Difference between revisions of "Rhododendron" - New World Encyclopedia

From New World Encyclopedia
 
(22 intermediate revisions by 4 users not shown)
Line 1: Line 1:
 +
{{Copyedited}}{{Images OK}}{{Approved}}
 
{{Taxobox
 
{{Taxobox
 
| name = ''Rhododendron''
 
| name = ''Rhododendron''
Line 23: Line 24:
 
*''[[Rhododendron subgenus Vireya|Vireya]]''  
 
*''[[Rhododendron subgenus Vireya|Vireya]]''  
 
}}
 
}}
'''''Rhododendron''''' is the common and [[genus]] name for a large and diverse group of woody [[shrub]]s and small (rarely large) [[tree]]s in the [[flowering plant]] family [[Ericaceae]]. It is a large genus with over 1000 [[species]], and subdivided into various subgenera (commonly 5, 8, or 9, depending on the taxonomic scheme). The trees may be [[deciduous]] or [[evergreen]] and the spirally arranged leaves can vary significantly in size according to species, from 1 to 2 centimeters (0.4-0.8 inches) up to 100 centimeters (39 inches). Members of two of the traditional subgenera, ''Pentanthera'' (deciduous), and ''Titsushi'' (evergreen), are known as [[azalea]]s.
+
'''''Rhododendron''''' is the common and [[genus]] name for a large and diverse group of woody [[shrub]]s and small (rarely large) [[tree]]s in the [[flowering plant]] family [[Ericaceae]]. It is a large genus with over 1000 [[species]], and subdivided into various subgenera (commonly eight, nine, or five, depending on the taxonomic scheme). The trees may be [[deciduous]] or [[evergreen]] and the spirally arranged leaves can vary significantly in size according to species, from one to two centimeters (0.4-0.8 inches) up to 100 centimeters (39 inches). Members of two of the traditional subgenera, ''Pentanthera'' (deciduous), and ''Titsushi'' (evergreen), are known as [[azalea]]s.
  
and most have showy flower displays. Indeed, the name rhododendron comes from the [[Greek language|Greek]] ''rhodos'', meaning "rose," and ''dendron'', meaning "tree."
+
Rhododendrons, including azaleas, generally have showy flower displays. Indeed, the name rhododendron comes from the [[Greek language|Greek]] ''rhodos'', meaning "rose," and ''dendron'', meaning "tree." As such, and combined with their often being evergreen, they often are cultivated as ornamental plants, such as around the foundations of houses or in woodland gardens, and even as specimen plants.
 +
 
 +
They also provide important [[ecology|ecological]] values. They provide food for a number of insects and an important source of [[pollen]] and [[nectar]] for many [[bumblebee]]s, who in turn [[pollination|pollinate]] the plants. However, rhododendrons also are poisonous to many grazing animals, such as [[horse]]s.
 +
{{toc}}
 +
Some species (e.g. ''[[Rhododendron ponticum]]'' in the [[United Kingdom]]) are invasive as introduced plants, spreading in woodland areas replacing the natural understory. ''R. ponticum'' is difficult to eradicate, as its roots can make new shoots.
  
 
==Overview and description==
 
==Overview and description==
 
+
[[Image:Rhododendron decorum ssp diaprepes -1.jpg|thumb|right|240px|''Rhododendron decorum'' subsp. ''diaprepes'']]
 
Ericaceae, the family to which the [[genus]] ''Rhododendron'' belongs, is known as the "heath family," and consists of herbs, shrubs, and trees with leaves that are alternate, simple, and estipulate, and mostly [[calcium oxide|lime]]-hating or [[calcifuge]] plants that thrive in [[acid]] [[soil]]s. In addition to rhododendrons, well-known examples include [[cranberry]], [[blueberry]], [[Erica|heath]], [[Calluna vulgaris|heather]], and [[huckleberry]].  
 
Ericaceae, the family to which the [[genus]] ''Rhododendron'' belongs, is known as the "heath family," and consists of herbs, shrubs, and trees with leaves that are alternate, simple, and estipulate, and mostly [[calcium oxide|lime]]-hating or [[calcifuge]] plants that thrive in [[acid]] [[soil]]s. In addition to rhododendrons, well-known examples include [[cranberry]], [[blueberry]], [[Erica|heath]], [[Calluna vulgaris|heather]], and [[huckleberry]].  
  
The ''Rhododendron'' [[genus]] is characterized by [[shrub]]s and small to (rarely) large [[tree]]s, the smallest species growing to 10 to 100 centimeters (3.9-39 inches) tall, and the largest, ''R. giganteum'', reported to over 30 meters (98 feet) tall (Francis 2006). The [[leaf|leaves]] are spirally arranged; leaf size can range from 1 to 2 centimeters (0.4-0.8 inches) to over 50 centimeters (20 inches), exceptionally 100 centimeters (39.4 inches) in ''R. sinogrande''. They may be either [[evergreen]] or [[deciduous]]. In some species the underside of the leaves is covered with scales (lepidote) or hairs (indumentum).  
+
The ''Rhododendron'' [[genus]] is characterized by [[shrub]]s and small to (rarely) large [[tree]]s, the smallest species growing to ten to 100 centimeters (3.9-39 inches) tall, and the largest, ''R. giganteum'', reported to over 30 meters (98 feet) tall (Francis 2006). The [[leaf|leaves]] are spirally arranged; leaf size can range from one to two centimeters (0.4-0.8 inches) to over 50 centimeters (20 inches), exceptionally 100 centimeters (39.4 inches) in ''R. sinogrande''. They may be either [[evergreen]] or [[deciduous]]. In some species the underside of the leaves is covered with scales (lepidote) or hairs (indumentum).  
  
 
Some of the best known species of rhododendrons are noted for their many clusters of large flowers. There are alpine species with small flowers and small leaves, and tropical species such as section ''Vireya'' that often grow as [[epiphyte]]s.  
 
Some of the best known species of rhododendrons are noted for their many clusters of large flowers. There are alpine species with small flowers and small leaves, and tropical species such as section ''Vireya'' that often grow as [[epiphyte]]s.  
  
some rhododendrons are known by the common name of [[azalea]]. These are flowering [[shrub]]s that are characterized by the lack of scales on the underside of the generally thin, soft, and pointed [[leaf|leaves]], and typically having terminal blooms (one flower per stem), [[flower]]s with five or six stamens, and with long straight hairs parallel to the leaf surface and along the midrid on the ventral surface of the leaf. Originally, azaleas were classed as a different genus of [[plant]], but now they are recognized as part of the rhododendrons. In the common classification of rhododendrons into eight or nine subgenera, azaleas comprise two of the subgenera: ''[[Pentanthera]]'' (deciduous) and ''Titsushi'' (evergreen).
+
Some rhododendrons are known by the common name of [[azalea]]. These are flowering [[shrub]]s that are characterized by the lack of scales on the underside of the generally thin, soft, and pointed [[leaf|leaves]], and typically having terminal blooms (one flower per stem), [[flower]]s with five or six stamens, and with long straight hairs parallel to the leaf surface and along the midrid on the ventral surface of the leaf. Originally, azaleas were classed as a different genus of [[plant]], but now they are recognized as part of the rhododendrons. In the common classification of rhododendrons into eight or nine subgenera, azaleas comprise two of the subgenera: ''[[Pentanthera]]'' (deciduous) and ''Titsushi'' (evergreen).
  
 
==Distribution==
 
==Distribution==
'''Rhododendron''' is a very widely distributed genus, occurring throughout most of the [[Northern Hemisphere]] except for dry areas, and extending into the [[Southern Hemisphere]] in southeastern [[Asia]] and northern [[Australasia]]. The highest species diversity is found in the [[Himalaya]]n mountains from [[Uttarakhand]], [[Nepal]] and [[Sikkim]] to [[Yunnan]] and [[Sichuan]], with other significant areas of diversity in the mountains of [[Indo-China]], [[Korea]], [[Japan]] and [[Taiwan]]. Rhododendron, also known as "Lali Guras" in local language is the national flower of [[Nepal]] and the state tree of the state of [[Uttarakhand]] in [[India]]. In Korea, [[Rhododendron schlippenbachii]] is particularly common (called 철쭉(choltjuk)). In addition, there are a significant number of tropical rhododendron species from southeast Asia to northern [[Australia]], with 55 known species in [[Borneo]] and 164 in [[New Guinea]]. <ref name="Argent 2006">Argent, G. ''Rhododendrons of subgenus Vireya''. 2006. Royal Horticultural Society. ISBN 1-902896-61-0</ref> Relatively fewer species occur in [[North America]] and [[Europe]].  Rhododendrons have not been found in South America or Africa.
+
Rhododendron is a very widely distributed genus, occurring throughout most of the [[Northern Hemisphere]] except for dry areas, and extending into the [[Southern Hemisphere]] in southeastern [[Asia]] and northern [[Australasia]].  
  
==Classification==
+
The highest species diversity is found in the [[Himalaya]]n mountains from [[Uttarakhand]], [[Nepal]], and [[Sikkim]] to [[Yunnan]] and [[Sichuan]], with other significant areas of diversity in the mountains of [[Indo-China]], [[Korea]], [[Japan]], and [[Taiwan]]. Rhododendron, also known as "Lali Guras" in local language, is the national flower of Nepal and the state tree of the state of Uttarakhand in [[India]]. In Korea, [[Rhododendron schlippenbachii]] is particularly common (called 철쭉 (choltjuk)). In addition, there are a significant number of tropical rhododendron species from southeast Asia to northern [[Australia]], with 55 known species in [[Borneo]] and 164 in [[New Guinea]] (Argent 2006).  
:When Linnaeus created the botanical grouping called genus Rhododendron in 1753, he created a separate genus for Azaleas containing 6 species. In 1796 Salisbury pointed out that Azalea and Rhododendron could not be maintained as distinct genera. In 1834, George Don subdivided the genus Rhododendron into 8 sections which were recognized until 2004. Azaleas comprised two of these sections, Subgenus Pentanthera typified by Rhododendron nudiflorum and Subgenus Tsutsusi typified by Rhododendron Tsutsusi.
 
  
http://www.rhodyman.net/rhodyhis.html#anchor282700
+
Relatively fewer species occur in [[North America]] and [[Europe]]. Rhododendrons have not been found in [[South America]] or [[Africa]].
http://www.rhodyman.net/rhodyhis.html#anchor282700
 
Rhododendron History
 
Henning, ? 2007  Hennings Home Page
 
:Based upon genetic and chemical data, in 2004, Loretta Goetsch, Andrew Eckert and Benjamin Hall of the University of Washington proposed and in 2005 published a recommendation that the Genus Rhododendron be revised and subdivided into five subgenera:
 
  
  1. Rhododendron (lepidotes)
+
==Classification==
  2. Hymenanthes (elepidotes & deciduous azaleas)
+
The genus ''Rhododendron'' was created by Linnaeus in 1753, but he placed the [[azalea]]s in another [[genus]]. However, it was later recognized that a separation between azaleas and the members of ''Rhododendron'' could not be justified and they were placed together. The ''Rhododendron'' genus was subdivided into eight sections in 1834 by George Don and these eight subcategories have been commonly recognized until today, with the azaleas comprising two of the sections, subgenus ''Pentanthera'' (typified by ''Rhododendron nudiflorum'') and subgenus ''Tsutsusi'' (typified by ''R. Tsutsusi'') (Henning 2007).
  3. Azaleastrum (evergreen azaleas)
 
  4. Choniastrum (R. campionae, R. honcockii, R. latoucheae, R. moulmainense, R. stamineum)
 
  5. Therorhodion (R. camtschaticum)
 
http://www.ingentaconnect.com/content/aspt/sb/2005/00000030/00000003/art00012
 
  
The Molecular Systematics of Rhododendron (Ericaceae): A Phylogeny Based Upon RPB2 Gene Sequences
+
However, other subcategories also have been recognized. It is not uncommon for nine subgenera to be recognized, with section Vireya within the large subgenera ''Rhododendron'' being raised to the subgenus level (Callard 2008). Furthermore, in 2004, based on genetic and chemical data, Goetsch et al. (2005) called for a new realignment in which the genus ''Rhododendron'' was subdivided into five subgenera (''Rhododendron, Hymenanthes, Azaleastrum, Choniastrum'' and ''Therorhodion''). Essentially, they call for the elimination of three subgenera and two sections that are present in the taxonomic system of Chamberlain et al. (1996) and the inclusiong of inclusion of section Pentanthera within subgenus Hymenanthes.
Authors: Goetsch, Loretta; Eckert, Andrew J.; Hall, Benjamin D.
 
Source: Systematic Botany, Volume 30, Number 3, July-September 2005 , pp. 616-626(11)
 
  
 +
[[Image:Rhododendron wardii var puralbum.jpg|thumb|250px|right|''Rhododendron wardii var. puralbum'']]
  
[[Image:Rhododendron wardii var puralbum.jpg|thumb|250px|right|''Rhododendron wardii var. puralbum'']]
+
Overall, the species today are organized by subgenus, section, subsection and series.  
The species are organized by subgenus, section, subsection and series. These are currently divided into four large and four small subgenera:
+
The traditional recognition of eight subgenera divides the genus into four large and four small subgenera:
 
*Subgenus ''[[Rhododendron subgenus Rhododendron|Rhododendron]]'' L.: small leaf or lepidotes (with scales on the underside of their leaves); several hundred species, type: ''[[Rhododendron ferrugineum]]''.  
 
*Subgenus ''[[Rhododendron subgenus Rhododendron|Rhododendron]]'' L.: small leaf or lepidotes (with scales on the underside of their leaves); several hundred species, type: ''[[Rhododendron ferrugineum]]''.  
 
**The tropical rhododendrons (sect. ''[[Rhododendron subgenus Vireya|Vireya]]'', about 300 species) are usually included as a section in this subgenus, but sometimes split off as a ninth subgenus.
 
**The tropical rhododendrons (sect. ''[[Rhododendron subgenus Vireya|Vireya]]'', about 300 species) are usually included as a section in this subgenus, but sometimes split off as a ninth subgenus.
Line 73: Line 68:
 
*Subgenus ''[[Rhododendron subgenus Therorhodion|Therorhodion]]'': one species, ''[[Rhododendron camtschaticum]]''.  
 
*Subgenus ''[[Rhododendron subgenus Therorhodion|Therorhodion]]'': one species, ''[[Rhododendron camtschaticum]]''.  
  
Chris Callard  2008  recognizes nine subgenera Subgenus Vireya
+
Recent genetic investigations have caused an ongoing realignment of species and groups within the genus, and also have caused the old genus ''[[Ledum]]'' to be reclassified within subgenus ''Rhododendron''. As noted, further realignment within subgenera is currently proposed by Goetsch et al. (2005).
http://www.vireya.net/classification.htm
 
he Classification of Vireya Rhododendrons
 
  
Recent genetic investigations have caused an ongoing realignment of species and groups within the genus, and also have caused the old genus ''[[Ledum]]'' to be reclassified within subgenus ''Rhododendron''. Further realignment within the subgenera is currently proposed <ref name="Goetsch et al.">Goetsch, L. A., Eckert, A. J. & Hall, B. D. (2005). [http://www.ingentaconnect.com/content/aspt/sb/2005/00000030/00000003/art00012 The molecular systematics of Rhododendron (Ericaceae): A Phylogeny based upon RPB2 gene sequences]. ''Sys. Bot''. 30(3): 616-626.</ref><ref>[http://www.flounder.ca/FraserSouth/Goetsch-Eckert-Hall.asp Summary of Goetsch-Eckert-Hall results]</ref>, including the merging of subgenus ''Hymenanthes'' into subgenus ''Pentanthera''.  
+
Rhododendrons are extensively hybridized in cultivation, and natural hybrids often occur in areas where species ranges overlap. There are over 28,000 [[cultivar]]s of rhododendron in the [http://www.rhs.org.uk/plants/registration_rhododendron.asp International Rhododendron Registry] held by the [[Royal Horticultural Society]]. Most have been bred for their flowers, but a few are of garden interest because of ornamental leaves and some for ornamental bark or stems.
  
Rhododendrons are extensively hybridized in cultivation, and natural hybrids often occur in areas where species ranges overlap. There are over 28,000 [[cultivar]]s of Rhododendron in the [http://www.rhs.org.uk/plants/registration_rhododendron.asp International Rhododendron Registry] held by the [[Royal Horticultural Society]]. Most have been bred for their flowers, but a few are of garden interest because of ornamental leaves and some for ornamental bark or stems.
+
[[Image:Garden with Rhododendrons.JPG|500px|thumb|right|| A garden with tall rhododendrons in Lynnwood, Washington]]
 
 
Some species (e.g. ''[[Rhododendron ponticum]]'' in the [[United Kingdom]]) are invasive as introduced plants, spreading in woodland areas replacing the natural understory. R. ponticum is difficult to eradicate, as its roots can make new shoots.
 
 
 
[[Image:Garden with Rhododendrons.JPG|500px|thumb|right|| A garden with tall Rhododendrons in Lynnwood, Washington]]
 
  
 
;Sample species:
 
;Sample species:
Line 108: Line 97:
  
 
==Toxicology==
 
==Toxicology==
<!-- Image with unknown copyright status removed: [[Image:Bee Collecting Pollen From Rhododendron.JPG|500px|thumb|right|| A Bee Collecting Pollen from a Rhododendron Flower]] —>
+
Some rhododendron species, including azaleas, are poisonous to grazing animals. These rhododendrons have a [[toxin]] called [[grayanotoxin]] in their [[pollen]] and [[nectar (plant)|nectar]]. People have been known to become ill from eating [[honey]] made by [[bee]]s feeding on rhododendron and azalea flowers. [[Xenophon]] described the odd behavior of [[Greeks|Greek]] soldiers after having consumed honey in a village surrounded by rhododendrons. Later, it was recognized that honey resulting from these plants have a slightly [[hallucinogenic]] and [[laxative]] effect.
  
Some species are poisonous to grazing animals. These Rhododendrons have a [[toxin]] called [[grayanotoxin]] in their [[pollen]] and [[nectar (plant)|nectar]]. People have been known to become ill from eating [[honey]] made by [[bee]]s feeding on rhododendron and azalea flowers. [[Xenophon]] described the odd behavior of [[Greeks|Greek]] soldiers after having consumed honey in a village surrounded by rhododendrons. Later, it was recognized that honey resulting from these plants have a slightly [[hallucinogenic]] and [[laxative]] effect.{{Fact|date=July 2008}}  The suspect rhododendrons are ''[[Rhododendron ponticum]]'' and ''[[Rhododendron luteum]]'' (formerly ''Azalea pontica''), both found in northern [[Asia Minor]]. Eleven similar cases have been documented in [[Istanbul]], [[Turkey]] during the 1980s <ref name="Poison honey">Nurhayat Sütlüpmar, Afife Mat and Yurdagül Satganoglu ''[http://www.springerlink.com/content/w322484735r06208/ Poisoning by toxic honey in Turkey]''. Archives of Toxicology. Volume 67, Number 2, pages 148-150, February, 1993 </ref>. Rhododendron is extremely toxic to horses, with some animals dying within a few hours of ingesting the plant, although most horses tend to avoid it if they have access to good forage.
+
The suspect rhododendrons are ''[[Rhododendron ponticum]]'' and ''[[Rhododendron luteum]]'' (formerly ''Azalea pontica''), both found in northern [[Asia Minor]]. Eleven similar cases have been documented in [[Istanbul]], [[Turkey]] during the 1980s (Sütlüpmar et al. 1993).  
 +
 
 +
Rhododendron is extremely toxic to horses, with some animals dying within a few hours of ingesting the plant, although most horses tend to avoid it if they have access to good forage.
  
 
==Use in landscaping==
 
==Use in landscaping==
Both species and hybrid rhododendrons (including azaleas) are used extensively as [[ornamental plant]]s in [[landscaping]] in many parts of the world, and many species and cultivars are grown commercially for the [[Nursery (horticulture)|nursery]] trade. Rhododendrons are often valued in landscaping for their structure, size, flowers, and the fact that many of them are evergreen <ref name=Huxley>Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan</ref>. Azaleas are frequently used around foundations and occasionally as hedges, and many larger-leafed rhododendrons lend themselves well to more informal plantings and woodland gardens, or as specimen plants. In some areas, larger rhododendrons can be pruned to encourage more tree-like form, with some species such as ''[[Rhododendron arboreum|R. arboreum]]'' and ''[[Rhododendron falconeri|R. falconeri]]'' eventually growing to 10-15 m or more tall. <ref name=Huxley/>
+
Both species and hybrid [[rhododendron]]s, including [[azalea]]s, are used extensively as [[ornamental plant]]s in [[landscaping]] in many parts of the world, and many species and [[cultivar]]s are grown commercially for the [[Nursery (horticulture)|nursery]] trade. Rhododendrons are often valued in landscaping for their structure, size, flowers, and the fact that many of them are evergreen (Huxley 1992). Azaleas are frequently used around foundations and occasionally as hedges, and many larger-leafed rhododendrons lend themselves well to more informal plantings and woodland gardens, or as specimen plants. In some areas, larger rhododendrons can be pruned to encourage more tree-like form, with some species such as ''[[Rhododendron arboreum|R. arboreum]]'' and ''[[Rhododendron falconeri|R. falconeri]]'' eventually growing to ten to 15 meters (33-49 feet) or more tall (Huxley 1992).
  
 
===Commercial growing===
 
===Commercial growing===
Rhododendrons are grown commercially in many areas for sale, and are occasionally collected in the wild, a practice now rare in most areas. Larger commercial growers often ship long distances; in the [[United States]] most of them are located on the west coast ([[Oregon]], [[Washington]] and [[California]]). Large-scale commercial growing often selects for different characteristics than hobbyist growers might, such as resistance to root rot when over-watered, ability to be forced into budding early, ease of rooting or other propagation, and saleability. <ref name=PCox>Peter A. Cox (1993). ''The Cultivation of Rhododendrons''. B. T. Batsford, London  ISBN 0-7134-5630-2 (pp80-1) </ref>
+
Rhododendrons are grown commercially in many areas for sale, and are occasionally collected in the wild, a practice now rare in most areas. Larger commercial growers often ship long distances; in the [[United States]] most of them are located on the west coast ([[Oregon]], [[Washington]], and [[California]]). Large-scale commercial growing often selects for different characteristics than hobbyist growers might, such as resistance to root rot when over-watered, ability to be forced into budding early, ease of rooting or other propagation, and saleability (Cox 1993).  
  
 
===Planting and care===
 
===Planting and care===
Like other [[Ericaceae|ericaceous]] plants, most rhododendrons prefer acid soils with a pH of roughly 4.5-5.5; some tropical Vireyas and a few other rhododendron species grow as [[epiphyte]]s and require a planting mix similar to [[orchids]]. Rhododendrons have fibrous roots and prefer well-drained soils high in organic material. In areas with poorly-drained or alkaline soils, rhododendrons are often grown in raised beds using mediums such as composted pine bark.<ref>[http://www.rhododendron.org/soil.htm Soil information for planting rhododendrons]</ref> Mulching and careful watering are important, especially before the plant is established.
+
Like other [[Ericaceae|ericaceous]] plants, most rhododendrons prefer acid soils with a pH of roughly 4.5 to 5.5. Rhododendrons have fibrous roots and prefer well-drained [[soil]]s high in organic material. In areas with poorly-drained or alkaline soils, rhododendrons are often grown in raised beds using mediums such as composted pine bark. Mulching and careful watering are important, especially before the plant is established. Some tropical Vireyas and a few other rhododendron species grow as [[epiphyte]]s and require a planting mix similar to [[orchid]]s.  
  
 
===Insects and diseases===
 
===Insects and diseases===
{{Main|List of rhododendron diseases}}
 
 
There are a number of insects that either target rhododendrons or will opportunistically attack them.  Rhododendron borers and various weevils are major pests of rhododendrons, and many caterpillars will attack rhododendrons.  Major diseases include ''[[Phytophthora]]'' root rot, stem and twig fungal dieback; [[Ohio State University Extension]] provides [http://www.ag.ohio-state.edu/~ohioline/hyg-fact/3000/3043.html information on maintaining health of rhododendrons]Rhododendrons can easily be suffocated by other plants.
 
 
''Rhododendron'' species are used as food plants by the [[larva]]e of some members of the Order [[Lepidoptera]] ([[butterflies]] and [[moths]]) (See [[List of Lepidoptera that feed on rhododendrons]]).
 
  
 +
There are a number of insects that either target rhododendrons or will opportunistically attack them. Rhododendron borers and various weevils are major pests of rhododendrons, and many [[caterpillar]]s will attack rhododendrons. Major diseases include ''[[Phytophthora]]'' root rot, stem and twig fungal dieback. ''Rhododendron'' species are used as food plants by the [[larva]]e of some members of the Order [[Lepidoptera]] ([[butterflies]] and [[moths]]).
  
 
== Gallery ==
 
== Gallery ==
 
<gallery>
 
<gallery>
Image:Rhododendron decorum ssp diaprepes -1.jpg|''Rhododendron decorum'' subsp. ''diaprepes''
 
 
Image:rhododendron2.jpg
 
Image:rhododendron2.jpg
Image:Wild Rhododendrons in Kashmir.jpg|Wild Rhododendrons in Kashmir by [[Edward Molyneux]]; painted before 1908
+
Image:Wild Rhododendrons in Kashmir.jpg|Wild rhododendrons in Kashmir by [[Edward Molyneux]]; painted before 1908
 
Image:Rhododendron 1 Fcb981.JPG
 
Image:Rhododendron 1 Fcb981.JPG
Image:Wild Rhododendrons in Whitwick Leicestershire 1155.JPG | Flowering Rhododendrons in Whitwick Leicestershire
+
Image:Wild Rhododendrons in Whitwick Leicestershire 1155.JPG | Flowering rhododendrons in Whitwick Leicestershire
 
</gallery>
 
</gallery>
  
 
==References==
 
==References==
*Cox, P. A. & Kenneth, N. E. ''The Encyclopedia of Rhododendron Species''. 1997. Glendoick Publishing. ISBN 0-9530533-0-X.
+
* Argent, G. 2006. ''Rhododendrons of subgenus Vireya''. Royal Horticultural Society. ISBN 1902896610.
*Davidian, H. H. ''The Rhododendron Species''. In four volumes from 1982-1995. Timber Press. ISBN 0-917304-71-3, ISBN 0-88192-109-2, ISBN 0-88192-168-8, ISBN 0-88192-311-7.
+
* Callard, C. 2008. [http://www.vireya.net/classification.htm The classification of vireya rhododendrons] ''Vireya''. Retrieved December 18, 2008.
 +
* Cox, P.A. 1993. ''The Cultivation of Rhododendrons''. London: B. T. Batsford. ISBN 0713456302.
 +
* Cox, P.A., and N.E. Kenneth. 1997. ''The Encyclopedia of Rhododendron Species''. Glendoick Publishing. ISBN 095305330X.
 +
* Davidian, H.H. 1982-1985. ''The Rhododendron Species''. In four volumes from 1982-1995. Timber Press. ISBN 0917304713.
 +
* Francis, R. 2006. Tree rhododendrons ''Wildeel.com''.
 +
* Fraser South Rhododendron Society. n.d. Excerpts from and comments on a paper published by: Loretta Goetsch, Andrew Eckert and Benjamin Hall ''Fraser South Rhododendron Society''.
 +
* Goetsch, L.A., A.J. Eckert, and B.D. Hall. 2005. [http://www.ingentaconnect.com/content/aspt/sb/2005/00000030/00000003/art00012 The molecular systematics of rhododendron (Ericaceae): A phylogeny based upon RPB2 gene sequences] ''Sys. Bot''. 30(3): 616-626. Retrieved December 18, 2008.
 +
* Henning. 2007. [http://www.rhodyman.net/rhodyhis.html#anchor282700 Rhododendron history] ''Henning Home Page''. Retrieved December 18, 2008.
 +
* Huxley, A. (ed.). 1992. ''The New Royal Horticultural Society Dictionary of Gardening''. London: Macmillan. ISBN 0333770188.
 +
* Sütlüpmar, N., A. Mat, and Y. Satganoglu. 1993. [http://www.springerlink.com/content/w322484735r06208/ Poisoning by toxic honey in Turkey] ''Archives of Toxicology'' 67(2): 148-150. Retrieved December 18, 2008.
  
<ref>
+
==External links==
Richard Francis 2006
+
All links retrieved December 8, 2022.
[http://wildeel.com/rhodotree.html Tree rhododendrons]</ref>.  
 
  
==External links==
 
* [http://www.lahana.org/blog/Rhododendrons.htm Rhododendrons from Turkey, Anatolia]
 
 
* [http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=128386 Flora of China: ''Rhododendron'']
 
* [http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=128386 Flora of China: ''Rhododendron'']
* [http://www.countrysideinfo.co.uk/rhododen.htm Description of damage caused by Rhododendrons in the UK]
 
 
* [http://www.wfu.edu/~kronka/kakbek/rhododendrons.html Information on rhododendrons at the Ericaceae web pages of Dr. Kron at Wake Forest University.]
 
* [http://www.wfu.edu/~kronka/kakbek/rhododendrons.html Information on rhododendrons at the Ericaceae web pages of Dr. Kron at Wake Forest University.]
* [http://www.vireya.net/ Information on Vireyas]
 
* [http://www.rhodophiles.com/ Information on Rhododendrons by Marc Colombel, founder of the Société Bretonne du Rhododendron.]
 
* [http://rhodyman.net/rahome.html Extensive information on rhododendron species: the history of their discovery, botanical details, toxicity, classification, cultural conditions, care for common problems, and suggestions for companion plants by Steve Henning.]
 
* [http://www.rhodo.citymax.com/page/page/627471.htm History of Rhododendrons]
 
* [http://scholar.lib.vt.edu/ejournals/JARS/v23n4/v23n4-black.htm Historical Survey of Rhododendron Collecting - With Emphasis on its Close Associations with Horticulture]
 
  
===Rhododendron Societies===
 
* [http://rhododendron.org/ American Rhododendron Society]
 
* [http://www.rhs.org.uk/plants/plant_groups/rhododendrons.asp UK Royal Horticultural Society]
 
* [http://www.rhodygarden.org/ Rhododendron Species Foundation and Botanical Garden]
 
* [http://honeybee.helsinki.fi/users/avainola/rhodo/club.htm Société Finlandaise du Rhododendron]
 
* [http://www.eisa.net.au/~mirra/australi.html Australian Rhododendron Society]
 
* [http://www.rhodo.org/wissen.php?lang=gb German Rhododendron Society]
 
  
 
[[Category:Life sciences]]
 
[[Category:Life sciences]]

Latest revision as of 20:52, 16 April 2023

Rhododendron
Rhododendron ponticum
Rhododendron ponticum
Scientific classification
Kingdom: Plantae
Division: Magnoliophyta
Class: Magnoliopsida
Order: Ericales
Family: Ericaceae
Genus: Rhododendron
L.
Subgenera
  • Azaleastrum
  • Candidastrum
  • Hymenanthes
  • Mumeazalea
  • Pentanthera (Azaleas)
  • Rhododendron
  • Therorhodion
  • Tsutsusi (Azaleas)
  • Vireya

Rhododendron is the common and genus name for a large and diverse group of woody shrubs and small (rarely large) trees in the flowering plant family Ericaceae. It is a large genus with over 1000 species, and subdivided into various subgenera (commonly eight, nine, or five, depending on the taxonomic scheme). The trees may be deciduous or evergreen and the spirally arranged leaves can vary significantly in size according to species, from one to two centimeters (0.4-0.8 inches) up to 100 centimeters (39 inches). Members of two of the traditional subgenera, Pentanthera (deciduous), and Titsushi (evergreen), are known as azaleas.

Rhododendrons, including azaleas, generally have showy flower displays. Indeed, the name rhododendron comes from the Greek rhodos, meaning "rose," and dendron, meaning "tree." As such, and combined with their often being evergreen, they often are cultivated as ornamental plants, such as around the foundations of houses or in woodland gardens, and even as specimen plants.

They also provide important ecological values. They provide food for a number of insects and an important source of pollen and nectar for many bumblebees, who in turn pollinate the plants. However, rhododendrons also are poisonous to many grazing animals, such as horses.

Some species (e.g. Rhododendron ponticum in the United Kingdom) are invasive as introduced plants, spreading in woodland areas replacing the natural understory. R. ponticum is difficult to eradicate, as its roots can make new shoots.

Overview and description

Rhododendron decorum subsp. diaprepes

Ericaceae, the family to which the genus Rhododendron belongs, is known as the "heath family," and consists of herbs, shrubs, and trees with leaves that are alternate, simple, and estipulate, and mostly lime-hating or calcifuge plants that thrive in acid soils. In addition to rhododendrons, well-known examples include cranberry, blueberry, heath, heather, and huckleberry.

The Rhododendron genus is characterized by shrubs and small to (rarely) large trees, the smallest species growing to ten to 100 centimeters (3.9-39 inches) tall, and the largest, R. giganteum, reported to over 30 meters (98 feet) tall (Francis 2006). The leaves are spirally arranged; leaf size can range from one to two centimeters (0.4-0.8 inches) to over 50 centimeters (20 inches), exceptionally 100 centimeters (39.4 inches) in R. sinogrande. They may be either evergreen or deciduous. In some species the underside of the leaves is covered with scales (lepidote) or hairs (indumentum).

Some of the best known species of rhododendrons are noted for their many clusters of large flowers. There are alpine species with small flowers and small leaves, and tropical species such as section Vireya that often grow as epiphytes.

Some rhododendrons are known by the common name of azalea. These are flowering shrubs that are characterized by the lack of scales on the underside of the generally thin, soft, and pointed leaves, and typically having terminal blooms (one flower per stem), flowers with five or six stamens, and with long straight hairs parallel to the leaf surface and along the midrid on the ventral surface of the leaf. Originally, azaleas were classed as a different genus of plant, but now they are recognized as part of the rhododendrons. In the common classification of rhododendrons into eight or nine subgenera, azaleas comprise two of the subgenera: Pentanthera (deciduous) and Titsushi (evergreen).

Distribution

Rhododendron is a very widely distributed genus, occurring throughout most of the Northern Hemisphere except for dry areas, and extending into the Southern Hemisphere in southeastern Asia and northern Australasia.

The highest species diversity is found in the Himalayan mountains from Uttarakhand, Nepal, and Sikkim to Yunnan and Sichuan, with other significant areas of diversity in the mountains of Indo-China, Korea, Japan, and Taiwan. Rhododendron, also known as "Lali Guras" in local language, is the national flower of Nepal and the state tree of the state of Uttarakhand in India. In Korea, Rhododendron schlippenbachii is particularly common (called 철쭉 (choltjuk)). In addition, there are a significant number of tropical rhododendron species from southeast Asia to northern Australia, with 55 known species in Borneo and 164 in New Guinea (Argent 2006).

Relatively fewer species occur in North America and Europe. Rhododendrons have not been found in South America or Africa.

Classification

The genus Rhododendron was created by Linnaeus in 1753, but he placed the azaleas in another genus. However, it was later recognized that a separation between azaleas and the members of Rhododendron could not be justified and they were placed together. The Rhododendron genus was subdivided into eight sections in 1834 by George Don and these eight subcategories have been commonly recognized until today, with the azaleas comprising two of the sections, subgenus Pentanthera (typified by Rhododendron nudiflorum) and subgenus Tsutsusi (typified by R. Tsutsusi) (Henning 2007).

However, other subcategories also have been recognized. It is not uncommon for nine subgenera to be recognized, with section Vireya within the large subgenera Rhododendron being raised to the subgenus level (Callard 2008). Furthermore, in 2004, based on genetic and chemical data, Goetsch et al. (2005) called for a new realignment in which the genus Rhododendron was subdivided into five subgenera (Rhododendron, Hymenanthes, Azaleastrum, Choniastrum and Therorhodion). Essentially, they call for the elimination of three subgenera and two sections that are present in the taxonomic system of Chamberlain et al. (1996) and the inclusiong of inclusion of section Pentanthera within subgenus Hymenanthes.

Rhododendron wardii var. puralbum

Overall, the species today are organized by subgenus, section, subsection and series. The traditional recognition of eight subgenera divides the genus into four large and four small subgenera:

  • Subgenus Rhododendron L.: small leaf or lepidotes (with scales on the underside of their leaves); several hundred species, type: Rhododendron ferrugineum.
    • The tropical rhododendrons (sect. Vireya, about 300 species) are usually included as a section in this subgenus, but sometimes split off as a ninth subgenus.
  • Subgenus Hymenanthes (Blume) K.Koch: large leaf or elepidotes (without scales on the underside of their leaves); about 140 species, type: Rhododendron degronianum.
  • Subgenus Pentanthera G.Don: deciduous azaleas; about 25 species, type Rhododendron luteum.
  • Subgenus Tsutsusi: evergreen azaleas, about 15 species; type Rhododendron indicum.
  • Subgenus Azaleastrum Planch.: five species; type Rhododendron ovatum.
  • Subgenus Candidastrum (Sleumer) Philipson & Philipson: one species; Rhododendron albiflorum.
  • Subgenus Mumeazalea: one species, Rhododendron semibarbatum.
  • Subgenus Therorhodion: one species, Rhododendron camtschaticum.

Recent genetic investigations have caused an ongoing realignment of species and groups within the genus, and also have caused the old genus Ledum to be reclassified within subgenus Rhododendron. As noted, further realignment within subgenera is currently proposed by Goetsch et al. (2005).

Rhododendrons are extensively hybridized in cultivation, and natural hybrids often occur in areas where species ranges overlap. There are over 28,000 cultivars of rhododendron in the International Rhododendron Registry held by the Royal Horticultural Society. Most have been bred for their flowers, but a few are of garden interest because of ornamental leaves and some for ornamental bark or stems.

A garden with tall rhododendrons in Lynnwood, Washington
Sample species
  • Rhododendron atlanticum
  • Rhododendron canadense
  • Rhododendron catawbiense
  • Rhododendron chapmanii
  • Rhododendron ferrugineum
  • Rhododendron groenlandicum
  • Rhododendron hirsutum
  • Rhododendron lochiae
  • Rhododendron luteum
  • Rhododendron macrophyllum
  • Rhododendron maximum
  • Rhododendron moulmainense
  • Rhododendron occidentale
  • Rhododendron ponticum
  • Rhododendron schlippenbachii
  • Rhododendron spinuliferum
  • Rhododendron tomentosum

A sample hybrid:

Toxicology

Some rhododendron species, including azaleas, are poisonous to grazing animals. These rhododendrons have a toxin called grayanotoxin in their pollen and nectar. People have been known to become ill from eating honey made by bees feeding on rhododendron and azalea flowers. Xenophon described the odd behavior of Greek soldiers after having consumed honey in a village surrounded by rhododendrons. Later, it was recognized that honey resulting from these plants have a slightly hallucinogenic and laxative effect.

The suspect rhododendrons are Rhododendron ponticum and Rhododendron luteum (formerly Azalea pontica), both found in northern Asia Minor. Eleven similar cases have been documented in Istanbul, Turkey during the 1980s (Sütlüpmar et al. 1993).

Rhododendron is extremely toxic to horses, with some animals dying within a few hours of ingesting the plant, although most horses tend to avoid it if they have access to good forage.

Use in landscaping

Both species and hybrid rhododendrons, including azaleas, are used extensively as ornamental plants in landscaping in many parts of the world, and many species and cultivars are grown commercially for the nursery trade. Rhododendrons are often valued in landscaping for their structure, size, flowers, and the fact that many of them are evergreen (Huxley 1992). Azaleas are frequently used around foundations and occasionally as hedges, and many larger-leafed rhododendrons lend themselves well to more informal plantings and woodland gardens, or as specimen plants. In some areas, larger rhododendrons can be pruned to encourage more tree-like form, with some species such as R. arboreum and R. falconeri eventually growing to ten to 15 meters (33-49 feet) or more tall (Huxley 1992).

Commercial growing

Rhododendrons are grown commercially in many areas for sale, and are occasionally collected in the wild, a practice now rare in most areas. Larger commercial growers often ship long distances; in the United States most of them are located on the west coast (Oregon, Washington, and California). Large-scale commercial growing often selects for different characteristics than hobbyist growers might, such as resistance to root rot when over-watered, ability to be forced into budding early, ease of rooting or other propagation, and saleability (Cox 1993).

Planting and care

Like other ericaceous plants, most rhododendrons prefer acid soils with a pH of roughly 4.5 to 5.5. Rhododendrons have fibrous roots and prefer well-drained soils high in organic material. In areas with poorly-drained or alkaline soils, rhododendrons are often grown in raised beds using mediums such as composted pine bark. Mulching and careful watering are important, especially before the plant is established. Some tropical Vireyas and a few other rhododendron species grow as epiphytes and require a planting mix similar to orchids.

Insects and diseases

There are a number of insects that either target rhododendrons or will opportunistically attack them. Rhododendron borers and various weevils are major pests of rhododendrons, and many caterpillars will attack rhododendrons. Major diseases include Phytophthora root rot, stem and twig fungal dieback. Rhododendron species are used as food plants by the larvae of some members of the Order Lepidoptera (butterflies and moths).

Gallery

References
ISBN links support NWE through referral fees

External links

All links retrieved December 8, 2022.

Credits

New World Encyclopedia writers and editors rewrote and completed the Wikipedia article in accordance with New World Encyclopedia standards. This article abides by terms of the Creative Commons CC-by-sa 3.0 License (CC-by-sa), which may be used and disseminated with proper attribution. Credit is due under the terms of this license that can reference both the New World Encyclopedia contributors and the selfless volunteer contributors of the Wikimedia Foundation. To cite this article click here for a list of acceptable citing formats.The history of earlier contributions by wikipedians is accessible to researchers here:

The history of this article since it was imported to New World Encyclopedia:

Note: Some restrictions may apply to use of individual images which are separately licensed.