Difference between revisions of "Starfish" - New World Encyclopedia
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{{Taxobox | {{Taxobox | ||
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| subdivision_ranks = [[Order (biology)|Orders]] | | subdivision_ranks = [[Order (biology)|Orders]] | ||
| subdivision = | | subdivision = | ||
| − | Brisingida (100 species<ref name=fossilgroups>[http://palaeo.gly.bris.ac.uk/Palaeofiles/Fossilgroups/asteroz2/index_f/mod_fm.html | + | Brisingida (100 species<ref name=fossilgroups>Sweet, E. 2005. [http://palaeo.gly.bris.ac.uk/Palaeofiles/Fossilgroups/asteroz2/index_f/mod_fm.html Modern Forms: Asteroids]. ''University of Bristol''. Retrieved July 31, 2007.</ref>)<br/> |
| − | Forcipulatida (300 species<ref name=fossilgroups/>)<br> | + | Forcipulatida (300 species<ref name=fossilgroups/>)<br/> |
| − | Paxillosida (255 species | + | Paxillosida (255 species<ref name=fossilgroups/>)<br/> |
| − | Notomyotida (75 species<ref name=fossilgroups/>)<br> | + | Notomyotida (75 species<ref name=fossilgroups/>)<br/> |
| − | Spinulosida (120 species<ref name=fossilgroups/>)<br> | + | Spinulosida (120 species<ref name=fossilgroups/>)<br/> |
| − | Valvatida (695 species<ref name=fossilgroups/>)<br> | + | Valvatida (695 species<ref name=fossilgroups/>)<br/> |
Velatida (200 species<ref name=fossilgroups/>) | Velatida (200 species<ref name=fossilgroups/>) | ||
}} | }} | ||
| − | '''Starfish''' | + | '''Starfish,''' or '''sea stars''' (a less confusing designation, since they are only very distantly related to [[fish]]), are marine [[invertebrate]]s belonging to the kingdom [[animalia]], phylum [[Echinoderm]]ata, and class '''Asteroidea,''' of which there are over 1800 known species. The names sea star and starfish also are (incorrectly) used for the closely related [[brittle star]]s, which make up the class Ophiuroidea. |
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| + | Characteristics of sea stars include that they exhibit a superficially [[symmetry (biology)#Radial symmetry|radial symmetry]], since they typically have five or more "arms" which radiate from an indistinct disk ([[symmetry (biology)#Special forms of radial symmetry|pentaradial symmetry]]), and tend to be covered in spines. Like other [[echinoderm]]s, starfish possess an endoskeleton, but do not rely on it for support and locomotion, instead using a hydraulic [[water]] vascular system that functions via many projections called tube feet, located on the ventral surface of the starfish's arms. | ||
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| + | Starfish are an essential part of marine [[food chain]]s, being eaten as larvae, and becoming voracious predators upon reaching adulthood, with their diet including [[mollusk]]s and other marine [[organism]]s. They even are able to evert their [[stomach]] in order to pull apart and digest their shelled prey. For humans, they are a source of aesthetic joy and their remarkable characteristics—movement based on a hydraulics system, ability to regenerate lost limbs and evert their stomach—add to the wonder of nature. Some starfish also are captured, dried out, and turned into popular souvenirs or decorations. | ||
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| + | While starfish can reproduce sexually, if an arm of the starfish is severed from the body and it contains a portion of the central [[nerve]] ring, then another starfish can regrow from the severed portion. Lack of knowledge of this capacity increased the role of starfish as pests to commercial [[clam]] and [[oyster]] beds when fisherman, in an effort to get rid of these predators, would cut them up and throw them back into the ocean. | ||
==Distribution== | ==Distribution== | ||
[[Image:Sandstar 300.jpg|thumb|left|200px|Giant sea star, ''Pisaster giganteus'']] | [[Image:Sandstar 300.jpg|thumb|left|200px|Giant sea star, ''Pisaster giganteus'']] | ||
| − | There are about 1,800 known living species of starfish, and | + | There are about 1,800 known living species of starfish, and they occur in all of the [[Earth]]'s oceans (though the greatest variety of starfish is found in the tropical Indo-Pacific). Habitats range from tropical [[coral]] reefs to kelp forests to the deep-sea floor, although none live within the [[water]] column; all species of starfish found are living as benthos (or on the ocean floor). |
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| + | Because [[echinoderm]]s need to maintain a delicate internal balance in their body, no starfish are found in freshwater environments. | ||
==External anatomy== | ==External anatomy== | ||
| − | [[Image:red-knobbed.starfish.arp.jpg| | + | [[Image:red-knobbed.starfish.arp.jpg|right|thumb|200px|Red-knobbed starfish ''Protoreaster linckii'', a starfish from the Indian Ocean]] |
| − | Starfish are composed of a central disc from which arms sprout in pentaradial symmetry | + | Starfish are composed of a central disc from which arms (usually five, but the number can vary) sprout in [[symmetry (biology)#Special forms of radial symmetry|pentaradial symmetry]]. The mouth is located underneath the starfish on the oral or ventral surface, while the anus is located on the top of the [[animal]]. The spiny upper surface covering the species is called the aboral (meaning the side opposite to the mouth) or dorsal surface. On this surface there is a structure called the madreporite (or the sieve plate), a small white spot located slightly off-center on the central disc, which acts as a [[water]] filter and supplies the starfish's water vascular system with water to move. |
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[[Image:Schmedelian pin-cushion sea star.jpg|thumb|left|200px|Schmedelian pin-cushion sea star on Meedhupparu house reef in the Maldives, ''Culcita schmedeliana'']] | [[Image:Schmedelian pin-cushion sea star.jpg|thumb|left|200px|Schmedelian pin-cushion sea star on Meedhupparu house reef in the Maldives, ''Culcita schmedeliana'']] | ||
| − | + | Also on the aboral surface, surrounding the spines, are small white objects known as pedicellarieae, which are like pairs of claws or jaws that serve to prevent encrusting organisms from colonizing the starfish. | |
| − | + | The radial canals (a part of the internal anatomy), which extend from the center of the starfish out towards the tips of the arms, have hollow, muscular tube feet branching from them. Each tube foot has a bulb-like ampulla, as well as a suckered podium (or foot part), which are a part of the water vascular system. Starfish also have a simple photoreceptor eye spot at the end of each arm that is able to "see" differences of [[light]] and dark, enabling the starfish to detect movement. Additional parts, like cribriform organs, that are present exclusively in ''Porcellanasteridae,'' are used to generate current in the burrows made by these starfish. | |
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| − | + | While starfish generally adhere to this basic body plan, there is great variety in the shape (ranging from nearly pentagonal like the Indo-pacific cushion star, ''Culcita novaeguineae,'' to gracile stars like those of the ''Zoroaster'' genus), the [[color]], and the morphology between each species. Some species have patterns that serve as camouflage or warning coloration, which include mosaic-like tiles formed by ossicles, stripes, interconnecting net between spines, and pustules with bright colors, mottles, or spots. | |
| − | + | [[Image:starfish.jpg|thumb|200px|right|The tube feet can be seen on this starfish.]] | |
| + | [[Image:Henricia-sanguinolenta.jpg|thumb|200px|right|Blood star ''(Henricia sanguinolenta)'' at the New England Aquarium, displaying its tube feet.]] | ||
==Internal anatomy== | ==Internal anatomy== | ||
| + | ===Hemal system=== | ||
[[Image:Asterias.png|left|400px|thumb|'''Dissection of''' '''''Asterias rubens'''''. 1 - Ambulacral ossicles and ampullae. 2 - Madreporite. 3 - Stone canal. 4 - Pyloric caecae. 5 - Rectal glands. 6 - Gonads.]] | [[Image:Asterias.png|left|400px|thumb|'''Dissection of''' '''''Asterias rubens'''''. 1 - Ambulacral ossicles and ampullae. 2 - Madreporite. 3 - Stone canal. 4 - Pyloric caecae. 5 - Rectal glands. 6 - Gonads.]] | ||
| − | + | [[Echinoderm]]s have a simple [[circulatory system]] (also called a hemal system) that is linked with their excretory system. In the starfish, the hemal system is used to distribute nutrients, with hemal channels forming rings around the mouth (the oral hemal ring), closer to the top of the starfish, and around the [[digestive system]] (the gastric hemal ring). The axial sinus, a portion of the body cavity, connects these three rings. Each ray of the starfish also has hemal channels running next to the gonads. | |
===Digestion and excretion=== | ===Digestion and excretion=== | ||
| − | Starfish digestion is carried out in two separate [[stomach]]s | + | Starfish digestion is carried out in two separate [[stomach]]s, the cardiac stomach and the pyloric stomach. The cardiac stomach, which is a sack-like stomach located at the center of the body, may be everted, or pushed out of the organism's body, and used to engulf and digest [[food]]. Some species take advantage of the great endurance of their [[water]] vascular systems to force open the shells of [[bivalve]] [[mollusk]]s, such as [[clam]]s and [[mussel]]s, and inject their stomachs into the shells. Once the stomach is inserted inside the shell, it digests the mollusk in place. The cardiac stomach is then brought back inside the body, and the partially digested food is moved to the pyloric stomach. Further digestion occurs in the intestine and waste is excreted through the anus on the aboral side of the body. |
| − | Because of this ability to digest food outside of its body, the sea star is able to hunt prey that are much larger than its mouth would otherwise allow including [[arthropod]]s, and even small [[fish]] in addition to mollusks. | + | Because of this ability to digest [[food]] outside of its body, the sea star is able to hunt prey that are much larger than its mouth would otherwise allow, including [[arthropod]]s, and even small [[fish]], in addition to mollusks. |
| − | Some | + | Some [[echinoderm]]s have been shown to live for several weeks without food under artificial conditions—it is believed that they may receive some nutrients from organic material dissolved in seawater. |
| − | Sea stars and other echinoderms have endoskeletons (internal | + | ===Skeletal system=== |
| − | + | Sea stars and other echinoderms have endoskeletons (internal [[skeleton]]s) that are composed of calcium carbonate plates called ossicles, which is one of the reasons some scientists are led to believe that echinoderms are very closely related to [[chordate]]s (animals with a hollow nerve chord that usually have vertebrae). | |
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===Nervous system=== | ===Nervous system=== | ||
[[Image:Star Fish at Montana de Oro.JPG|thumb|Bat star, ''Asterina miniata'']] | [[Image:Star Fish at Montana de Oro.JPG|thumb|Bat star, ''Asterina miniata'']] | ||
| − | + | Starfish have a simple [[nervous system]] that lacks a true [[brain]], and consists partially of a [[nerve]] plexus (a network of interlacing nerves), which lies within as well as below the [[skin]]. The esophagus is also surrounded by a number of nerve rings, which send radial nerves that are often parallel with the branches of the [[water]] vascular system. The ring nerves and radial nerves coordinate the starfish's balance and directional systems. | |
| + | |||
| + | Although [[echinoderm]]s do not have many well-defined sensory inputs, they are sensitive to touch, light, temperature, orientation, and the status of water around them. The tube feet, spines, and pedicellariae found on starfish are sensitive to touch, while eye spots on the ends of the rays are light-sensitive. | ||
==Behavior== | ==Behavior== | ||
===Diet=== | ===Diet=== | ||
[[Image:Starfishmussel.jpg|thumb|right|Starfish consuming a mussel]] | [[Image:Starfishmussel.jpg|thumb|right|Starfish consuming a mussel]] | ||
| − | Most species are generalist predators, some eating | + | Most species of starfish are generalist predators, some eating [[bivalve]]s like [[mussel]]s, [[clam]]s, and [[oyster]]s, or any [[animal]] too slow to evade the attack (such as dying [[fish]]). Some species are detritivores, eating decomposed animal and [[plant]] material, or organic films attached to substrate. Others may consume [[coral]] polyps (the best-known example for this is the infamous ''Acanthaster planci,'' or crown-of-thorns starfish, which also secretes toxins from its spines), [[sponge]]s, or even suspended particles and [[plankton]] (starfish from the Order Brisingida). |
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| + | The process of feeding or capture may be aided by special parts, such as specialized tube feet (used by the ''Pisaster brevispinus'' or short-spined pisaster) capable of extending deep into the soft substrata in order to haul out prey (usually clams) from within (Nybakken 1997). | ||
===Reproduction=== | ===Reproduction=== | ||
[[Image: Ochre sea star on beach, Olympic National Park USA.jpg|thumb|Ochre sea star, ''Pisaster ochraceus'' on beach at Olympic National Park, USA.]] | [[Image: Ochre sea star on beach, Olympic National Park USA.jpg|thumb|Ochre sea star, ''Pisaster ochraceus'' on beach at Olympic National Park, USA.]] | ||
| − | Starfish are capable of both sexual and asexual reproduction | + | Starfish are capable of both [[sexual reproduction|sexual]] and [[reproduction#asexual reproduction|asexual]] [[reproduction]], with individual starfish being male or female. Fertilization takes place externally, with both male and female releasing their gametes into the environment. Resulting fertilized [[embryo]]s form part of the [[plankton|zooplankton]]. |
| − | Starfish are developmentally ([[embryo]]logically) known as | + | Starfish are developmentally ([[embryo]]logically) known as deuterostomes. Their embryo initially develops [[symmetry (biology)#bilateral symmetry|bilateral symmetry]], leading some scientists to believe that starfish share a common ancestor with [[chordate]]s. Later development takes a very different path as the developing starfish settles out of the zooplankton and develops its characteristic radial symmetry. Some species reproduce cooperatively, using environmental signals to coordinate the timing of gamete release, while in other species one to one pairing is more normal. |
| − | Some species of starfish also reproduce asexually by fragmentation, often with part of an arm becoming detached and eventually developing into an independent individual starfish. This has led to some notoriety. Starfish can be pests to fishermen who make their living on the capture of clams and other | + | Some species of starfish are also able to reproduce asexually by fragmentation, often with part of an arm becoming detached and eventually developing into an independent individual starfish. This has led to some notoriety. Starfish can be pests to fishermen who make their living on the capture of clams and other [[mollusk]]s at sea as starfish prey on these. The fishermen presumably would kill the starfish by chopping them up and disposing of them at sea, ultimately leading to their increased numbers until the issue was better understood. A starfish arm can only regenerate into a whole new organism if some of the central [[nerve]] ring of the starfish is part of the chopped off arm. |
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===Locomotion=== | ===Locomotion=== | ||
| − | [[Image:Stachelhaeuter fg01.jpg|thumb|left|The underside of a sea star. | + | [[Image:Stachelhaeuter fg01.jpg|thumb|left|The underside of a sea star. The inset shows a magnified view of the tube feet.]] |
| − | Starfish move using a water vascular system. Water comes into the system via the madreporite. It is then circulated from the stone canal to the ring canal and into the radial canals. The radial canals carry water to the ampullae and provide suction to the tube feet. The tube feet latch on to surfaces and move in a wave, with one body section attaching to the surfaces as another releases. | + | Starfish, like other [[echinoderm]]s, move using a water vascular system. Water comes into the system via the madreporite. It is then circulated from the stone canal to the ring canal and into the radial canals. The radial canals carry water to the ampullae and provide suction to the tube feet. The tube feet latch on to surfaces and move in a wave, with one body section attaching to the surfaces as another releases. |
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| + | Much of the adhesion of starfish to surface is [[chemical]], with the tube foot secreting substances that either bond with surfaces or break down the bonds with surfaces, allowing the tube foot to move. This is so that extra [[energy]] is not expended by the starfish in order to stay attached by exerting its muscles. As a result of all this, most starfish cannot move quickly. However, some burrowing species like starfish from genus ''Astropecten'' and ''Luidia'' are quite capable of rapid, creeping motion—it "glides" across the ocean floor. This motion results from their pointed tube feet adapted specially for excavating local area of sand. | ||
[[Image:Starfish Roentgen X-Ray 01 Nevit.jpg|thumb|Starfish skeleton]] | [[Image:Starfish Roentgen X-Ray 01 Nevit.jpg|thumb|Starfish skeleton]] | ||
===Regeneration=== | ===Regeneration=== | ||
[[Image:Starfishmissinglegsn2.jpg|thumb|left|A sunflower starfish is regenerating his arms that were lost to predators]] | [[Image:Starfishmissinglegsn2.jpg|thumb|left|A sunflower starfish is regenerating his arms that were lost to predators]] | ||
| − | Some species of starfish have the ability to regenerate lost arms and can regrow an entire new arm in time. Most species must have the central part of the body intact to be able to regenerate, but a few can grow an entire starfish from a single ray | + | Some species of starfish have the ability to regenerate lost arms and can regrow an entire new arm in time. Most species must have the central part of the body intact to be able to regenerate, but a few (such as the red and blue ''Linkia'' star) can grow an entire starfish from a single ray. The regeneration of these stars is possible due to the vital organs kept in their arms.[[Image:Guildings's sea star.jpg|thumb|right|Guilding's sea star on Meedhupparu house reef in the [[Maldives]], ''Linkia guildingi'']] |
==Geological history== | ==Geological history== | ||
| − | Fossil starfish and brittle stars are first known from rocks of [[Ordovician]] age indicating that two groups probably diverged in the [[Cambrian]]. However, Ordovician examples of the two groups show many similarities and can be difficult to distinguish. Complete fossil starfish are very rare, but where they do occur they may be abundant | + | [[Fossil]] starfish and brittle stars are first known from rocks of the [[Ordovician]] age indicating that two groups probably diverged in the [[Cambrian]]. However, Ordovician examples of the two groups show many similarities and can be difficult to distinguish. Complete fossil starfish are very rare, but where they do occur they may be abundant. |
| − | == | + | Most fossil starfish consist of scattered individual plates or segments of arms, since the endoskeleton is composed of multiple plates that fall apart and are scattered after death and the decay of the soft parts of the creature. Scattered starfish ossicles are relatively common in the [[Cretaceous]] [[Chalk]] Formation of [[England]]. |
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| − | + | ==Notes== | |
| − | + | <references/> | |
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==References== | ==References== | ||
| − | + | * Blake, D. B., and T. E. Guensburg. 2005. Implications of a new early Ordovician asteroid (Echinodermata) for the phylogeny of Asterozoans. ''Journal of Paleontology'' 79 (2): 395-399. | |
| − | * Blake | + | * California Academy of Sciences. [http://www.calacademy.org/research/izg/echinoderm/classify.htm Classification of the Extant Echinodermata]. ''California Academy of Sciences''. Retrieved August 21, 2007. |
| − | * Gilbertson, | + | * Cornell University. 2007. [http://instruct1.cit.cornell.edu/courses/biog105/labs/deuts/echinoderms.html BioG 105/106: Phylum Echinodermata]. ''Cornell University, Autotutorial Introductory Biology''. Retrieved August 21, 2007. |
| − | * Shackleton, | + | * Gilbertson, L. 1999. ''Zoology Lab Manual''. New York: McGraw Hill Companies. ISBN 007237716X |
| − | * Solomon, E.P. | + | * Hickman, C. P, L. S. Roberts, A. Larson, H. l'Anson, and D. J. Eisenhour. 2006. ''Integrated Principles of Zoology''. New York: McGraw Hill. ISBN 0071115935 |
| − | * Sutton | + | * Nybakken, J. W. 1997. ''Marine Biology: An Ecological Approach.'' Addison-Wesley Longman. ISBN 0673994511 |
| − | * | + | * Shackleton, J. D. 2005. Skeletal homologies, phylogeny and classification of the earliest asterozoan echinoderms. ''Journal of Systematic Palaeontology'' 3 (1): 29-114. |
| + | * Solomon, E. P., L. R.Berg, and D. W. Martin. 2002. ''Biology.'' ISBN 0534391753 | ||
| + | * Stevenson, J. 2006. [http://users.bigpond.net.au/je.st/starfish/index.html Starfish—sea stars home Page]. ''Sealife''. Retrieved August 21, 2007. | ||
| + | * Sutton, M. D., D. E. G. Briggs, D. J. Siveter, D. J. Siveter, and D. J. Gladwell. 2005. A starfish with three-dimensionally preserved soft parts from the Silurian of England. ''Proceedings of the Royal Society, London B'' 272 (1567): 1001-1006. | ||
| + | * Wrey, G. A. 1999. Echinodermata. Spiny-skinned animals: Sea urchins, starfish, and their allies. ''Tree of Life Project''. | ||
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{{credit|144910722}} | {{credit|144910722}} | ||
Latest revision as of 04:39, 28 April 2023
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 "Asteroidea" from Ernst Haeckel's Kunstformen der Natur, 1904
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Brisingida (100 species[1]) |
Starfish, or sea stars (a less confusing designation, since they are only very distantly related to fish), are marine invertebrates belonging to the kingdom animalia, phylum Echinodermata, and class Asteroidea, of which there are over 1800 known species. The names sea star and starfish also are (incorrectly) used for the closely related brittle stars, which make up the class Ophiuroidea.
Characteristics of sea stars include that they exhibit a superficially radial symmetry, since they typically have five or more "arms" which radiate from an indistinct disk (pentaradial symmetry), and tend to be covered in spines. Like other echinoderms, starfish possess an endoskeleton, but do not rely on it for support and locomotion, instead using a hydraulic water vascular system that functions via many projections called tube feet, located on the ventral surface of the starfish's arms.
Starfish are an essential part of marine food chains, being eaten as larvae, and becoming voracious predators upon reaching adulthood, with their diet including mollusks and other marine organisms. They even are able to evert their stomach in order to pull apart and digest their shelled prey. For humans, they are a source of aesthetic joy and their remarkable characteristics—movement based on a hydraulics system, ability to regenerate lost limbs and evert their stomach—add to the wonder of nature. Some starfish also are captured, dried out, and turned into popular souvenirs or decorations.
While starfish can reproduce sexually, if an arm of the starfish is severed from the body and it contains a portion of the central nerve ring, then another starfish can regrow from the severed portion. Lack of knowledge of this capacity increased the role of starfish as pests to commercial clam and oyster beds when fisherman, in an effort to get rid of these predators, would cut them up and throw them back into the ocean.
Distribution
There are about 1,800 known living species of starfish, and they occur in all of the Earth's oceans (though the greatest variety of starfish is found in the tropical Indo-Pacific). Habitats range from tropical coral reefs to kelp forests to the deep-sea floor, although none live within the water column; all species of starfish found are living as benthos (or on the ocean floor).
Because echinoderms need to maintain a delicate internal balance in their body, no starfish are found in freshwater environments.
External anatomy
Starfish are composed of a central disc from which arms (usually five, but the number can vary) sprout in pentaradial symmetry. The mouth is located underneath the starfish on the oral or ventral surface, while the anus is located on the top of the animal. The spiny upper surface covering the species is called the aboral (meaning the side opposite to the mouth) or dorsal surface. On this surface there is a structure called the madreporite (or the sieve plate), a small white spot located slightly off-center on the central disc, which acts as a water filter and supplies the starfish's water vascular system with water to move.
Also on the aboral surface, surrounding the spines, are small white objects known as pedicellarieae, which are like pairs of claws or jaws that serve to prevent encrusting organisms from colonizing the starfish.
The radial canals (a part of the internal anatomy), which extend from the center of the starfish out towards the tips of the arms, have hollow, muscular tube feet branching from them. Each tube foot has a bulb-like ampulla, as well as a suckered podium (or foot part), which are a part of the water vascular system. Starfish also have a simple photoreceptor eye spot at the end of each arm that is able to "see" differences of light and dark, enabling the starfish to detect movement. Additional parts, like cribriform organs, that are present exclusively in Porcellanasteridae, are used to generate current in the burrows made by these starfish.
While starfish generally adhere to this basic body plan, there is great variety in the shape (ranging from nearly pentagonal like the Indo-pacific cushion star, Culcita novaeguineae, to gracile stars like those of the Zoroaster genus), the color, and the morphology between each species. Some species have patterns that serve as camouflage or warning coloration, which include mosaic-like tiles formed by ossicles, stripes, interconnecting net between spines, and pustules with bright colors, mottles, or spots.
Internal anatomy
Hemal system
Echinoderms have a simple circulatory system (also called a hemal system) that is linked with their excretory system. In the starfish, the hemal system is used to distribute nutrients, with hemal channels forming rings around the mouth (the oral hemal ring), closer to the top of the starfish, and around the digestive system (the gastric hemal ring). The axial sinus, a portion of the body cavity, connects these three rings. Each ray of the starfish also has hemal channels running next to the gonads.
Digestion and excretion
Starfish digestion is carried out in two separate stomachs, the cardiac stomach and the pyloric stomach. The cardiac stomach, which is a sack-like stomach located at the center of the body, may be everted, or pushed out of the organism's body, and used to engulf and digest food. Some species take advantage of the great endurance of their water vascular systems to force open the shells of bivalve mollusks, such as clams and mussels, and inject their stomachs into the shells. Once the stomach is inserted inside the shell, it digests the mollusk in place. The cardiac stomach is then brought back inside the body, and the partially digested food is moved to the pyloric stomach. Further digestion occurs in the intestine and waste is excreted through the anus on the aboral side of the body.
Because of this ability to digest food outside of its body, the sea star is able to hunt prey that are much larger than its mouth would otherwise allow, including arthropods, and even small fish, in addition to mollusks.
Some echinoderms have been shown to live for several weeks without food under artificial conditions—it is believed that they may receive some nutrients from organic material dissolved in seawater.
Skeletal system
Sea stars and other echinoderms have endoskeletons (internal skeletons) that are composed of calcium carbonate plates called ossicles, which is one of the reasons some scientists are led to believe that echinoderms are very closely related to chordates (animals with a hollow nerve chord that usually have vertebrae).
Nervous system
Starfish have a simple nervous system that lacks a true brain, and consists partially of a nerve plexus (a network of interlacing nerves), which lies within as well as below the skin. The esophagus is also surrounded by a number of nerve rings, which send radial nerves that are often parallel with the branches of the water vascular system. The ring nerves and radial nerves coordinate the starfish's balance and directional systems.
Although echinoderms do not have many well-defined sensory inputs, they are sensitive to touch, light, temperature, orientation, and the status of water around them. The tube feet, spines, and pedicellariae found on starfish are sensitive to touch, while eye spots on the ends of the rays are light-sensitive.
Behavior
Diet
Most species of starfish are generalist predators, some eating bivalves like mussels, clams, and oysters, or any animal too slow to evade the attack (such as dying fish). Some species are detritivores, eating decomposed animal and plant material, or organic films attached to substrate. Others may consume coral polyps (the best-known example for this is the infamous Acanthaster planci, or crown-of-thorns starfish, which also secretes toxins from its spines), sponges, or even suspended particles and plankton (starfish from the Order Brisingida).
The process of feeding or capture may be aided by special parts, such as specialized tube feet (used by the Pisaster brevispinus or short-spined pisaster) capable of extending deep into the soft substrata in order to haul out prey (usually clams) from within (Nybakken 1997).
Reproduction
Starfish are capable of both sexual and asexual reproduction, with individual starfish being male or female. Fertilization takes place externally, with both male and female releasing their gametes into the environment. Resulting fertilized embryos form part of the zooplankton.
Starfish are developmentally (embryologically) known as deuterostomes. Their embryo initially develops bilateral symmetry, leading some scientists to believe that starfish share a common ancestor with chordates. Later development takes a very different path as the developing starfish settles out of the zooplankton and develops its characteristic radial symmetry. Some species reproduce cooperatively, using environmental signals to coordinate the timing of gamete release, while in other species one to one pairing is more normal.
Some species of starfish are also able to reproduce asexually by fragmentation, often with part of an arm becoming detached and eventually developing into an independent individual starfish. This has led to some notoriety. Starfish can be pests to fishermen who make their living on the capture of clams and other mollusks at sea as starfish prey on these. The fishermen presumably would kill the starfish by chopping them up and disposing of them at sea, ultimately leading to their increased numbers until the issue was better understood. A starfish arm can only regenerate into a whole new organism if some of the central nerve ring of the starfish is part of the chopped off arm.
Locomotion
Starfish, like other echinoderms, move using a water vascular system. Water comes into the system via the madreporite. It is then circulated from the stone canal to the ring canal and into the radial canals. The radial canals carry water to the ampullae and provide suction to the tube feet. The tube feet latch on to surfaces and move in a wave, with one body section attaching to the surfaces as another releases.
Much of the adhesion of starfish to surface is chemical, with the tube foot secreting substances that either bond with surfaces or break down the bonds with surfaces, allowing the tube foot to move. This is so that extra energy is not expended by the starfish in order to stay attached by exerting its muscles. As a result of all this, most starfish cannot move quickly. However, some burrowing species like starfish from genus Astropecten and Luidia are quite capable of rapid, creeping motion—it "glides" across the ocean floor. This motion results from their pointed tube feet adapted specially for excavating local area of sand.
Regeneration
Some species of starfish have the ability to regenerate lost arms and can regrow an entire new arm in time. Most species must have the central part of the body intact to be able to regenerate, but a few (such as the red and blue Linkia star) can grow an entire starfish from a single ray. The regeneration of these stars is possible due to the vital organs kept in their arms.
Geological history
Fossil starfish and brittle stars are first known from rocks of the Ordovician age indicating that two groups probably diverged in the Cambrian. However, Ordovician examples of the two groups show many similarities and can be difficult to distinguish. Complete fossil starfish are very rare, but where they do occur they may be abundant.
Most fossil starfish consist of scattered individual plates or segments of arms, since the endoskeleton is composed of multiple plates that fall apart and are scattered after death and the decay of the soft parts of the creature. Scattered starfish ossicles are relatively common in the Cretaceous Chalk Formation of England.
Notes
ReferencesISBN links support NWE through referral fees
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