshrews of the order [[Scandentia]], even though the treeshrews were at one time considered primates. These three orders make up the [[Euarchonta]] [[clade]]. This clade combines with the [[Glires]] clade (made up of the [[Rodentia]] and [[Lagomorpha]]) to form the Euarchontoglires clade. Variously, both Euarchonta and Euarchontoglires are ranked as superorders. Also, some scientists consider Dermoptera a suborder of Primates and call the "true" primates the suborder [[Euprimates]].{{cite needed}}
 
  
−
[[Euarchontoglires]]
+
Other lineages of lower primates once inhabited Earth. During the [[Eocene]], most of the northern continents were dominated by two dominant groups, the adapids and the omomyids. The former is considered a member of Strepsirrhini, but it does not have a tooth-comb like modern lemurs. The latter was related closely to tarsiers, monkeys, and apes. Adapids survived until 10 mya; omomyids, on the other hand, perished 20 million years earlier.
−
    ├─[[Glires]]
 
−
    │    ├─rodents ([[Rodentia]])
 
−
    │    └─rabbits, hares, pikas ([[Lagomorpha]])
 
−
    └─[[Euarchonta]]
 
−
          ├─treeshrews ([[Treeshrew|Scandentia]])
 
−
          └─N.N.
 
−
              ├─flying lemurs ([[Colugo|Dermoptera]])
 
−
              └─N.N.
 
−
                    ├─[[Plesiadapiformes]] (extinct)
 
−
                    └─primates (Primates)
 
  
−
===Classification===
+
The Aye-aye is difficult to place in Strepsirrhini. Its family, Daubentoniidae, could be a lemuriform primate and its ancestors split from lemur line more recently than the lemurs and lorises split, about 50 mya. Otherwise, it is sister to all of the other strepsirrhines, in which case it evolved away from the main strepsirrhine line between 50 and 63 mya. 
−
[[Image:Ring_tailed_lemurs.jpg|200px|right|thumb|[[Ring-tailed Lemur]], a [[strepsirrhine]] primate]]
+
[[Image:EmperorTamarin.jpg|200px|left|thumb|Emperor Tamarin, a New World monkey]]
−
In older classifications, the Primates were divided into two superfamilies: Prosimii and Anthropoidea. The Prosimii included all of the [[prosimian]]s: all of Strepsirrhini plus the tarsiers. The Anthropoidea contained all of the [[simian]]s.
 
  
−
In modern, cladistic reckonings, the Primate order is also a true clade. The suborder [[Strepsirrhini]], the "[[rhinarium|wet-nosed]]" primates, split off from the primitive primate line about 63 [[mya (unit)|million years ago (mya)]]. The seven strepsirhine families are the four related [[lemur]] families and the three remaining families that include the [[loris]]es, the [[Aye-aye]], the [[galago]]s, and the [[potto]]s.<ref name=MSW3>{{MSW3 Groves|pages=111-184}}</ref> Some classification schemes wrap the [[Lepilemuridae]] into the [[Lemuridae]] and the [[Galagidae]] into the [[Lorisidae]], yielding a three-two family split instead of the four-three split as presented here.<ref name=MSW3/> Other lineages of lower primates inhabited Earth. During the [[Eocene]], most of the northern continents were dominated by two dominant groups, the [[adapid]]s and the [[omomyid]]s. The former is considered a member of Strepsirrhini, but it does not have a [[tooth-comb]] like modern lemurs. The latter was related closely to tarsiers, monkeys, and apes. Adapids survived until 10 mya; omomyids on the other hand perished 20 million years earlier.
+
The suborder Haplorrhini, the "dry-nosed" primates, is composed of two sister clades. The prosimian tarsiers in family Tarsiidae (monotypic in its own infraorder Tarsiiformes), represent the most primitive division at about 58 mya. The Simiiformes infraorder contains the two parvorders: The New World monkeys in one, and the Old World monkeys, humans, and the other [[ape]]s in the other. This division is considered to have happened about 40 mya. However, about 30 mya, it is thought that these three groups split from the main haplorrhine lineage. One group stayed in Asia and are closest in kin to the "dawn monkey" ''Eosimias''. The second stayed in Africa, where they developed into the Old World primates. The third rafted to South America to become the New World monkeys. Mysteriously the aboriginal Asian Haplorrhini vanished from record once Africa collided with Eurasia 24 mya. Apes and monkeys spread into Europe and Asia. Close behind came lorises and tarsiers, also African castaways. The first hominid fossils were discovered in Northern Africa and date back 7 mya. Modern humans did not appear until 0.2 mya, eventually becoming the most prevalent primate and mammal on Earth.
  
−
The Aye-aye is difficult to place in Strepsirrhini.<ref name=MSW3/> Its family, Daubentoniidae, could be a lemuriform primate and its ancestors split from lemur line more recently than the lemurs and lorises split, about 50 mya. Otherwise it is sister to all of the other strepsirrhines, in which case in evolved away from the main strepsirrhine line between 50 and 63 mya. 
+
The discovery of new species happens at a rate of a few each year, and the evaluation of current populations as distinct species is in flux. Colin Groves (2001) lists about 350 species of primates in ''Primate Taxonomy''. The recently published third edition of ''Mammal Species of the World'' (MSW 2005) lists 376 species. But even MSW3's list falls short of current understanding as its collection cutoff was in 2003. Notable new species not listed in MSW3 include Cleese's Woolly Lemur (named after [[United Kingdom|British]] [[actor]] and [[lemur]] enthusiast [[John Cleese]]) and the GoldenPalace.com Monkey (whose name was put up for auction).
−
[[Image:EmperorTamarin.jpg|200px|left|thumb|[[Emperor Tamarin]], a [[New World monkey]]]]
 
−
The suborder [[Haplorrhini]], the "dry-nosed" primates, is composed of two sister clades.<ref name=MSW3/> The [[prosimian]] tarsiers in family [[Tarsiidae]] (monotypic in its own infraorder [[Tarsiiformes]]), represent the most primitive division at about 58 mya. The [[Simiiformes]] infraorder contains the two parvorders: the [[New World monkey]]s in one, and the [[Old World monkey]]s, [[human]]s and the other [[ape]]s in the other.<ref name=MSW3/> This division happened about 40 mya. However about 30 mya, three groups split from the main haplorrhine lineage. One group stayed in Asia and are closest in kin to the "dawn monkey" ''[[Eosimias]]''. The second stayed in Africa, where they developed into the Old World primates. The third rafted to South America to become the New World monkeys. Mysteriously the aboriginal Asian Haplorrhini vanished from record once Africa collided with Eurasia 24 mya. Apes and monkeys spread into Europe and Asia. Close behind came lorises and tarsiers, also African castaways. The first hominid fossils were discovered in Northern Africa and date back 7 mya. Modern humans did not appear until 0.2 mya, eventually becoming the most prevalent primate and mammal on Earth.
 
−
 
 
−
The discovery of new species happens at a rate of a few new species each year, and the evaluation of current populations as distinct species is in flux. [[Colin Groves]] lists about 350 species of primates in ''Primate Taxonomy'' in 2001.<ref name=PT>''Primate Taxonomy'' (Smithsonian Institute Press, 2001), Colin Groves (ISBN 1-56098-872-X )</ref> The recently published third edition of ''Mammal Species of the World'' (2005) lists 376 species.<ref name=MSW3/> But even MSW3's list falls short of current understanding as its collection cutoff was in 2003. Notable new species not listed in MSW3 include [[Cleese's Woolly Lemur]] (named after [[United Kingdom|British]] [[actor]] and [[lemur]] enthusiast [[John Cleese]]) and the [[GoldenPalace.com Monkey]] (whose name was put up for auction).
 
  
 
====Extant primate families====
 
====Extant primate families====
 
* '''ORDER PRIMATES'''
 
* '''ORDER PRIMATES'''
−
** '''Suborder [[Strepsirrhini]]''': non-tarsier prosimians
+
** '''Suborder Strepsirrhini:''' non-tarsier prosimians
−
*** Infraorder [[Lemuriformes]]
+
*** Infraorder Lemuriformes
−
**** Superfamily [[Cheirogaleidae|Cheirogaleoidea]]
+
**** Superfamily Cheirogaleoidea
−
***** Family [[Cheirogaleidae]]: dwarf lemurs and mouse-lemurs (24 species)
+
***** Family Cheirogaleidae: dwarf lemurs and mouse-lemurs (24 species)
−
**** Superfamily [[Lemuroidea]]
+
**** Superfamily Lemuroidea
−
***** Family [[Lemuridae]]: lemurs (19 species)
+
***** Family Lemuridae: [[lemur]]s (19 species)
−
***** Family [[Lepilemuridae]]: sportive lemurs (11 species)
+
***** Family Lepilemuridae: sportive lemurs (11 species)
−
***** Family [[Indriidae]]: woolly lemurs and allies (12 species)
+
***** Family Indriidae: woolly lemurs and allies (12 species)
−
*** Infraorder [[Aye-aye|Chiromyiformes]]
+
*** Infraorder Chiromyiformes
−
**** Family [[Aye-aye|Daubentoniidae]]: Aye-aye (1 species)
+
**** Family Daubentoniidae: Aye-aye (1 species)
−
*** Infraorder [[Lorisiformes]]
+
*** Infraorder Lorisiformes
−
**** Family [[Lorisidae]]: lorises, pottos and allies (9 species)
+
**** Family Lorisidae: lorises, pottos and allies (9 species)
−
**** Family [[Galagidae]]: galagos (19 species)
+
**** Family Galagidae: galagos (19 species)
−
** '''Suborder [[Haplorrhini]]''': tarsiers, monkeys and apes
+
** '''Suborder Haplorrhini:''' tarsiers, monkeys and apes
−
*** Infraorder [[tarsier|Tarsiiformes]]
+
*** Infraorder Tarsiiformes
−
**** Family [[tarsier|Tarsiidae]]: tarsiers (7 species)
+
**** Family Tarsiidae: tarsiers (7 species)
−
*** Infraorder [[Simiiformes]]
+
*** Infraorder Simiiformes
−
**** Parvorder [[Platyrrhini]]: New World monkeys
+
**** '''Parvorder Platyrrhini:''' '''New World [[monkey]]s'''
−
***** Family [[Cebidae]]: marmosets, tamarins, capuchins and squirrel monkeys (56 species)
+
***** Family Cebidae: marmosets, tamarins, capuchins and squirrel monkeys (56 species)
−
***** Family [[Aotidae]]: night or owl monkeys (douroucoulis) (8 species)
+
***** Family Aotidae: night or owl monkeys (douroucoulis) (8 species)
−
***** Family [[Pitheciidae]]: titis, sakis and uakaris (41 species)
+
***** Family Pitheciidae: titis, sakis and uakaris (41 species)
−
***** Family [[Atelidae]]: howler, spider and woolly monkeys (24 species)
+
***** Family Atelidae: howler, spider and woolly monkeys (24 species)
−
**** Parvorder [[Catarrhini]]
+
**** '''Parvorder Catarrhini'''
−
***** Superfamily [[Old World monkey|Cercopithecoidea]]
+
***** '''Superfamily Cercopithecoidea'''
−
****** Family [[Old World monkey|Cercopithecidae]]: Old World monkeys (135 species)
+
****** '''Family Cercopithecidae:''' '''Old World [[monkey]]s''' (135 species)
−
***** Superfamily [[Hominoidea]]
+
******* Subfamily Cercopithecinae
−
****** Family [[Hylobatidae]]: gibbons or "lesser apes" (13 species)
+
******** Tribe Cercopithecini (Allen's Swamp Monkey, talapoins, Patas Monkey, Silver Monkey, Guenons, Owl-faced Monkey, etc.)
−
****** Family [[Hominidae]]: humans and other great apes (7 species)
+
******** Tribe Papionini (Macaques, mangabeys, Gelada, Hamadryas Baboon, Olive Baboon, Yellow Baboon, mandrills, etc.)
 +
******* Subfamily Colobinae
 +
******** African group (colobus)
 +
******** Langur (leaf monkey) group (langurs, leaf monkeys, surilis)
 +
******** Odd-Nosed group (doucs, snub-nosed monkeys, Proboscis Monkey, Pig-tailed Langur)
 +
***** '''Superfamily Hominoidea:'''  '''[[Ape]]s'''
 +
****** '''Family Hylobatidae:''' [[gibbon]]s or '''lesser apes''' (13 species)
 +
******* Genus ''Hylobates''
 +
******* Genus ''Hoolock''
 +
******* Genus ''Symphalangus''
 +
******* Genus ''Nomascus''
 +
****** '''Family Hominidae:''' humans and other '''great apes''' (7 species)
 +
****** Genus ''Pongo:'' [[orangutan]]s
 +
****** Genus ''[[Gorilla]]:'' gorillas
 +
****** Genus ''Homo:'' [[human]]s
 +
****** Genus ''Pan:'' [[chimpanzee]]s
  
 
====Some prehistoric primates====
 
====Some prehistoric primates====
−
*''[[Adapis]]'', an adapid
+
*''Adapis,'' an adapid
−
*''[[Australopithecus]]'', a human-like animal
+
*''Australopithecus,'' a human-like animal
−
*''[[Branisella boliviana]]'', an early New World monkey
+
*''Branisella boliviana,'' an early New World monkey
−
*''[[Dryopithecus]]'', an early ape
+
*''Dryopithecus,'' an early ape
−
*''[[Eosimias]]'', an early catarrhine
+
*''Eosimias,'' an early catarrhine
−
*''[[Sahelanthropus tchadensis]]'', a possible ancestor of humans
+
*''Sahelanthropus tchadensis,'' a possible ancestor of humans
−
*''[[Aegyptopithecus zeuxis]]'', an early haplorrhine
+
*''Aegyptopithecus zeuxis,'' an early haplorrhine
−
*''[[Pliopithecus]]'', ancestor of the modern gibbons
+
*''Pliopithecus,'' ancestor of the modern gibbons
−
*''[[Gigantopithecus]]'', the largest ape
+
*''Gigantopithecus,'' the largest ape
−
*''[[Godinotia]]'', an adapid
+
*''Godinotia,'' an adapid
−
*''[[Megaladapis]]'', a giant lemur
+
*''Megaladapis,'' a giant lemur
−
*''[[Notharctus]]'', an adapid
+
*''Notharctus,'' an adapid
−
*''[[Plesiopithecus teras]]'', a relative of lorises and galagos
+
*''Plesiopithecus teras,'' a relative of lorises and galagos
−
*''[[Protopithecus brasiliensis]]'', a giant New World monkey
+
*''Protopithecus brasiliensis,'' a giant New World monkey
−
*''[[Sivapithecus]]'', an early ape
+
*''Sivapithecus,'' an early ape
−
*''[[Tielhardina]]'', the earliest haplorrhines
+
*''Tielhardina,'' the earliest haplorrhines
−
*''[[Victoriapithecus]]'', an early Old World monkey
+
*''Victoriapithecus,'' an early Old World monkey
−
*''[[Pierolapithecus catalaunicus]]'', a possible ancestor of large apes
+
*''Pierolapithecus catalaunicus,'' a possible ancestor of large apes
  
−
==Primate hybrids==
+
==Legal status==
−
In "The Variation Of Animals And Plants Under Domestication" [[Charles Darwin]] noted: "Several members of the family of [[Lemur]]s have produced hybrids in the Zoological Gardens."
+
Humans are the only apes recognized as "persons" and protected in law by the [[United Nations]] ''Universal Declaration of Human Rights'' and by all governments, though to varying degrees.  
−
 
 
−
Many [[gibbon]]s are hard to identify based on fur coloration and are identified either by song or genetics. These morphological ambiguities have led to [[hybrid]]s in zoos. Zoo gibbons usually come from the black market pet trade in Southeast Asia, which transported gibbons across countries all over the region. As a result, perhaps as much as 95% of zoo gibbons are of unknown geographic origin. As most zoos rely on morphological variation or labels that are impossible to verify to assign species and subspecies names, it is unfortunately common for gibbons to be misidentified and housed together. For example, some collections' supposedly pure breeding pairs were actually mixed pairs or hybrids from previous mixed pairs.  The hybrid offspring were sent to other gibbon breeders and led to further hybridization in captive gibbons. Within-genus hybrids also occur in wild gibbons where the ranges overlap ([[Agile Gibbon]]s and [[Pileated Gibbon]]s x [[Lar Gibbon]]s, [[Agile Gibbon]]s x [[Müller's Bornean Gibbon]], [[Yellow-cheeked Gibbon]]s x [[Northern White-cheeked Gibbon]]s).
 
−
 
 
−
Intergeneric gibbon hybridizations have only occurred in captivity. [[Silvery Gibbon]]s (''Hylobates moloch'') and Müller's Bornean Gibbon (''H. muelleri'') have hybridized with [[Siamang]]s (''Symphalangus syndactylus'') in captivity - a female Siamang produced hybrid "Siabon" offspring on 2 occasions when housed with a male gibbon; only one hybrid survived.
 
−
 
 
−
[[Anubis Baboon]]s and [[Hamadryas Baboon]]s have hybridized in the wild where their ranges meet. A [[Rheboon]] is a captive-bred [[Rhesus Macaque]]/Hamadryas Baboon hybrid with a [[baboon]]-like body shape and [[macaque]]-like tail.
 
−
 
 
−
Different macaque species can interbreed. In "The Variation Of Animals And Plants Under Domestication" Charles Darwin wrote: ''A Macacus, according to Flourens, bred in Paris; and more than one species of this genus has produced young in London, especially the Macacus rhesus, which everywhere shows a special capacity to breed under confinement. Hybrids have been produced both in Paris and London from this same genus.''  The [[Japanese Macaque]] (''Macaca fuscata'') has interbred with the introduced [[Taiwanese Macacque]] (''M. cyclopis'') when the latter escaped into the wild from private zoos.  
 
  
−
Various hybrid monkeys are bred within the pet trade, for example:
+
The non-human apes are not classified as persons. This has generated much debate, particularly through the Great Ape Project ''Declaration on Great Apes,'' which argues for their personhood. Many argue that the other apes' cognitive capacity in itself, as well as their close [[gene|genetic]] relationship to human beings, dictates an acknowledgment of personhood.  
−
* Hybrid [[capuchin monkey]]s e.g. [[Tufted Capuchin]] (''Cebus apella'') x [[Weeper Capuchin]] (''C. olivaceus'')
 
−
* [[Liontail Macaque]] x [[Pigtail Macaque]] hybrids
 
−
* [[Rhesus Macaque]] x [[Stumptail Macaque]] hybrids.
 
−
 
 
−
Among [[Old World monkey]]s, natural hybridization is not uncommon. There numerous field reports of hybrid monkeys and detailed studies of zones where species overlap and hybrids occur.
 
−
 
 
−
Among the great apes, [[Sumatran Orangutan|Sumatran]] and [[Bornean Orangutan|Bornean]] [[orangutan]]s are considered separate species with anatomical differences, producing sterile or poorly fertile hybrids. Hybrid orangutans are genetically weaker, with lower survival rates than pure animals.
 
−
 
 
−
==Legal status==
 
−
[[Image:Monkey5.jpg|right|thumb|300px|Monkeys imported for experimentation in a crate. Credit: [[BUAV]]]]
 
−
Humans are recognized as [[person]]s and protected in law by the [[United Nations]] [[Universal Declaration of Human Rights]]<ref name=UN>[http://www.un.org/Overview/rights.html UN Declaration of Human Rights]</ref> and by all governments, though to varying degrees. Non-human primates are not classified as persons, which means their individual interests have no formal recognition or protection. The status of non-human primates has generated much debate, particularly through the [[Great Ape Project]]<ref name=GA>[http://www.greatapeproject.org/declaration.html ''Declaration on Great Apes''], Great Ape Project</ref> which argues for their [[personhood]].
 
  
−
Thousands of primates are used every year around the world in scientific [[Animal testing|experiments]] because of their psychological and physiological similarity to [[human]]s. The species most commonly used are chimpanzees, [[baboon]]s, [[marmoset]]s, [[macaque]]s, and African [[green monkey]]s. In the [[European Union]], around 10,000 were used in 2004, with 4,652 experiments conducted on 3,115 non-human primates in the UK alone in 2005.<ref name=BUAV>[http://www.buav.org/campaigns/primates British Union for the Abolition of Vivisection - Primates]</ref> As of 2004, 3,100 non-human great apes were living in captivity in the [[United States]], in [[zoo]]s, [[circus]]es, and [[Laboratory|laboratories]], 1,280 of them being used in experiments.<ref name=GA/> European campaign groups such as the [[BUAV]] are seeking a ban on all primate use in experiments as part of the [[European Union]]'s current review of existing law on animal experimentation.
+
However, despite genetic and morphological similarities between the other great apes and [[human being]]s, the gulf between these species is qualitatively immense. References in the ''Universal Declaration of Human Rights'' itself is indicative of many fundamental aspects of ''Homo sapiens'' that are unthinkable for [[chimpanzee]]s, [[gorilla]]s, and [[organgutan]]s. Among these are rights related to [[religion]], politics, freedom of [[conscience]], the arts, scientific advancement, government, and so forth.  
  
−
==See also==
+
One role that many humans see for themselves is as stewards of creation, with the responsibility to care for other living organisms. Despite this, however, thousands of primates are used every year around the world in scientific experiments because of their similarity to humans. The species most commonly used are chimpanzees, [[baboon]]s, [[marmoset]]s, [[macaque]]s, and African green monkeys. However, in general, there are many more laws protecting primates, and [[mammal]]s in general, than are protecting [[invertebrate]]s, [[fish]], [[amphibian]]s, and [[reptile]]s.
−
*[[Arboreal theory]]
 
−
*[[Primatology]]
 
  
 
==References==
 
==References==
−
{{Sisterlinks|Primate}}
 
−
{{Wikispecies|Primates}}
 
−
{{Wikibookspar|Dichotomous Key|Primates}}
 
−
<references/>
 
  
−
* {{note|PQ}}''Primates in Question'' (Smithsonian Institute Press, 2003), Robert W. Shumaker & Benjamin B. Beck (ISBN 1-58834-176-3 )
+
* Groves, C. 2001. ''Primate Taxonomy.'' Smithsonian Institute Press. ISBN 1-56098-872-X
 +
* Groves, C., D. E. Wilson, and D. M. Reeder, eds. 2005. ''Mammal Species of the World,'' 3rd edition, Johns Hopkins University Press. ISBN 0-801-88221-4
 +
* Mayr, E. 2001. ''What Evolution Is''. New York: Basic Books. ISBN 0465044263
 +
* Mootnick, A., and C. P. Groves. 2005. A new generic name for the hoolock gibbon (Hylobatidae). ''International Journal of Primatology'' 26:971-976.
 +
* Shumaker, R. W., and B. B. Beck. 2003. ''Primates in Question''. Smithsonian Institute Press. ISBN 1-58834-176-3
 +
* Wood, T. C. 2006. [http://www.bryancore.org/bsg/opbsg/007.pdf The chimpanzee genome and the problem of biological similarity]. ''Occas. papers of the BSG'' 7:1-18.
  
−
==External links==
 
−
* [http://primate-brain.org High-Resolution Cytoarchitectural Primate Brain Atlases]
 
−
* [http://pin.primate.wisc.edu  Primate Info Net]
 
−
* [http://www.buav.org/campaigns/primates/index.html Primates in scientific experimentation]
 
−
* [http://www.pri.kyoto-u.ac.jp/ Primate Research Institute, Kyoto University]
 
  
−
{{credit|74156174}}
+
{{credit5|Primate|74156174|Prosimian|72379726|New_World_monkey|64833388|Old_World_monkey|73605584|Ape|73830647}}
 
[[Category:Life sciences]]
 
[[Category:Life sciences]]
 +
[[Category:Animals]]
 +
[[Category:Mammals]]
 +
[[Category:Primates]]

Latest revision as of 15:45, 7 November 2011

Primates
Olive baboon.jpg
Olive Baboon
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Subclass: Eutheria
(unranked) Euarchontoglires
Superorder: Euarchonta
Order: Primates
Linnaeus, 1758
Families

A primate (L. prima, first) is any mammal of the biological order Primates, the group that contains all the species commonly related to the lemurs, monkeys, and apes, with the latter category including humans. Primates are characterized by being anatomically unspecialized, with limbs capable of performing a variety of functions, refined five-digit hands and feet adapted for grasping (including opposable thumbs), comparatively flattened snouts, and prolonged pre- and postnatal development, among other features.

The Primates order is divided informally into three main groupings: Prosimians, monkeys of the New World, and monkeys and apes of the Old World. The prosimians are species whose bodies most closely resemble that of the early proto-primates. The most well known of the prosimians, the lemurs, are located on the island of Madagascar and, to a lesser extent, on the Comoros Islands, isolated from the rest of the world. The New World monkeys include the familiar capuchin, howler, and squirrel monkeys. They live exclusively in the Americas. Discounting humans, the rest of the simians (monkeys and apes), the Old World monkeys, and the apes, inhabit Africa and southern and central Asia, although fossil evidence shows many species once existed in Europe as well.

Based on purely biological aspects (DNA, proteins, morphology, etc.), it is clear the humans are primates. Indeed, humans and chimpanzees share more than 98 percent identity by various molecular comparisons (protein sequences, allele differences, DNA nucleotide sequences) (Wood 2006). However, humans are also defined in cultural, spiritual, psychological, and behavioral terms that make them qualitatively very different from other primates. Despite this, effort is sometimes made to consider humans "just primates," to the extent that efforts are actually being made to legally define other primates as "persons."

Primates are found all over the world. Non-human primates occur mostly in Central and South America, Africa, and southern Asia. A few species exist as far north in the Americas as southern Mexico, and as far north in Asia as northern Japan. The earliest fossils of primates date to the late Cretaceous period (Mayr 2001).

The English singular, primate, is a back-formation from the Latin name Primates, which itself was the plural of the Latin primas ("one of the first, excellent, noble").

Physical description

All primates have five fingers (pentadactyl), a generalized dental pattern, and an unspecialized body plan. Another distinguishing feature of primates is fingernails. Opposing thumbs are also a characteristic primate feature, but are not limited to this order; opossums, for example, also have opposing thumbs. In primates, the combination of opposing thumbs, short fingernails (rather than claws), and long, inward-closing fingers is considered a relic of the ancestral practice of brachiating through trees. Forward-facing color binocular vision would also have been useful for the brachiating ancestors, particularly for finding and collecting food, although some studies suggest it was more useful in courtship. All primates, even those that lack the features typical of other primates (like lorises), share eye orbit characteristics, such as a postorbital bar, that distinguish them from other taxonomic orders.

Old World species tend to have significant sexual dimorphism. This is characterized most in size difference, with males being up to a bit more than twice as heavy as females. New World species generally form pair bonds and these species (including tamarins and marmosets) generally do not show a significant size difference between the sexes.

Species Female Male
Gorilla 105 kg (231 lb) 205 kg (452 lb)
Human 62.5 kg (137.5 lb) 78.4 kg (172 lb)
Patas Monkey 5.5 kg (12 lb) 10 kg (22 lb)
Proboscis Monkey 9 kg (20 lb) 19 kg (42 lb)
Pygmy Marmoset 120 g (4.2 oz) 140 g (5 oz)

Habitat

Many modern species of primates live mostly in trees and hardly ever come to the ground. Other species are partially terrestrial, such as baboons and the Patas Monkey. Only a few species are fully terrestrial, such as the Gelada and Gorilla.

Primates live in a diverse number of forested habitats, including rainforests, mangrove forests, and mountain forests to altitudes of over 3000 m. Although most species are generally shy of water, a few are fine swimmers and are comfortable in swamps and watery areas, including the Proboscis Monkey, De Brazza's Monkey, and Allen's Swamp Monkey, which even has small webbing between its fingers. Some primates, such the Rhesus Macaque and the Hanuman Langur, are common in cities and villages.

Types of primates

The three basic groups of primates are prosimians, New World monkeys, and Old World monkeys and apes. The monkeys (New World monkeys and Old World monkeys) and apes are also referred to as simians.


Prosimians

The Ring-tailed Lemur (Lemur catta) is a prosimian of the family Lemuridae.

Prosimians are generally considered the most primitive extant (living) primates, representing forms that were ancestral to monkeys and apes. With the exception of the tarsiers, all of the prosimians are in the suborder Strepsirrhini. These include the lemurs, Aye-aye, and lorises. The tasiers are placed in the suborder Haplorrhini (with the monkeys and apes). Due to this reason, the classification is not considered valid in terms of phylogeny, as they do not share a unique last common ancestor or anatomical traits.

New World monkeys

The New World monkeys are the four families of primates that are found in Central and South America: The Cebidae (marmosets, tamarins, capuchins, and squirrel monkeys), Aotidae (night or owl monkeys), Pitheciidae (titis, sakis, and uakaris), and Atelidae (howler, spider, and woolly monkeys).

Spider monkey A. geoffroyi

The four families are ranked together as the Platyrrhini parvorder, placing them in a different grouping from the Old World monkeys and the apes.

All New World monkeys differ slightly from Old World monkeys in many aspects, but the most prominent difference is the nose, which is the feature most commonly used to distinguish between the two groups. The scientific name for New world monkey, Platyrrhini, means "flat nosed," therefore the noses are flatter, with side facing nostrils, compared to the narrow noses of the Old World monkey. Most New World monkeys have long, often prehensile tails. Many are small, arboreal, and nocturnal, so knowledge of them is less comprehensive than that of the more easily observed Old World monkeys. Unlike most Old World monkeys, many New World monkeys form monogamous pair bonds and show substantial paternal care of young.

Old World monkeys and apes

Old World monkeys

Old World monkey: Crab-eating Macaque from Malaysia

The Old World monkeys, or Cercopithecidae family, are placed in the superfamily Cercopithecoidea in the clade Catarrhini. From the point of view of superficial appearance, they are unlike apes in that most have tails (the family name means "tailed ape"), and unlike the New World monkeys in that their tails are never prehensile (adapted to be able to grasp and hold objects). Technically, the distinction of catarrhines from platyrrhines (New World monkeys) depends on the structure of the nose, and the distinction of Old World monkeys from apes depends on dentition.

The Old World monkeys are native to Africa and Asia today, but are also known from Europe in the fossil record. They include many of the most familiar species of non-human primates, such as baboons, macaques, and colobus monkeys. There are about 135 known species of Old World monkeys.

Two subfamilies of Old World monkeys are recognized, the Cercopithecinae, which are mainly African, but include the diverse genus of macaques, which are Asian and North African, and the Colobinae, which includes most of the Asian genera, as well as the African colobus monkeys.

Apes

Ape: Lar Gibbon (Hylobatidae family)

Apes are the members of the Hominoidea superfamily of primates. Under the taxonomic scheme that is currently common, there are two families of hominoids:

  • The family Hylobatidae consists of 4 genera and 12 species of gibbons, collectively known as the "lesser apes";
  • The family Hominidae, consisting of gorillas, chimpanzees, orangutans, and humans, is collectively known as the "great apes."

Awkwardly, the term "hominid" is still used with the specific meaning of extinct animals more closely related to humans than the other great apes (for example, Australopithecines).

In another classification, historically common, members of the Hominoidea are divided into three families, the Hylobatidae (gibbons), the Pongidae (orangutans, gorillas, chimpanzees, and bonobos), and the Hominidae (humans). In this case, the term hominid accurately applies to all human and humanlike forms, while great apes applies only to gorillas, chimpanzees, and orangutans.

Some researchers place gorillas and chimpanzees (and the related bonobos) into the Panidae family, while orangutans remain in the Pongidae family, and humans in the Hominidae. The bonobos are sometimes considered a variety of chimpanzees and are referred to as pygmy chimpanzees.

The eight extant (living) genera of hominoids are the four great ape genera (Homo (humans), Pan (chimpanzees), Gorilla, and Pongo (orangutans)), and the four genera of gibbons (Hylobates, Hoolock, Nomascus, and Symphalangus). The genus for the hoolock gibbons was changed from Bunopithecus to Hoolock (Mootnick and Groves 2005).

A few other primates have the word "ape" in their common names, but they are not regarded as true apes.

Both great apes and lesser apes fall within parvorder Catarrhini, which also includes the Old World monkeys of Africa and Eurasia. Within this group, both families of apes can be distinguished from these monkeys by the number of cusps on their molars (apes have five—the "Y-5" molar pattern, Old World monkeys have only four in a "bilophodont" pattern). Apes have more mobile shoulder joints and arms, ribcages that are flatter front-to-back, and a shorter, less mobile spine compared to Old World monkeys. These are all anatomical adaptations to vertical hanging and swinging locomotion (brachiation) in the apes. All living members of the Hylobatidae and Hominidae are tailless, and thus humans have been referred to as tailless, bipedal, primates. However there are also primates in other families that lack tails, and at least one (the Pig-Tailed Langur) that has been known to walk significant distances bipedally.

Except for gorillas and humans, all true apes are agile climbers of trees. They are best described as omnivorous, their diet consisting of fruit, grass seeds, and in most cases some quantities of meat and invertebrates—either hunted or scavenged—along with anything else available and easily digested. The apes are native to Africa and Asia, although humans have spread to all parts of the world.

Most ape species are rare or endangered. The chief threat to most of the endangered species is loss of tropical rainforest habitat, though some populations are further imperiled by hunting.

Many people do not consider the term "apes" to include humans, although biologists generally do consider humans as apes. The terms "non-human apes" or "non-human great apes" is used with increasing frequency to both show the relationship of humans to the other species classified in the families Hylobatidae and Hominidae, while maintaining the distinction of humans.

Classification

The Primate order lies in a tight clustering of related orders (the Euarchontoglires) within the Eutheria, a subclass of Mammalia. Recent molecular genetic research on primates, flying lemurs, and treeshrews has shown that the two species of flying lemur (Dermoptera) are more closely related to the primates than the treeshrews of the order Scandentia, even though the treeshrews were at one time considered primates. These three orders make up the Euarchonta clade. This clade combines with the Glires clade (made up of the Rodentia, rodents, and Lagomorpha, rabbits) to form the Euarchontoglires clade. Variously, both Euarchonta and Euarchontoglires are ranked as superorders. Also, some scientists consider Dermoptera a suborder of Primates and call the "true" primates the suborder Euprimates. 

Euarchontoglires
    ├─Glires
    │    ├─rodents (Rodentia)
    │    └─rabbits, hares, pikas (Lagomorpha)
    └─Euarchonta
         ├─treeshrews (Scandentia)
         └─N.N.
              ├─flying lemurs (Dermoptera
              └─N.N.
                   ├─Plesiadapiformes (extinct)
                   └─primates (Primates)


Ring-tailed Lemur, a strepsirrhine primate

In older classifications, the Primates were divided into two superfamilies: Prosimii and Anthropoidea. The Prosimii included all of the prosimians: All of Strepsirrhini plus the tarsiers. The Anthropoidea contained all of the simians.

In modern cladistic reckonings, the Primate order is also a true clade. The suborder Strepsirrhini, the "wet-nosed" primates, split off from the primitive primate line about 63 million years ago (mya). The seven strepsirhine families are the four related lemur families and the three remaining families that include the lorises, the Aye-aye, the galagos, and the pottos. Some classification schemes wrap the Lepilemuridae into the Lemuridae and the Galagidae into the Lorisidae, yielding a three-two family split instead of the four-three split as presented here.

Other lineages of lower primates once inhabited Earth. During the Eocene, most of the northern continents were dominated by two dominant groups, the adapids and the omomyids. The former is considered a member of Strepsirrhini, but it does not have a tooth-comb like modern lemurs. The latter was related closely to tarsiers, monkeys, and apes. Adapids survived until 10 mya; omomyids, on the other hand, perished 20 million years earlier.

The Aye-aye is difficult to place in Strepsirrhini. Its family, Daubentoniidae, could be a lemuriform primate and its ancestors split from lemur line more recently than the lemurs and lorises split, about 50 mya. Otherwise, it is sister to all of the other strepsirrhines, in which case it evolved away from the main strepsirrhine line between 50 and 63 mya.

Emperor Tamarin, a New World monkey

The suborder Haplorrhini, the "dry-nosed" primates, is composed of two sister clades. The prosimian tarsiers in family Tarsiidae (monotypic in its own infraorder Tarsiiformes), represent the most primitive division at about 58 mya. The Simiiformes infraorder contains the two parvorders: The New World monkeys in one, and the Old World monkeys, humans, and the other apes in the other. This division is considered to have happened about 40 mya. However, about 30 mya, it is thought that these three groups split from the main haplorrhine lineage. One group stayed in Asia and are closest in kin to the "dawn monkey" Eosimias. The second stayed in Africa, where they developed into the Old World primates. The third rafted to South America to become the New World monkeys. Mysteriously the aboriginal Asian Haplorrhini vanished from record once Africa collided with Eurasia 24 mya. Apes and monkeys spread into Europe and Asia. Close behind came lorises and tarsiers, also African castaways. The first hominid fossils were discovered in Northern Africa and date back 7 mya. Modern humans did not appear until 0.2 mya, eventually becoming the most prevalent primate and mammal on Earth.

The discovery of new species happens at a rate of a few each year, and the evaluation of current populations as distinct species is in flux. Colin Groves (2001) lists about 350 species of primates in Primate Taxonomy. The recently published third edition of Mammal Species of the World (MSW 2005) lists 376 species. But even MSW3's list falls short of current understanding as its collection cutoff was in 2003. Notable new species not listed in MSW3 include Cleese's Woolly Lemur (named after British actor and lemur enthusiast John Cleese) and the GoldenPalace.com Monkey (whose name was put up for auction).

Extant primate families

  • ORDER PRIMATES
    • Suborder Strepsirrhini: non-tarsier prosimians
      • Infraorder Lemuriformes
        • Superfamily Cheirogaleoidea
          • Family Cheirogaleidae: dwarf lemurs and mouse-lemurs (24 species)
        • Superfamily Lemuroidea
          • Family Lemuridae: lemurs (19 species)
          • Family Lepilemuridae: sportive lemurs (11 species)
          • Family Indriidae: woolly lemurs and allies (12 species)
      • Infraorder Chiromyiformes
        • Family Daubentoniidae: Aye-aye (1 species)
      • Infraorder Lorisiformes
        • Family Lorisidae: lorises, pottos and allies (9 species)
        • Family Galagidae: galagos (19 species)
    • Suborder Haplorrhini: tarsiers, monkeys and apes
      • Infraorder Tarsiiformes
        • Family Tarsiidae: tarsiers (7 species)
      • Infraorder Simiiformes
        • Parvorder Platyrrhini: New World monkeys
          • Family Cebidae: marmosets, tamarins, capuchins and squirrel monkeys (56 species)
          • Family Aotidae: night or owl monkeys (douroucoulis) (8 species)
          • Family Pitheciidae: titis, sakis and uakaris (41 species)
          • Family Atelidae: howler, spider and woolly monkeys (24 species)
        • Parvorder Catarrhini
          • Superfamily Cercopithecoidea
            • Family Cercopithecidae: Old World monkeys (135 species)
              • Subfamily Cercopithecinae
                • Tribe Cercopithecini (Allen's Swamp Monkey, talapoins, Patas Monkey, Silver Monkey, Guenons, Owl-faced Monkey, etc.)
                • Tribe Papionini (Macaques, mangabeys, Gelada, Hamadryas Baboon, Olive Baboon, Yellow Baboon, mandrills, etc.)
              • Subfamily Colobinae
                • African group (colobus)
                • Langur (leaf monkey) group (langurs, leaf monkeys, surilis)
                • Odd-Nosed group (doucs, snub-nosed monkeys, Proboscis Monkey, Pig-tailed Langur)
          • Superfamily Hominoidea: Apes
            • Family Hylobatidae: gibbons or lesser apes (13 species)
              • Genus Hylobates
              • Genus Hoolock
              • Genus Symphalangus
              • Genus Nomascus
            • Family Hominidae: humans and other great apes (7 species)
            • Genus Pongo: orangutans
            • Genus Gorilla: gorillas
            • Genus Homo: humans
            • Genus Pan: chimpanzees

Some prehistoric primates

  • Adapis, an adapid
  • Australopithecus, a human-like animal
  • Branisella boliviana, an early New World monkey
  • Dryopithecus, an early ape
  • Eosimias, an early catarrhine
  • Sahelanthropus tchadensis, a possible ancestor of humans
  • Aegyptopithecus zeuxis, an early haplorrhine
  • Pliopithecus, ancestor of the modern gibbons
  • Gigantopithecus, the largest ape
  • Godinotia, an adapid
  • Megaladapis, a giant lemur
  • Notharctus, an adapid
  • Plesiopithecus teras, a relative of lorises and galagos
  • Protopithecus brasiliensis, a giant New World monkey
  • Sivapithecus, an early ape
  • Tielhardina, the earliest haplorrhines
  • Victoriapithecus, an early Old World monkey
  • Pierolapithecus catalaunicus, a possible ancestor of large apes

Legal status

Humans are the only apes recognized as "persons" and protected in law by the United Nations Universal Declaration of Human Rights and by all governments, though to varying degrees.

The non-human apes are not classified as persons. This has generated much debate, particularly through the Great Ape Project Declaration on Great Apes, which argues for their personhood. Many argue that the other apes' cognitive capacity in itself, as well as their close genetic relationship to human beings, dictates an acknowledgment of personhood.

However, despite genetic and morphological similarities between the other great apes and human beings, the gulf between these species is qualitatively immense. References in the Universal Declaration of Human Rights itself is indicative of many fundamental aspects of Homo sapiens that are unthinkable for chimpanzees, gorillas, and organgutans. Among these are rights related to religion, politics, freedom of conscience, the arts, scientific advancement, government, and so forth.

One role that many humans see for themselves is as stewards of creation, with the responsibility to care for other living organisms. Despite this, however, thousands of primates are used every year around the world in scientific experiments because of their similarity to humans. The species most commonly used are chimpanzees, baboons, marmosets, macaques, and African green monkeys. However, in general, there are many more laws protecting primates, and mammals in general, than are protecting invertebrates, fish, amphibians, and reptiles.

References
ISBN links support NWE through referral fees

  • Groves, C. 2001. Primate Taxonomy. Smithsonian Institute Press. ISBN 1-56098-872-X
  • Groves, C., D. E. Wilson, and D. M. Reeder, eds. 2005. Mammal Species of the World, 3rd edition, Johns Hopkins University Press. ISBN 0-801-88221-4
  • Mayr, E. 2001. What Evolution Is. New York: Basic Books. ISBN 0465044263
  • Mootnick, A., and C. P. Groves. 2005. A new generic name for the hoolock gibbon (Hylobatidae). International Journal of Primatology 26:971-976.
  • Shumaker, R. W., and B. B. Beck. 2003. Primates in Question. Smithsonian Institute Press. ISBN 1-58834-176-3
  • Wood, T. C. 2006. The chimpanzee genome and the problem of biological similarity. Occas. papers of the BSG 7:1-18.


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