Difference between revisions of "Buffalo" - New World Encyclopedia

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The word '''buffalo''' refers to certain animals of the Bovidae, [[bovid]], family:
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[[Image:African Buffalo.jpg|400px|thumb|Herd of African buffalo]]
  
* African buffalo, ''Syncerus caffer''.
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'''Buffalo''' (plural: buffalo, buffaloes, or, less commonly, buffalos) is the common name for certain animals of the Bovidae family ([[bovid]]s):
* Asian buffalo, ''Bubalus'', a genus that includes the domesticated water buffalo.
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* It is also sometimes used to refer to the American [[bison]], ''Bison bison''.
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* African buffalo, ''Syncerus caffer.''
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* Asian buffalo, ''Bubalus,'' a genus that includes the domesticated water buffalo.
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* It is also sometimes used to refer to the [[American bison]], ''Bison bison.''
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All of these are even-toed [[ungulate]]s (hoofed [[mammal]]s). They are all ruminants, which means that they have complex digestive systems that allow them to live on a diet of [[grass]], [[leaf|leaves]], and other low-grade [[plant]] material, utilizing [[symbiosis|symbiotic]] microorganisms to break down [[cellulose]], while providing shelter and food for the microbes. All buffalo belong to the subfamily Bovinae, along with domestic [[cattle]], ''Bos taurus''.
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The domesticated water buffalo, a type of Asian buffalo, provides great value to [[human]]s, including pulling carts, pulling plows to prepare [[rice]] fields, and providing milk, meat, and leather, and fertilizer. In India, more milk is produced from water buffalo than from cattle. Unlike the African buffalo, which has not been domesticated and can be dangerous to humans, the water buffalo have been known to be docile and form bonds with their human owners.
  
 
==African Buffalo==
 
==African Buffalo==
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{{Taxobox_phylum_entry | taxon = [[Chordate|Chordata]]}}
 
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{{Taxobox_classis_entry | taxon = [[mammal|Mammalia]]}}
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{{Taxobox_ordo_entry | taxon = [[Ungulate#Artiodactyla: Even-toed ungulates|Artiodactyla]]}}
 
{{Taxobox_familia_entry | taxon = [[Bovidae]]}}
 
{{Taxobox_familia_entry | taxon = [[Bovidae]]}}
 
{{Taxobox_genus_entry | taxon = '''''Syncerus'''''}}<br/>{{Taxobox authority | author = [[Brian Houghton Hodgson|Hodgson]] | date = 1847}}
 
{{Taxobox_genus_entry | taxon = '''''Syncerus'''''}}<br/>{{Taxobox authority | author = [[Brian Houghton Hodgson|Hodgson]] | date = 1847}}
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The '''African buffalo''' or '''Cape buffalo''' (''Syncerus caffer'') is a large animal; 1 to 1.7 meters (3 to 5.5 feet) high at the shoulder, 2 to 3 meters (7 to 10 feet) long, and with a weight of 500 to 1000 kilograms (1,100 to 2,200 lbs). Males are somewhat larger than females. Both sexes have large horns. Newborn African buffalo are covered with thick reddish hair which becomes thinner and darker with age so that mature animals are dark brown or black (Nowak 1983).
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The '''African buffalo''' or '''Cape buffalo''' (''Syncerus caffer'') is a large animal, 1 to 1.7 meters (3 to 5.5 feet) high at the shoulder, 2 to 3 meters (7 to 10 feet) long, and with a weight of 500 to 1000 kilograms (1,100 to 2,200 lbs). Males are somewhat larger than females. Both sexes have large horns. Newborn African buffalo are covered with thick reddish hair, which becomes sparser and darker with age so that mature animals are dark brown or black (Nowak 1983).
[[Image:African_Buffalo_Drawing_historic.jpg|left|thumb|African Buffalo, 19th century drawing]]
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The Cape Buffalo is not closely related to the Asian [[Water Buffalo]], but its ancestry remains unclear. It is a very powerful creature, demanding respect from even a pride of [[lion]]s when paths cross. Other than humans, they have few natural [[predator]]s and are capable of defending themselves against (and sometimes killing) [[lion]]s. Lions do kill and eat buffalo occasionally, but it typically takes multiple lions to bring down a single adult buffalo; fewer when it is injured or very old. The [[leopard]] and [[spotted hyena]] are a threat only to newborn calves. Crossbreeding with domestic [[cattle]] has had only limited success, and the African Buffalo remains a wild animal.
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African buffalo are found over much of [[Africa]] south of the [[Sahara Desert]], in habitats ranging from open savanna (grassland with some trees) to thick [[forest]]s. Those living in forests tend to be smaller than those living on the savannas. Some of the nations in which African buffalo are now found are [[Ethiopia]], [[Somalia]], [[Zambia]], [[Zimbabwe]], [[Namibia]], [[Botswana]], [[Mozambique]], [[South Africa]], [[Kenya]], and [[Tanzania]].
Known as one of the "[[Big Five Game|big five]]" in Africa, the Cape Buffalo can be a volatile and formidable beast, goring, trampling and killing several people a year. It is reputedly the most dangerous game animal, with the possible exception of the [[hippopotamus]].
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African buffalo tend to live in herds, although some older males live alone. Forest living buffalo herds most commonly contain around a dozen animals; while on the savannas herds are mostly in the range of 50 to 500, with groups of up to 2,000 or 3,000 sometimes coming together in good grazing conditions (Huffman 2006).
Cape Buffalo occur from open savannah to thickly wooded country, and wallow when the opportunity presents itself. They are found in [[Ethiopia]], [[Somalia]], [[Zambia]], [[Zimbabwe]], [[Namibia]], [[Botswana]], [[Mozambique]], [[South Africa]], [[Kenya]] and [[Tanzania]].
 
  
The main herd consists of both sexes and all ages, though bachelor groups are also found. A male is recognizable by the thickness of his horns, and is called the "Boss." Bulls mature at eight years of age. Cows first calve at five years of age, after a [[gestation]] period of 11.5 months. Older bulls forced out of herds are commonly found in bachelor groups.
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[[Image:African_Buffalo_Drawing_historic.jpg|left|thumb|African Buffalo, nineteenth-century drawing]]
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African buffalo are formidable fighters. Both males and females will attack and kill [[lion]]s and other predators using both their horns and hooves. Many human hunters have also been killed by them. Because of this, sports hunters consider them one of the "Big Five" African animals most dangerous and challenging to hunt. The others are [[elephant]]s, lions, [[rhinoceros]]es, and [[leopard]]s.  
  
==Bubalus==
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Although in the past the population of African buffalo was in the tens of millions, it now stands at about 900,000; mostly in the savannas of eastern Africa. Some of the reasons for this decline in numbers are hunting, both for food and for sport; the conversion of their habitat to farmland and [[cattle]] pasture; droughts; and the introduction of foreign pests and [[diseases]], especially the cattle disease rinderpest. It is not considered to be in danger of [[extinction]] as long as it remains protected in parks and reserves, although habitat loss continues (Huffman 2006; IUCN 2006).
'''''Bubalus''''' is a genus of [[bovine]]s, the English name of which is buffalo. Species that belong to this genus are:
 
  
* '''Subgenus Bubalus'''
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The African buffalo has never been successfully domesticated.
** [[Water Buffalo]], ''Bubalus bubalus''
 
** ''[[Bubalus cebuensis]]'' (extinct)
 
** [[Tamaraw]], ''Bubalus mindorensis''
 
* '''Subgenus Anoa'''
 
** [[Anoa|Lowland Anoa]], ''Bubalus depressicornis''
 
** [[Anoa|Mountain Anoa]], ''Bubalus quarlesi''
 
  
Some [[zoologist]]s split the Water Buffalo into two species.
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==Asian buffalo==
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'''Asian buffalo''' are members of the genus ''Bubalus.'' They are:
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* Subgenus Bubalus
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** '''Water Buffalo,''' ''Bubalus bubalus''
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** '''Tamaraw,''' ''Bubalus mindorensis''
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* Subgenus Anoa
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** '''Lowland Anoa,''' ''Bubalus depressicornis''
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** '''Mountain Anoa,''' ''Bubalus quarlesi''
  
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These are described below.
  
 
===Water buffalo===
 
===Water buffalo===
 
{{Taxobox_begin | color = pink | name = Water Buffalo}}
 
{{Taxobox_begin | color = pink | name = Water Buffalo}}
{{Taxobox_image | image = [[Image:Water buffalo bathing.jpg|240px]] | caption = A family of Water Buffalo bathing <br/>in a sinkhole, [[Taiwan]]}}
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{{Taxobox_image | image = [[Image:Two asiatic water buffalos in zoo tierpark friedrichsfelde berlin germany.jpg|240px]] | caption = Domestic Water Buffalo}}
 
{{Taxobox_begin_placement | color = pink}}
 
{{Taxobox_begin_placement | color = pink}}
 
{{Taxobox_regnum_entry | taxon = [[Animal]]ia}}
 
{{Taxobox_regnum_entry | taxon = [[Animal]]ia}}
 
{{Taxobox_phylum_entry | taxon = [[Chordate|Chordata]]}}
 
{{Taxobox_phylum_entry | taxon = [[Chordate|Chordata]]}}
 
{{Taxobox_classis_entry | taxon = [[mammal|Mammalia]]}}
 
{{Taxobox_classis_entry | taxon = [[mammal|Mammalia]]}}
{{Taxobox_ordo_entry | taxon = [[Artiodactyla]]}}
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{{Taxobox_ordo_entry | taxon = [[Ungulate#Artiodactyla: Even-toed ungulates|Artiodactyla]]}}
 
{{Taxobox_familia_entry | taxon = [[Bovidae]]}}
 
{{Taxobox_familia_entry | taxon = [[Bovidae]]}}
 
{{Taxobox_subfamilia_entry | taxon = [[Bovinae]]}}
 
{{Taxobox_subfamilia_entry | taxon = [[Bovinae]]}}
{{Taxobox_genus_entry | taxon = '''''[[Bubalus]]'''''}}
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{{Taxobox_genus_entry | taxon = '''''Bubalus'''''}}
 
{{Taxobox_species_entry | taxon = '''''B. bulbalis'''''}}
 
{{Taxobox_species_entry | taxon = '''''B. bulbalis'''''}}
 
{{Taxobox_end_placement}}
 
{{Taxobox_end_placement}}
{{Taxobox_section_binomial_parens | color = pink | binomial_name = Bubalus arnee | author = [[Robert Kerr (writer)|Kerr]] | date = 1792}}
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{{Taxobox_section_binomial_parens | color = pink | binomial_name = Bubalus bulbalis | author = [[Robert Kerr (writer)|Kerr]] | date = 1792}}
 
{{Taxobox_end}}
 
{{Taxobox_end}}
  
The '''Water Buffalo''' is a very large [[ungulate]] and a member of the [[bovinae|bovine subfamily]]. The '''Arni''' or Wild Buffalo survives in the wild in [[Pakistan]], [[India]], [[Bangladesh]], [[Nepal]], [[Bhutan]], [[Vietnam]], [[China]], [[Philippine]], [[Taiwan]], [[Indonesia]], and [[Thailand]]. The [[domestication|domestic]] Buffalo are very widespread  in [[Asia]], [[South America]], [[North Africa]] and [[Europe]]. They are [[feral]] in northern [[Australia]]. Wild-living populations also exist in much of [[South-east Asia]] but their origins are uncertain. They may be descendants of wild Water Buffalo, formerly domesticated ferals or a mixture. In Asia, the population of wild Water Buffalo has become sparse, and there is fear that no pure-bred wild water buffalo exists. Adult Water Buffalo range in size from 300 kg to 600 kg for the domestic bred, but true wild water buffaloes can grow larger. Adult females typically weigh around 800 kg, and males up to 1200 kg. The average weight of an adult male is 900 kg, and the average height at the shoulder is about 1.7m. One distinction between wild buffaloes and domesticated version is that the wild species lacks the round belly found in domesticated buffalo. Due to interbreeding with feral and domestic species, weights can vary greatly even in populations which are in in close proximity.  Interbreeding with domesticated buffaloes is the major cause of extinction of wild buffaloes. Buffaloes are believed to have originated in [[South Asia]].
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The wild '''water buffalo,''' sometimes called the '''Arni,''' ''Bubalus bulbalis,'' is somewhat larger than the African buffalo. An adult male typically stands 1.5 to 1.9 meters (5 to 6.25 feet) tall at the shoulder and weighs 700 to 1200 kg (1,500 to 2,600 lbs), while females are smaller. The spread of their horns is the largest of any living [[ungulate]]. They are covered with sparse hair and range in color from light gray to black. Their skin is very thick, which helps to protect them from [[insect]]s and from thorns (Nowak 1983). The wild water buffalo is native to the area of [[India]] and Southeast [[Asia]], and has been domesticated.  
 
 
The classification of the Water Buffalo is uncertain. Some authorities list a single [[species]], ''Bubalus bubalis'' with three [[subspecies]], the River Buffalo (''B. bubalis bubalis'') of South Asia and the [[Carabao]] or Swamp Buffalo (''B. bubalis carabanesis'') of the [[Philippines]] and Southeast Asia, and the Arni, or Wild Buffalo, (''B. bubalis arnee''). Others regard these as closely-related, but separate, species. The Swamp Buffalo is primarily found in the eastern half of Asia and has 48 chromosomes. The River Buffalo is mostly found in western half of Asia, and has 50 chromosomes. Fertile offspring occur between the two. It does not readily hybridise with [[cattle]] which have 60 chromosomes.
 
  
Domestic animal expert Juliet Clutton-Brock wrote (Clutton-Brock 1999):
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Wild water buffalo prefer to live near freshwater and mostly live in marshland and along the banks of [[river]]s. They are good swimmers and spend much of their time in the water, which keeps them cool and protects them from biting insects. They can graze on water plants while swimming and even dive underwater to feed on them (Massicot 2004). Their feet are large and the toes can spread apart to give them better footing in mud.
  
:More than half the people of the world depend on rice for their staple diet and it is the water buffalo that enables rice to be cultivated and threshed with the greatest efficiency and economy.
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[[Image:Carabao.jpg|200px|thumb|left|A Carabao, a smaller breed of water buffalo found in the Philippines]]
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Wild water buffalo are now very rare. It is thought that less than 4,000 remain and their numbers are declining. Some of the problems are illegal hunting, loss of habitat, introduced [[disease]]s and parasites, and interbreeding with domesticated buffalo (IUCN 2006).
  
[[Milk]] from both of these animals is used by many human populations, and is the traditional raw material for [[mozzarella cheese]] and [[curd]] due to its higher fat content. The chief dairy breed of Buffalo is the [[Murrah Buffalo|Murrah breed]]. Buffalo meat, sometimes called "Carabeef", is often passed off as beef in certain regions and is also a major source of export revenue for India which has the largest population of buffaloes in the world. However, in many Asian regions, buffalo meat is less preferred due to its toughness. Water buffalo hide provides a tough and useful [[leather]] often used for shoes and [[motorcycle]] [[helmets]].
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The water buffalo was first domesticated in Southeast Asia, at least 6,000 years ago; it is not sure exactly when since little [[archaeology|archaeological]] evidence has been found. It seems that buffalo were used early for the plowing of [[rice]] fields and the threshing of rice. Domestic animal expert Juliet Clutton-Brock wrote (1999):
  
[[Image:Indonesia-Bull.jpg|thumbnail|left|Water Buffalo in Indonesia]]Buffaloes, apart from its use as [[draught]] and [[milch]] animals, are also used to pull bullock carts in developing world. Their dung is used as a fertilizer and as a fuel when dried. Sometimes, people will also race on buffaloes, though riding them can be very difficult. A few have also found use as pack animals carrying loads even for special forces.
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<blockquote>More than half the people of the world depend on rice for their staple diet and it is the water buffalo that enables rice to be cultivated and threshed with the greatest efficiency and economy.</blockquote>
  
'''Asia'''. Asia is the native home of the water buffalo, with 95% of the world population of water buffalo. Many Asian countries depend on the water buffalo as its primary bovine species. It is valuable for its meat and milk as well as the labour it performs. As of 1992 the Asian population was estimated at 141 million. The fat content of buffalo milk is the highest amongst farm animals and the butterfat is a major source of [[ghee]] in some Asian countries. Its success in Asia is evident by its extensive range. Both variants occur in Asia. River Buffalo are found in elevations of 2,800 m in Nepal, and Swamp Buffalo are found throughout the lowland tropics. Part of their success is due to their ability to thrive on poor foodstuffs and yet be valuable economically. Moreover they are much better suited to plough the muddy [[paddy]] fields as they are better adapted than common cattle (''[[Bos taurus]]'') to move in swamps.
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Water buffalo changed under domestication. Over many generations they became smaller so that today their average weight is around 300 to 600 kg (650 to 1,300 lbs). Their bodies became rounder and their horns smaller. They also became slower and more docile. Water buffalo form a bond with their human owners and can be herded even by children. There are even stories of water buffalo protecting their masters by fending off attacks by [[crocodile]]s and [[tiger]]s (Olson 2006).
  
In the wild, very few pockets of buffalo exist. The Indian wild buffalo is found in India, Pakistan, Bangladesh, Nepal, Bhutan and Thailand. This buffalo has the largest horns of any living animal. The average spread is about 1 m (3 ft 3 in), but one bull shot in 1955 had horns measuring 4.24 m (13 ft 11 in) from tip to tip along the outside curve across the forehead. These massive battering rams protect the beast in the wild efficiently enough that even its predator seldom attacks a fully grown male. Tiger is their only predator, which can kill a full-grown male weighing 1000-1200kg, but only an experienced tiger will take them on, often by hamstringing their hind legs first. When faced by a tiger, they form a line facing the predator and charge with noses out and horns laid back.  Like its other family members, the [[tamaraw]] and the [[anoa]], the wild buffalo is a very dangerous animal known to charge without being provoked. Together with its large size, this bad temper makes wild water buffalo the most dangerous bovine on earth to humans. It is generally found in swamplands and grass jungles and moves in herds.  
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Like [[cattle]], domestic water buffalo have many uses. They have been used to pull carts and carry loads, although they travel more slowly than oxen, [[donkey]]s, and [[horse]]s and need to stop and get into water to cool off on hot days. They have provided [[milk]] and meat for food and leather, bone, and horn for materials. Their manure is used to fertilize the rice fields and is used for fuel.
  
Today, the estimated population of wild water buffaloes are about 4000, but, this number takes into account all wild population, including feral herds and hybrid buffaloes. In fact, it is possible that no true wild specimens exist anymore.  
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[[Image:Water buffalo bathing.jpg|thumb|left|Water buffalo bathing]]
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Over the years, domestic water buffaloes spread around the world to areas to which they were suited, mainly lowlands in tropical and subtropical areas with wet climates, especially rice growing areas. By 4,500 years ago, they were found in [[India]] and around the year 600 they were introduced to [[Egypt]] and southern [[Europe]]. In more recent times, they have become common in the [[Philippines]] and other Pacific islands and in [[South America]], especially [[Brazil]]. In the early 1800s, they were introduced to [[Australia]] where many escaped or were abandoned and gave rise to a large feral population in the northern part of the country.
  
[[Image:Army buffalo.jpg|thumb|A buffalo in service with an unidentified army's special forces]]
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[[Image:Asiatic water buffalo in zoo tierpark friedrichsfelde berlin germany.jpg|thumb|Water buffalo]]
The water buffalo, or "[[Carabao]]," is the national animal of the Philippines.
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There are two main types of domestic water buffalo, the ''swamp buffalo'' type originally from East Asia and the ''river buffalo'' type from India. Swamp buffalo have straighter, more swept back horns than river buffalo, more like wild water buffalo, and they prefer to wallow in mud, while river buffalo prefer clear running water (as their names suggest). River buffalo are used more for milk production than for work.  
  
'''Australia'''. Introduced into the [[Northern Territory]] early in the 19th century as a beast of burden, it quickly escaped and is now [[feral]]. As a result of its feral status it may be hunted. [[Melville Island, Northern Territory|Melville Island]] is a popular hunting location, where a steady population of up to 4,000 individuals exist.  Buffaloes are also found in [[Arnhem Land]] and the Top End. Safari outfits run out of [[Darwin, Northern Territory|Darwin]] to Melville Island and other locations in the [[Top End]] often with the use of [[bush pilot]]s. The government has unsuccessfully attempted several eradication programs. The buffaloes live mainly in freshwater marshes and [[billabong]]s, and their range can be quite expansive during the [[Wet season]]. They have developed a different appearance from the Indonesian buffaloes from which they descend.
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Buffalo milk is richer than cow's milk and is traditionally used in [[Italy]] to make mozzarella [[cheese]]. In traditional societies, water buffalo were not killed for meat unless they were unable to work because of age or injury. In these cases, the meat was tough and not very palatable; however, when buffalo are raised expressly for meat, the product compares with that of [[cattle]]. Water buffalo leather is very thick and strong and is used to make, among other things, shoes and motorcycle helmets (Olson 2006).
  
'''Europe and Middle East'''. Introduced into North Africa and the Near East by [[600]] AD, the water buffalo was brought to Europe with returning [[Crusade]]rs in the [[Middle Ages]], and herds can be found in [[Bulgaria]] and [[Italy]]. As in Asia, buffaloes of the [[Middle East]] and [[Europe]] live on coarse vegetation on the marginal land traditionally available to [[peasants]]. They are an economic asset by serving as a protein source, draft animal, and storage of family or household wealth. In some areas, they also provide occasional recreation at annual racing festivals. These buffalo are mostly River Buffaloes; due to genetic isolation have adopted a distinct appearance. Buffalo milk is used in Italy and elsewhere for the production of [[cheese]], including [[mozzarella]].
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In 1992, the [[United Nations]] Food and Agriculture Organization (FAO) estimated the world water buffalo population at 148 million. About 95 percent live in Asia, with India and [[China]] having the largest number. In India, more milk for human consumption is produced by water buffalo than by cattle. In [[Thailand]], the Philippines, and [[Malaysia]], the number of water buffalo seems to be decreasing, while in most other southeastern Asian countries it is increasing. In the [[United States]], except for [[Hawaii]], water buffalo have only recently been introduced as farm animals and number in the few thousands (Ligda 1996).
  
 
===Tamaraw===
 
===Tamaraw===
The '''Tamaraw''' (''Bubalus mindorensis''; previously ''Anoa mindorensis''), '''Tamarao''' or '''Mindoro Dwarf Buffalo''' is a [[bovine]] [[endemic (ecology)|endemic]] to the island of [[Mindoro]] in the [[Philippines]]. It is believed, however, to have once also thrived on the greater island of [[Luzon]].  It was originally found all over Mindoro, from sea level up to the mountains, but because of human habitation, hunting, and logging, it is now restricted to only a few remote grassy plains and is now an [[endangered species]].
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The '''tamaraw,''' ''Bubalus mindorensis''' (also '''tamarau,''' '''tamarao,''' or '''Mindoro Dwarf Buffalo''') is native to the island of Mindoro in the [[Philippines]]. It is believed, however, to have once also thrived on the greater island of Luzon.  
  
Contrary to common belief and past classification, the Tamaraw is not a subspecies of the local [[Carabao]], which is only slightly larger, or the common [[Water Buffalo]]. In contrast to the Carabao, it has a number of distinguishing characteristics: it is slightly hairier, has light markings on its face, is not gregarious, and has shorter horns that are somewhat V-shaped.
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The tamaraw was originally found all over Mindoro, from sea level up to the mountains, but because of human habitation, hunting, and logging, it is now restricted to only a few remote grassy plains and is now an [[endangered species]]. In 2000, its population was estimated to be less than 200 (IUCN 2006).
  
Though the national animal of the Philippines is the Carabao, the Tamaraw is closely considered a national symbol of the Philippines. It symbolizes sturdiness amidst a small frame, which most Filipinos possess. An image of the Tamaraw could be found on the 1980 to early 1990 One-[[Philippine Peso|Peso]] coins.
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The tamaraw is smaller than other buffalo, about one meter (3 feet) tall at the shoulder and weighing 200 to 300 kg (450 to 650 lbs). They have more hair than water buffalo and are dark brown to grayish black in color. Their horns are swept back and about 35 to 45 cm (14 to 18 inches) long. They tend to live alone, with the exception of mothers and young, rather than in herds. They can be aggressive and have been know to attack humans (Huffman 2006).
 
 
During the wake of the Asian van popularity in 1990s, [[Toyota]] Motors released an Asian van called [[Tamaraw FX]] in the Philippines. It was widely patronized by taxi operators and was immediately turned into a staple mode of transportation much like a cross of the taxi and the local [[jeepney]].
 
 
 
The tamaraw is also the mascot of the varsity teams of the [[Far Eastern University]] ([[FEU Tamaraws]]) in the [[University Athletics Association of the Philippines]], and of the [[Toyota Super Corollas|Toyota Tamaraws]] of the [[Philippine Basketball Association]].
 
  
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The tamaraw is considered a national symbol of the [[Philippines]]. It symbolizes sturdiness in a small frame, which Filipinos are said to possess. The tamaraw was pictured on the Philippine one peso coin from 1980 to the early 1990s.
  
 
===Anoa===
 
===Anoa===
There are two species of '''anoa''': the '''Mountain Anoa''' (''Bubalus quarlesi'') and the '''Lowland Anoa''' (''Bubalus depressicornis''). Both live in undisturbed forest, and although they are essentially miniature [[Water Buffalo]], are similar in appearance to a deer, weighing 150&ndash;300&nbsp;kg (330&ndash;660&nbsp;lb).  
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[[image:Lowland_Anoa.JPG|thumb|Lowland anoa]]
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There are two species of '''anoa''': the '''mountain anoa''' (''Bubalus quarlesi'') and the '''lowland anoa''' (''Bubalus depressicornis''). Both live in undisturbed forest, and are similar in appearance to a [[deer]], weighing 150 to 300 kg (330 to 660 lbs). Because of their small size (the lowland anoa is only 1 meter tall), they are sometimes called pygmy buffalo or miniature water buffalo.
  
Both are found on the island of [[Sulawesi]] in [[Indonesia]]; the Mountain Anoa is also found on the nearby island of [[Butung]]. They apparently live singly or in pairs, rather than herds, except when the cows are about to give birth. One young is born per year.
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Both are found on the island of Sulawesi in [[Indonesia]]; the mountain anoa is also found on the nearby island of Butung. They apparently live singly or in pairs, rather than herds, except when the females are about to give birth. One young is born per year.
[[image:Lowland_Anoa.JPG|thumb|left|250px|A Lowland Anoa]]
 
  
Both species of anoa have been classified as endangered since the 1960s and the population continues to decrease. It is believed unlikely that there are more than 5000 animals of each species remaining. Reasons for the decline of the anoa include hunting for hide, horns and meat by the indigenous peoples (though they were rarely hunted by indigenous peoples before the introduction of modern firearms); and loss of habitat due to the advancement of settlement. Currently, hunting is the more serious factor in most areas. The anoa do not appear to be adaptable to humans.
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Both species of anoa have been classified as [[endangered species|endangered]] since the 1960s and the population continues to decrease. It is believed unlikely that there are more than 5,000 animals of each species remaining. Reasons for the decline of the anoa include hunting for hide, horns, and meat by the indigenous peoples (though they were rarely hunted by indigenous peoples before the introduction of modern firearms); and loss of habitat due to the advancement of settlement. Currently, hunting is the more serious factor in most areas.
  
Mountain Anoa are also known as Anoa de Montana, Anoa de Quarle, Anoa des Montagnes, Anoa Pegunungan, and Quarle's Anoa. Lowland Anoa are also known as Anoa de Ilanura or Anoa des Plaines.
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The lowland anoa stands one meter (3 feet) tall at the shoulder, and is the smallest of all wild buffalo. The horns of anoa are peculiar for their upright direction and comparative straightness, although they have the same triangular section as in other buffalo. White spots are sometimes present below the eyes, and there may be white markings on the legs and back; and the absence or presence of these white markings may be indicative of distinct races. The horns of the females are very small. The nearest relatives of the anoa appear to be certain [[extinction|extinct]] buffalo, of which the remains are found in the Siwalik Hills of northern [[India]].
 
 
Lowland Anoa stands but little over a yard at the shoulder, and is the most diminutive of all wild cattle. It is nearly allied to the larger Asiatic buffaloes, showing the same reversal of the direction of the hair on the back. The horns are peculiar for their upright direction and comparative straightness, although they have the same triangular section as in other buffaloes. White spots are sometimes present below the eyes, and there may be white markings on the legs and back; and the absence or presence of these white markings may be indicative of distinct races. The horns of the cows are very small. The nearest allies of the anoa appear to be certain extinct buffaloes, of which the remains are found in the Siwalik Hills of northern India. In habits the animal appears to resemble the Indian buffalo.  
 
  
 
==References==
 
==References==
* Clutton-Brock, J. 1999. ''A Natural History of Domesticated Mammals''. Cambridge UK : Cambridge University Press. ISBN 0521634954.
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* Clutton-Brock, J. 1999. ''A Natural History of Domesticated Mammals.'' Cambridge, UK: Cambridge University Press. ISBN 0521634954
* Huffman, B. 2006. [http://www.ultimateungulate.com ''The ultimate ungulate page'']. ''UltimateUngulate.com''. Retrieved (date).  
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* Huffman, B. 2006. [http://www.ultimateungulate.com The Ultimate Ungulate Page.] ''UltimateUngulate.com.'' Retrieved January 9, 2007.  
* International Union for Conservation of Nature and Natural Resources (IUCN). 2006. [http://www.iucnredlist.org/search/details.php/21251/all ''Syncerus caffer''], [http://www.iucnredlist.org/search/details.php/3129/summ ''Bubalus bubalis''], [http://www.iucnredlist.org/search/details.php/3128/all ''Bubalus quarlesi''], [http://www.iucnredlist.org/search/details.php/3126/doc ''Bubalus depressicornis''] [http://www.iucnredlist.org/search/details.php/3127/all ''Bubalus mindorensis'']. Retrieved  
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* International Union for Conservation of Nature and Natural Resources (IUCN). 2006. [http://www.iucnredlist.org/search/details.php/21251/all ''Syncerus caffer''], [http://www.iucnredlist.org/search/details.php/3129/summ ''Bubalus bubalis''], [http://www.iucnredlist.org/search/details.php/3128/all ''Bubalus quarlesi''], [http://www.iucnredlist.org/search/details.php/3126/doc ''Bubalus depressicornis''], [http://www.iucnredlist.org/search/details.php/3127/all ''Bubalus mindorensis'']. Retrieved January 9, 2007.
* Ligda, D.J. 1996. ''The Water Buffalo''. Website. [http://ww2.netnitco.net/users/djligda/waterbuf.htm]
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* Ligda, D. J. 1996. [http://ww2.netnitco.net/users/djligda/waterbuf.htm ''The Water Buffalo.''] Retrieved January 9, 2007.
* Massicot, P. 2004. Animal Info. Website [http://www.animalinfo.org/species/artiperi/bubaarne.htm Wild Asian (Water) Buffalo]
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* Massicot, P. 2004. [http://www.animalinfo.org/species/artiperi/bubaarne.htm Wild Asian (Water) Buffalo.] ''Animalinfo.com.'' Retrieved January 9, 2007.
* Nowak, R.M. and Paradiso, J.L. 1983. ''Walker's Mammals of the World''. Baltimore, Maryland, USA: The Johns Hopkins University Press. ISBN 0801825253
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* Nowak, R. M., and J. L. Paradiso. 1983. ''Walker's Mammals of the World.'' Baltimore: Johns Hopkins University Press. ISBN 0801825253
* Olson, T.J. 2006. ''Turkey Creek Water Buffalo'' [http://home.valornet.com/pcwdb] Accessed
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* Olson, T. J. 2006. [http://home.valornet.com/pcwdb ''Turkey Creek Water Buffalo.''] Retrieved January 9, 2007.
* Raudiansky, S. 1992. ''The Covenant of the Wild''. New York: William Morrow and Company, Inc. ISBN 0688096107.
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* Voelker, W. 1986. ''The Natural History of Living Mammals.'' Medford, NJ: Plexus Publishing. ISBN 0937548081
* Voelker, W. 1986. ''The Natural History of Living Mammals''. Medford, New Jersey, USA: Plexus Publishing, Inc. ISBN 0937548081.
 
 
 
  
 
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{{credit5|African_Buffalo|86857788|Bubalus|88479826|Water_Buffalo|88705572|Tamaraw|88705526|Anoa|88705552}}
 
[[Category:Life sciences]]
 
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[[Category:Animals]]
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[[Category:Mammals]][[Category:Ungulates]]

Latest revision as of 21:28, 6 February 2020

Herd of African buffalo

Buffalo (plural: buffalo, buffaloes, or, less commonly, buffalos) is the common name for certain animals of the Bovidae family (bovids):

  • African buffalo, Syncerus caffer.
  • Asian buffalo, Bubalus, a genus that includes the domesticated water buffalo.
  • It is also sometimes used to refer to the American bison, Bison bison.

All of these are even-toed ungulates (hoofed mammals). They are all ruminants, which means that they have complex digestive systems that allow them to live on a diet of grass, leaves, and other low-grade plant material, utilizing symbiotic microorganisms to break down cellulose, while providing shelter and food for the microbes. All buffalo belong to the subfamily Bovinae, along with domestic cattle, Bos taurus.

The domesticated water buffalo, a type of Asian buffalo, provides great value to humans, including pulling carts, pulling plows to prepare rice fields, and providing milk, meat, and leather, and fertilizer. In India, more milk is produced from water buffalo than from cattle. Unlike the African buffalo, which has not been domesticated and can be dangerous to humans, the water buffalo have been known to be docile and form bonds with their human owners.

African Buffalo

African Buffalo
Conservation status: Lower risk (cd)
African Buffalo
African Buffalo
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Bovidae
Genus: Syncerus
Hodgson, 1847
Species: S. caffer
Binomial name
Syncerus caffer
(Sparrman, 1779)
Subspecies

S. c. caffer
S. c. nanus
S. c. brachyceros
S. c. mathewsi

The African buffalo or Cape buffalo (Syncerus caffer) is a large animal, 1 to 1.7 meters (3 to 5.5 feet) high at the shoulder, 2 to 3 meters (7 to 10 feet) long, and with a weight of 500 to 1000 kilograms (1,100 to 2,200 lbs). Males are somewhat larger than females. Both sexes have large horns. Newborn African buffalo are covered with thick reddish hair, which becomes sparser and darker with age so that mature animals are dark brown or black (Nowak 1983).

African buffalo are found over much of Africa south of the Sahara Desert, in habitats ranging from open savanna (grassland with some trees) to thick forests. Those living in forests tend to be smaller than those living on the savannas. Some of the nations in which African buffalo are now found are Ethiopia, Somalia, Zambia, Zimbabwe, Namibia, Botswana, Mozambique, South Africa, Kenya, and Tanzania.

African buffalo tend to live in herds, although some older males live alone. Forest living buffalo herds most commonly contain around a dozen animals; while on the savannas herds are mostly in the range of 50 to 500, with groups of up to 2,000 or 3,000 sometimes coming together in good grazing conditions (Huffman 2006).

African Buffalo, nineteenth-century drawing

African buffalo are formidable fighters. Both males and females will attack and kill lions and other predators using both their horns and hooves. Many human hunters have also been killed by them. Because of this, sports hunters consider them one of the "Big Five" African animals most dangerous and challenging to hunt. The others are elephants, lions, rhinoceroses, and leopards.

Although in the past the population of African buffalo was in the tens of millions, it now stands at about 900,000; mostly in the savannas of eastern Africa. Some of the reasons for this decline in numbers are hunting, both for food and for sport; the conversion of their habitat to farmland and cattle pasture; droughts; and the introduction of foreign pests and diseases, especially the cattle disease rinderpest. It is not considered to be in danger of extinction as long as it remains protected in parks and reserves, although habitat loss continues (Huffman 2006; IUCN 2006).

The African buffalo has never been successfully domesticated.

Asian buffalo

Asian buffalo are members of the genus Bubalus. They are:

  • Subgenus Bubalus
    • Water Buffalo, Bubalus bubalus
    • Tamaraw, Bubalus mindorensis
  • Subgenus Anoa
    • Lowland Anoa, Bubalus depressicornis
    • Mountain Anoa, Bubalus quarlesi

These are described below.

Water buffalo

Water Buffalo
Two asiatic water buffalos in zoo tierpark friedrichsfelde berlin germany.jpg
Domestic Water Buffalo
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Bovidae
Subfamily:: Bovinae
Genus: Bubalus
Species: B. bulbalis
Binomial name
Bubalus bulbalis
(Kerr, 1792)

The wild water buffalo, sometimes called the Arni, Bubalus bulbalis, is somewhat larger than the African buffalo. An adult male typically stands 1.5 to 1.9 meters (5 to 6.25 feet) tall at the shoulder and weighs 700 to 1200 kg (1,500 to 2,600 lbs), while females are smaller. The spread of their horns is the largest of any living ungulate. They are covered with sparse hair and range in color from light gray to black. Their skin is very thick, which helps to protect them from insects and from thorns (Nowak 1983). The wild water buffalo is native to the area of India and Southeast Asia, and has been domesticated.

Wild water buffalo prefer to live near freshwater and mostly live in marshland and along the banks of rivers. They are good swimmers and spend much of their time in the water, which keeps them cool and protects them from biting insects. They can graze on water plants while swimming and even dive underwater to feed on them (Massicot 2004). Their feet are large and the toes can spread apart to give them better footing in mud.

A Carabao, a smaller breed of water buffalo found in the Philippines

Wild water buffalo are now very rare. It is thought that less than 4,000 remain and their numbers are declining. Some of the problems are illegal hunting, loss of habitat, introduced diseases and parasites, and interbreeding with domesticated buffalo (IUCN 2006).

The water buffalo was first domesticated in Southeast Asia, at least 6,000 years ago; it is not sure exactly when since little archaeological evidence has been found. It seems that buffalo were used early for the plowing of rice fields and the threshing of rice. Domestic animal expert Juliet Clutton-Brock wrote (1999):

More than half the people of the world depend on rice for their staple diet and it is the water buffalo that enables rice to be cultivated and threshed with the greatest efficiency and economy.

Water buffalo changed under domestication. Over many generations they became smaller so that today their average weight is around 300 to 600 kg (650 to 1,300 lbs). Their bodies became rounder and their horns smaller. They also became slower and more docile. Water buffalo form a bond with their human owners and can be herded even by children. There are even stories of water buffalo protecting their masters by fending off attacks by crocodiles and tigers (Olson 2006).

Like cattle, domestic water buffalo have many uses. They have been used to pull carts and carry loads, although they travel more slowly than oxen, donkeys, and horses and need to stop and get into water to cool off on hot days. They have provided milk and meat for food and leather, bone, and horn for materials. Their manure is used to fertilize the rice fields and is used for fuel.

Water buffalo bathing

Over the years, domestic water buffaloes spread around the world to areas to which they were suited, mainly lowlands in tropical and subtropical areas with wet climates, especially rice growing areas. By 4,500 years ago, they were found in India and around the year 600 they were introduced to Egypt and southern Europe. In more recent times, they have become common in the Philippines and other Pacific islands and in South America, especially Brazil. In the early 1800s, they were introduced to Australia where many escaped or were abandoned and gave rise to a large feral population in the northern part of the country.

Water buffalo

There are two main types of domestic water buffalo, the swamp buffalo type originally from East Asia and the river buffalo type from India. Swamp buffalo have straighter, more swept back horns than river buffalo, more like wild water buffalo, and they prefer to wallow in mud, while river buffalo prefer clear running water (as their names suggest). River buffalo are used more for milk production than for work.

Buffalo milk is richer than cow's milk and is traditionally used in Italy to make mozzarella cheese. In traditional societies, water buffalo were not killed for meat unless they were unable to work because of age or injury. In these cases, the meat was tough and not very palatable; however, when buffalo are raised expressly for meat, the product compares with that of cattle. Water buffalo leather is very thick and strong and is used to make, among other things, shoes and motorcycle helmets (Olson 2006).

In 1992, the United Nations Food and Agriculture Organization (FAO) estimated the world water buffalo population at 148 million. About 95 percent live in Asia, with India and China having the largest number. In India, more milk for human consumption is produced by water buffalo than by cattle. In Thailand, the Philippines, and Malaysia, the number of water buffalo seems to be decreasing, while in most other southeastern Asian countries it is increasing. In the United States, except for Hawaii, water buffalo have only recently been introduced as farm animals and number in the few thousands (Ligda 1996).

Tamaraw

The tamaraw,' Bubalus mindorensis (also tamarau, tamarao, or Mindoro Dwarf Buffalo) is native to the island of Mindoro in the Philippines. It is believed, however, to have once also thrived on the greater island of Luzon.

The tamaraw was originally found all over Mindoro, from sea level up to the mountains, but because of human habitation, hunting, and logging, it is now restricted to only a few remote grassy plains and is now an endangered species. In 2000, its population was estimated to be less than 200 (IUCN 2006).

The tamaraw is smaller than other buffalo, about one meter (3 feet) tall at the shoulder and weighing 200 to 300 kg (450 to 650 lbs). They have more hair than water buffalo and are dark brown to grayish black in color. Their horns are swept back and about 35 to 45 cm (14 to 18 inches) long. They tend to live alone, with the exception of mothers and young, rather than in herds. They can be aggressive and have been know to attack humans (Huffman 2006).

The tamaraw is considered a national symbol of the Philippines. It symbolizes sturdiness in a small frame, which Filipinos are said to possess. The tamaraw was pictured on the Philippine one peso coin from 1980 to the early 1990s.

Anoa

Lowland anoa

There are two species of anoa: the mountain anoa (Bubalus quarlesi) and the lowland anoa (Bubalus depressicornis). Both live in undisturbed forest, and are similar in appearance to a deer, weighing 150 to 300 kg (330 to 660 lbs). Because of their small size (the lowland anoa is only 1 meter tall), they are sometimes called pygmy buffalo or miniature water buffalo.

Both are found on the island of Sulawesi in Indonesia; the mountain anoa is also found on the nearby island of Butung. They apparently live singly or in pairs, rather than herds, except when the females are about to give birth. One young is born per year.

Both species of anoa have been classified as endangered since the 1960s and the population continues to decrease. It is believed unlikely that there are more than 5,000 animals of each species remaining. Reasons for the decline of the anoa include hunting for hide, horns, and meat by the indigenous peoples (though they were rarely hunted by indigenous peoples before the introduction of modern firearms); and loss of habitat due to the advancement of settlement. Currently, hunting is the more serious factor in most areas.

The lowland anoa stands one meter (3 feet) tall at the shoulder, and is the smallest of all wild buffalo. The horns of anoa are peculiar for their upright direction and comparative straightness, although they have the same triangular section as in other buffalo. White spots are sometimes present below the eyes, and there may be white markings on the legs and back; and the absence or presence of these white markings may be indicative of distinct races. The horns of the females are very small. The nearest relatives of the anoa appear to be certain extinct buffalo, of which the remains are found in the Siwalik Hills of northern India.

References
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