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'''Paleoanthropology''', a sub-discipline of  [[Anthropology]]. Studying [[Hominidae|hominid]] [[fossil]] evidence, such as [[Petrifaction|petrified]] bones and [[footprint]]s, and even incorporating knoweldge of current primate species, paleoanthropologits essentially study the origin of human beings as a biological spieces.  
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'''Paleoanthropology''' is a sub-discipline of  [[anthropology]] and [[paleontology]], and is also known as '''human anthropology.''' Studying [[Hominidae|hominid]] [[fossil]] evidence, such as [[Petrifaction|petrified]] bones and [[footprint]]s, as well as [[Artifact (archaeology)|artifact]]s such as [[tool]]s, and even incorporating knowledge of current [[primate]] species, paleoanthropologists essentially study the origin of [[human being]]s.
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Of great interest to this field is the theory of [[human evolution]], including the pattern and process of evolution, or the search for the "missing link" between the [[ape]]s and humans. Many paleoanthropologists have made their name through finding fossils purported to be of such an intermediate species, such as [[Davidson Black]], who discovered "[[Peking Man]]," [[Eugene Dubois]] with "[[Java Man]]," [[Richard Leakey]] and his "[[Turkana Boy]]," and [[Donald Johanson]] who discovered the 3.2 million year old [[Australopithecine]] fossil "[[Lucy (Australopithecus)| Lucy]]." Beyond the excitement of finding such examples, these scientists also debate the geographical origins of humankind, with each find supporting or disproving the various theories. Historically, a major source of controversy has been the process by which humans have developed, whether by a force with a random component (natural selection) or by the creative force of a Creator [[God]]. [[Abrahamic religion]]s believe in a single-point origin of modern humans, beginning with an "[[Adam and Eve]]."
  
 
==Etymology==
 
==Etymology==
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The word '''paleoanthropology''' is an academic creation that combines the [[Ancient Greek Language|Ancient Greek]] ''paleo,'' which refers to prehistoric time periods, with "anthropology," itself a combination of Greek words which mean "study of man."<ref>Dictionary.com, [http://dictionary.reference.com/browse/Paleoanthropology Paleoanthropology,] ''Dictionary.com Unabridged (v 1.1)''. Retrieved May 05, 2008. </ref>
  
The word paleoanthropology is an academic creation that combines the [[Ancient Greek Language|Ancient Greek]] ''paleo'', which refers to prehistoric time periods, with anthropology, itself a combination of Greek words which mean "study of man".<ref>Paleoanthropology. (n.d.). Dictionary.com Unabridged (v 1.1). Retrieved May 05, 2008, from Dictionary.com website: http://dictionary.reference.com/browse/Paleoanthropology</ref>
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Paleoanthropology is actually a sub-division of two different larger fields of study: [[Anthropology]] and [[paleontology]], and is sometimes known as '''human paleontology.''' Anthropology is concerned with the study of humankind's cultural and biological evolution through all time. Paleontology is the study of [[prehistory|prehistoric]] life forms on Earth through the examination of [[fossil]]s. Hence, paleoanthropology studies the prehistoric ancestors of humankind, referred to in a group as ''hominids''. The discipline often overlaps with [[geology]] (the study of rocks and rock formations) as well as with [[botany]], [[biology]], [[zoology]], and [[ecology]]—fields concerned with [[Organism|life forms]] and how they interact.
 
 
==Summary==
 
  
 
==Origins==
 
==Origins==
===Nineteenth century===
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[[Image:Neanderthal child.jpg|thumb|right|200px|Reconstruction of a Neanderthal child, made using modern techniques of computer-assisted paleoanthropology from the Gibraltar 2 Neanderthal specimen.]]
The science arguably began in the late 1800s when important discoveries occurred which led to the study of [[human evolution]]. The discovery of the [[Neanderthal]] in [[Germany]], [[Thomas Huxley]]'s ''[[Evidence as to Man's Place in Nature]]'', and [[Charles Darwin]]'s ''[[The Descent of Man]]'' were all important to early paleoanthropological research.
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The modern field of paleoanthropology (study of human origins) began in the nineteenth century with the discovery of "[[Neanderthal]] man" (the eponymous skeleton was found in 1856, but there had been finds elsewhere since 1830). However, fossils of Neanderthals were widely misinterpreted as skeletons of modern humans with deformation or [[disease]].<ref>S.J. Gould, "Men of the Thirty-third Division," ''Natural History'' (1990): 12-24.</ref>
  
The modern field of paleoanthropology began in the [[19th century]] with the discovery of "[[Neanderthal]] man" (the eponymous [[skeleton]] was found in 1856, but there had been finds elsewhere since 1830), and with evidence of so-called [[caveman|cave men]].  The idea that humans are similar to certain [[great ape]]s had been obvious to people for some time, but the idea of the biological evolution of species in general was not legitimized until after [[Charles Darwin]] published ''[[On the Origin of Species]]'' in 1859.
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==Research activities==
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Paleoanthropologists usually operate in one of two arenas: searching for physical remains and evidence in the field, or analyzing finds in a laboratory. In the field, discovering physical remains and other [[fossil]]s follows painstaking procedures similar to those [[archaeology|archaeologists]] use when uncovering cultural remains.  
  
Though Darwin's first book on evolution did not address the specific question of [[human evolution]]&mdash; "light will be thrown on the origin of man and his history," was all Darwin wrote on the subject&mdash; the implications of evolutionary theory were clear to contemporary readers.
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Areas where evidence is thought to be buried are systematically noted for geological data before layers of earth are removed slowly. Noting the condition and details of the location of the find is just as crucial as uncovering fossils. Once remains are discovered, they are usually sent to a laboratory or research center where they are carefully studied, using chemical and physical dating methods, [[X-Ray]]s, [[MRI]]s, and other special tools. Paleoanthropologists are most interested in noting how the finds are similar and how they are different from already established ancestral lines.
  
Debates between [[Thomas Huxley]] and [[Richard Owen]] focused on the idea of human evolution. Huxley convincingly illustrated many of the similarities and differences between humans and apes in his 1863 book ''[[Evidence as to Man's Place in Nature]]''. By the time Darwin published his own book on the subject, ''[[Descent of Man]]'', it was already a well-known interpretation of his theory&mdash; and the interpretation which made the theory highly controversial. Even many of Darwin's original supporters (such as [[Alfred Russel Wallace]] and [[Charles Lyell]]) balked at the idea that human beings could have evolved their apparently boundless mental capacities and moral sensibilities through [[natural selection]].
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[[Africa]] and [[Asia]] are two of the most popular sites for paleoanthropologists in the field, for they have historically yielded the oldest and most promising evidence. However, as recent finds in [[South America]] and [[Europe]] push the date of humankind's origin further back, paleoanthropologists can be found worldwide seeking to make a discovery of ancient human remains.
  
===Earlier===
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==Theories and findings==
Since the time of [[Carolus Linnaeus]], the great apes were considered the closest relatives of human beings, based on morphological similarity. In the [[19th century]], it was speculated that their closest living relatives were [[chimpanzee]]s and [[gorilla]]s, and based on the natural range of these creatures, it was surmised humans share a [[common ancestor]] with [[Africa]]n apes and that [[fossil]]s of these ancestors would ultimately be found in Africa.<ref name="name">[http://www.evoled.org/lessons/human.htm Human Evolution: Background Information] by Kerry Bright. ''National Science Foundation'' at the ''University of Montana''. Retrieved October 28, 2007.</ref>
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[[Image:Charles Darwin 01.jpg|thumb|right|200px|Charles Darwin (1809-1882) in his later years.]]
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The science of paleoanthropology is based upon the scientific theories of [[Human evolution]]. These involve both the pattern of evolution (descent with modification, the non-causal relations between ancestral and descendant species) and the process of evolution (various theories involving mechanisms and causes for the pattern observed, including fundamental concepts such as natural selection, punctuated equilibrium, and design, and specific scenarios, such as those involving movement from trees, use of tools, "out of Africa," and so forth).
  
===Asia===
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The idea that humans are similar to certain great apes had been obvious to people for some time, but the idea of the biological evolution of species in general was not significantly advanced until after [[Charles Darwin]] published ''On the Origin of Species,'' in 1859. Though Darwin's first book on evolution did not address the specific question of human evolution—"light will be thrown on the origin of man and his history" was all Darwin wrote on the subject—the implications of evolutionary theory were clear to contemporary readers.  
Prior to today's general acceptance of Africa as the root of genus ''Homo,'' [[19th century]] naturalists sought after the origin of man in Asia.  So-called "dragon bones" (fossil bones and teeth) from Chinese apothecary shops were known, but was not until the early 1900s that German paleontologist, [[Max Schlosser]], first described a single human tooth from [[Beijing]]. Although Schlosser (1903) was very cautious, identifying the tooth only as “?''Anthropoide g. et sp. indet''?,” he was hopeful that future work would discover a new anthropoid in [[China]].
 
  
Eleven years later, the Swedish geologist [[Johan Gunnar Andersson]] was sent to China as a mining adviser and soon developed an interest in “dragon bones.” It was he who, in 1918, discovered the sites around [[Zhoukoudian]], a village about 50 kilometers southwest of Beijing proper. However, because of the sparse nature of the initial finds, the site was abandoned.  Work did not resume until 1921, when the Austrian paleontologist, Otto Zdansky, fresh with his doctoral degree from Vienna, came to Beijing to work for Andersson. Zdansky conducted short-term excavations at Locality 1 in 1921 and 1923, and recovered only two teeth of significance (one premolar and one molar) that he subsequently described, cautiously, as “?''Homo sp.''” (Zdansky, 1927). With that done, Zdansky returned to Austria and suspended all fieldwork.
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Debates between [[Thomas Huxley]] and [[Richard Owen]] focused on the idea of human evolution. Huxley convincingly illustrated many of the similarities and differences between humans and apes in his 1863 book, ''Evidence as to Man's Place in Nature''. By the time Darwin published his own book on the subject, ''Descent of Man,'' it was already a well-known interpretation of his theory--and the interpretation helped make the theory of natural selection highly controversial. Even many of Darwin's original supporters (such as [[Alfred Russel Wallace]] and [[Charles Lyell]]) balked at the idea that human beings could have evolved their apparently boundless mental capacities and moral sensibilities through natural selection.
  
News of the fossil hominin [[teeth]] delighted the scientific community in Beijing, and plans soon began to formulate aimed at developing a larger, more systematic project at Zhoukoudian. At the epicenter of excitement was [[Davidson Black]], a Canadian-born anatomist working at the [[Peking Union Medical College]]. Black shared Andersson’s interest, as well as his view that central Asia was a promising home for early humankind. In late 1926, Black channeled a formal proposal through the [[Union Medical College]] to the [[Rockefeller Foundation]] concerning financial support for the systematic excavation at Zhoukoudian, and the establishment of an institute for the study of human biology in China. The [[Zhoukoudian Project]] came into existence in the spring of 1927, and two years later, the [[Cenozoic Research Laboratory]] of the Geological Survey of China was formally established. Being the first institution of its kind, the Cenozoic Laboratory opened up new avenues for the studies of paleogeology and paleontology in China. Moreover, the Laboratory was the precursor of the [[Institute of Vertebrate Paleontology and Paleoanthropology]] (IVPP) of the Chinese Academy of Science,  which took its modern form after 1949.
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Substantial evidence has been marshaled for the fact that humans have descended from common ancestors by a process of branching (descent with modification) and for a primate origin of humans. However, proposals for the specific ancestral-descendant relationships and for the process leading to humans tend to be speculative. And, while the theory of natural selection typically is central to scientific explanations for the process, evidence for natural selection being the directive or creative force is limited to extrapolation from the microevolutionary level (changes within the level of species).
  
The first of the major project finds should be attributed to the young [[Sweden|Swedish]] paleontologist, [[Anders Birger Bohlin]], then serving as the Field Adviser at Zhoukoudian. He recovered a left, lower molar which Black (1927) identified as unmistakably human (it compared favorably to the previous find made by Zdansky), and subsequently coined it ''[[Sinanthropus pekinensis]]'', Black and Zdansky 1927. The news was at first met with skepticism, and many scholars had reservations that a single tooth was sufficient to justify the naming of a new type of early hominin. Yet within a little more than two years, in the winter of 1929, [[Pei Wenzhong]], then the Field Director of Zhoukoudian, unearthed the first, complete calvaria of Peking Man. Twenty-seven years after Schlosser’s initial description, the antiquity of early humans in East Asia was no longer a speculation, but a reality.
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Since the time of [[Carolus Linnaeus]] in the eighteenth century, the [[great ape]]s were considered the closest relatives of human beings, based on morphological similarity. In the nineteenth century, it was speculated that the closest living relatives to human beings were [[chimpanzee]]s and [[gorilla]]s. Based on the natural range of these creatures, it was surmised that humans share a common ancestor with other African apes and that fossils of these ancestors would ultimately be found in [[Africa]].
  
Excavations continued at the site and remained fruitful until the outbreak of the [[Second Sino-Japanese War]] in 1937. The decade long research yielded a wealth of faunal and lithic materials, as well as hominin fossils. These included 5 more complete calvaria, 9 large cranial fragments, 6 facial fragments, 14 partial mandibles, 147 isolated teeth, and 11 postcranial elements-estimated to represent as least 40 individuals. Evidence of fire, marked by ash lenses and burned bones and stones, were apparently also present (Black, 1931), albeit recent studies have challenged this view (Weiner et al., 1998; Weiner et al., 1999). [[Franz Weidenreich]] came to Beijing soon after Black’s untimely death in 1934, and took over the academic affairs of studying the hominin specimens.
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It was not until the 1920s that hominid [[fossil]]s were discovered in Africa. In 1924, [[Raymond Dart]] described ''Australopithecus africanus''. The specimen was the ''Taung Child,'' an ''australopithecine'' infant discovered in a [[cave]] deposit at Taung, [[South Africa]]. The remains were a remarkably well-preserved tiny skull and an endocranial cast of the individual's brain. Although the [[brain]] was small (410 cm3), its shape was rounded, unlike that of chimpanzees and gorillas, and more like a modern human brain. Also, the specimen exhibited short canine teeth, and the position of the foramen magnum (the hole in the skull where the spine enters) was evidence of bipedal locomotion. All of these traits convinced Dart that the Taung baby was a bipedal human ancestor, a transitional form between apes and humans.<ref>Frances Wheelhouse, ''Dart: Scientist and Man of Grit'' (Hornsby, Australia: Transpareon Press, 2001, ISBN 0908021216).</ref> However, the prevailing view of the time was that a large brain evolved before bipedality, it being thought that [[intelligence]] on par with modern humans was a prerequisite to bipedalism. Another twenty years would pass before Dart's claims were taken seriously, following the discovery of more fossils that resembled his find.  
  
===Africa===
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{{:Human evolution/Species chart}}
Following the tragic loss of the Peking Man materials in late 1941, scientific endeavors at Zhoukoudian and elsewhere slowed. For the most part, it was because of the lack of funding. Conversely, frantic search for the missing fossils also figured prominently, and efforts continued well into the 1950s. Nonetheless, after the establishment of the [[People’s Republic of China]] in 1949, excavations resumed at Zhoukoudian. But as the story goes, with political instability and social unrest brewing in China, beginning in 1966, and major discoveries at [[Olduvai Gorge]] and East Turkana ([[Koobi Fora]]), the paleoanthropological spotlights inevitably shifted westward to East Africa. While China re-opened its doors to the West in the late 1970s, national policy calling for self-reliance, coupled with a widened language barrier, thwarted all the possibilities of renewed scientific relationships. Indeed, [[Harvard]] anthropologist K. C. Chang noted, “international collaboration (in developing nations very often a disguise for Western domination) became a thing of the past” (1977: 139).
 
  
Of course, work did not stop everywhere. Back in South Africa, a notable and rare find came to light in 1924. In a cave site at [[Taung]], Professor [[Raymond Dart]] discovered a remarkably well-preserved juvenile specimen (face and brain endocast) and named it ''[[Australopithecus africanus]]'' (''Australopithecus'' = Southern Ape). Although the brain was small (410 cm³), its shape was rounded, unlike that of chimpanzees and gorillas, and more like a modern human brain. In addition, the specimen exhibited short [[canine tooth|canine teeth]], and the [[foramen magnum]] was more anteriorly placed, hinting a [[bipedal]] mode of locomotion. All of these traits convinced Dart that the Taung child was a bipedal human ancestor, a transitional form between ape and man. Another 20 years would pass before Dart's claims were taken seriously, following the discovery of additional australopith fossils in Africa that resemble his type specimen. The prevailing view of the time was that a large brain evolved before bipedality. It was thought that intelligence on par with modern humans was a prerequisite to bipedalism.
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==Debates==
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While there are many aspects of human evolution that paleoanthropologist agree upon, there are several puzzles in regards to the human evolutionary line. One of the most contested issues is from where humans originated. There are two dominant views on the issue of human origins, the '''Out of Africa''' position and the '''multiregional position.''' There are also various combinations of these ideas.
  
Today, the australopiths are considered to be the last common ancestors leading to genus ''Homo'', the group to which modern humans belong. Both australopiths and ''Homo sapiens'' are part of the tribe [[Hominini]], but recent morphological data have brought into doubt the position of ''A. africanus'' as a direct ancestor of modern humans.
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The Out of Africa (or Out of Africa II, or replacement) model holds that there was a [[human migration|migration]] of [[Homo erectus]] (or [[Homo ergaster]]) out of Africa and into Europe and Asia, but that these populations did not subsequently contribute significant amounts of genetic material (or, some say, contributed absolutely nothing) to later populations along the lineage to Homo sapiens.<ref name=kreger>C.D. Kreger, Homo sapiens: Introduction, Archaeology.info  (2005).</ref> Later, approximately 200,000 years ago, there was a second migration of hominids out of Africa, and this was modern [[Homo sapiens]] that replaced the populations that then occupied Europe and Asia.<ref name=kreger/> This view maintains a specific speciation event that led to ''Homo sapiens'' in Africa, and this is the modern human.
  
The australopiths were originally grouped based on size as either [[gracile]] or [[robust]]. The robust variety of ''Australopithecus'' has since been renamed as ''[[Paranthropus]]'' (''[[P. robustus]]'' from [[South Africa]], and ''[[P. boisei]]'' and ''[[P. aethiopicus]]'' from East Africa). In the 1930s, when the robust specimens were first described, the ''Paranthropus'' genus was used. During the 1960s, the robust variety was moved into ''Australopithecus''. The recent consensus has been back to the original classification as a separate genus.
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The multiregional (or continuity) camp hold that since the origin of ''Homo erectus,'' there have been populations of hominids living in the Old World and that these all contributed to successive generations in their regions.<ref name=kreger/> According to this view, hominids in [[China]] and [[Indonesia]] are the most direct ancestors of modern East Asians, and those in Africa are the most direct ancestors of modern Africans. The European populations either gave rise to modern Europeans or contributed significant genetic material to them, while their origins were in Africa or West Asia.<ref name=kreger/> According to this model, there is genetic flow to allow for the maintenance of one species, but not enough to prevent racial differentiation.
  
While Ian Tattersall once noted (''[[Nature (journal)|Nature]]'' 2006, 441:155) that paleoanthropology is distinguished as the "branch of science [that] keeps its primary data secret," it is perhaps more accurate to observe that primary physical evidence in paleoanthropology is among the most difficult to obtain.
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Overall, the disagreement between these two camps has caused "fairly severe strife within the paleoanthropologist community."<ref name=kreger/> Multiregionalism is often "portrayed as a racist theory," while Out of Africa II "has often been portrayed as a religiously motivated idea" that strives to align with the biblical story of [[Genesis]].<ref name=kreger/>
  
==Paradigm==
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Other issues facing paleoanthropology deal with how current evidence is used to create theories. A major contention is that the fossil record remains fragmentary. No fossils of hominids have been found for the period between 6 and 13 million years ago (mya), the time when branching between the chimpanzee and human lineages is expected to have taken place.<ref name=mayr>Ernst Mayr, ''What Evolution Is'' (New York: Basic Books, 2001, ISBN 0465044255).</ref> Furthermore, as author Ernst Mayr notes "most hominid fossils are extremely incomplete. They may consist of part of a mandible, or the upper part of a skull without face and teeth, or only part of the extremities."<ref name=mayr/> Even the famous "[[Lucy (Australopithecus)|Lucy]]" finding ''(Australopithecus afarensis)'' was only a 40 percent complete female skeleton and lacked a head.<ref>S.J. Gould, "Lucy on the Earth in stasis," ''Natural History,'' (1994): 12-20.</ref>
  
==Major Discoveries==
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Coupled with this is a recurrent problem that interpretation of fossil evidence is heavily influenced by personal beliefs and [[prejudice]]s. Fossil evidence often allows a variety of interpretations, since the individual specimens may be reconstructed in a variety of ways.<ref name=wells>Jonathan Wells, ''Icons of Evolution''. (Washington, DC: Regnery Publishing, 2000, ISBN 0895262762)</ref> As Mayr notes, "subjectivity is inevitable in the reconstruction of the missing parts," and virtually all hominid finds and interpretations are "somewhat controversial!"<ref name=mayr/> Another author recounts several examples where the pieces of fossils found offered a variety of reconstructions that were sometimes dramatically different, such as long face versus a short face, a heavy brow, a missing forehead. Different interpretations of two sections of a fossil skull and how to place one of those pieces led Roger Lewin to recount, "How you held it really depended on your preconceptions. It was very interesting what people did with it."<ref name=wells/>
  
 
==Renowned paleoanthropologists==
 
==Renowned paleoanthropologists==
*[[Robert Ardrey]] (1908-1980), whose ''African Genesis'' (1961), ''The Territorial Imperative'' (1966), ''The Social Contract'' (1970), and ''The Hunting Hypothesis'' (1976) detail the mid-[[20th century]] transition in paleoanthropological studies and [[methodology]].
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*[[Robert Ardrey]] (1908-1980), wrote ''African Genesis'' (1961), ''The Territorial Imperative'' (1966), ''The Social Contract'' (1970), and ''The Hunting Hypothesis'' (1976) detailing the mid-twentieth century transition in paleoanthropologist studies and [[methodology]].
*[[Lee R. Berger|Lee Berger]] (1965 - )
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[[Image:Zhoukoudian Museum July2004.jpg|250px|right|thumb|[[Zhoukoudian]] Peking Man Site—the Museum (taken in July 2004). At the center: What Peking Man looked like.]]
*[[Davidson Black]] (1884-1934)
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*[[Davidson Black]] (1884-1934) discovered Sinanthropus pekinensis (now ''Homo erectus pekinensis'') or the “[[Peking Man]].” Despite the fact that Africa was later found to be the origin of humankind, proving Black's theory of an Asian origination wrong, Black's work greatly advanced our knowledge of the development of human beings in Asia.<ref>Encyclopædia Britannica, Davidson Black (2008).</ref>
*[[Robert Broom]] (1866-1951)
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*[[J. Desmond Clark]] (1916-2002)
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*[[Carleton S. Coon]] (1904-1981), in such works as ''The Origin of Races'' (1962), ''The Story of Man'' (1954), and  "The Races of Europe" (1939), concluded that sometimes different racial types had annihilated other types while in other cases warfare and/or settlement had only led to the partial displacement of racial types.
*[[Carleton S. Coon]] (1904-1981)
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*[[Raymond Dart]] (1893-1988)
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*[[Eugene Dubois]] (1858-1940) discovered several fossils of seemingly hominid origin, and called his finds ''Pithecanthropus erectus,'' or [[Java Man]]—"a species in between humans and apes." Later, they were classified as ''Homo erectus.'' What Dubois found in 1890s was a set of teeth, a skullcap, and a left femur (thigh bone). The femur suggested that its owner had walked erect. From the teeth and the skull, Dubois argued that the specimen was exactly between humans and apes on the evolutionary timeline.<ref>Pat Shipman, ''The Man who Found the Missing Link: The Extraordinary Life of Eugene Dubois'' (Diane Publishing Co, 2001, ISBN 075679160X).</ref>
*[[Eugene Dubois]] (1858-1940)
 
*[[Johann Carl Fuhlrott]] (1803-1877)
 
*[[Donald C. Johanson]] (1943- )
 
*[[Kamoya Kimeu]] (1940- )
 
*[[Gustav Heinrich Ralph von Koenigswald]] (1902-1982)
 
*[[Louis Leakey]] (1903-1972)
 
*[[Meave Leakey]] (1942- )
 
*[[Mary Leakey]] (1913-1996)
 
*[[Richard Leakey]] (1944- )
 
*[[André Leroi-Gourhan]] (1911-1986)
 
*[[Kenneth Oakley]] (1911-1981)
 
*[[Pierre Teilhard de Chardin]] (1881-1955)
 
*[[Phillip V. Tobias]] (1925- )
 
*[[Franz Weidenreich]] (1873-1948)
 
*[[Milford H. Wolpoff]] (1942- )
 
*[[Tim White]] (1950- )
 
*[[Ian Tattersall]] (1945- )
 
  
{{:Human evolution/Species chart}}
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*[[Johann Carl Fuhlrott]] (1803-1877) is famous for the discovery of the Neanderthal 1, a Neanderthal specimen found during an archaeology dig in August 1856.
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[[Image:Louis Leakey.jpg|thumb|200 px|Louis Leakey examining skulls from [[Olduvai Gorge]]]]
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*[[Louis Leakey]] (1903-1972) was one of the most renowned paleoanthropologists of all time. Among Leakey’s many extraordinary finds was the 1959 unearthing of Zinjanthropus, a robust hominid that hinted at the great complexity of humankind's evolutionary roots. Leakey called it ''Zinjanthropus boisei,'' and believed that it belonged in the line of direct human ancestors. Later, however, it was classified as australopithecus. In 1964, he and his wife discovered the skull and hand of what was recognized as a new species—''Homo habilis,'' or “the human who used tools.” Using the carbon-14 dating technique, researchers from the University of California at Berkeley have estimated that the site where they were found, and the bones themselves, were 1.75 million years old.<ref>Sonia Cole, ''Leakey's Luck: The Life of Louis Seymour Bazett Leakey, 1903-1972'' (Harcourt, 1975, ISBN 0151494568).</ref>
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*[[Mary Leakey]] (1913-1996) is as famous as her husband Louis. Mary discovered a set of footprints discovered at the Laetoli site located 27 miles (45 kilometers) south of [[Olduvai Gorge]]. The site is [[Pliocene]], dated by the Potassium-argon method to 3.7 million years ago. The footprints, preserved in powdery ash from an eruption of the 20 kilometers distant Sadiman [[volcano]], demonstrate that these hominids walked upright habitually, as there are no knuckle-impressions. The feet do not have the mobile big toe of apes; instead, they have an arch (the bending of the sole of the foot) typical of modern humans. The discovery caused serious debate among scientists, requiring them to change their theories concerning the evolution of bipedalism.<ref name=leakeys>Mary Bowman-Kruhm, ''The Leakeys: A Biography'' (Westport, CT: Greenwood Press, 2005, ISBN 0313329850).</ref>
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*[[Richard Leakey]] (1944- ), the son of Mary and Louis Leakey, built upon the legacy of his parents. In 1969, his discovery of a cranium of ''Australopithecus boisei'' caused great excitement. A Homo habilis skull (ER 1470) and a ''Homo erectus'' skull (ER 3733), discovered in 1972 and 1975, respectively, were among the most significant finds of Leakey's early expeditions. In 1978, an intact cranium of Homo erectus (KNM-ER 3883) was discovered. In 1984, he made his most important discovery—"[[Turkana Boy]]"—the nearly complete skeleton of a young ''Homo erectus'' who died 1.6 million years ago. It was one of the first well-preserved skeletons of that origin ever found.<ref name=leakeys/>
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*[[André Leroi-Gourhan]] (1911-1986), created theories about human evolution involving the notion that the transition to [[biped|bipedality]] freed the hands for grasping, and the face for gesturing and speaking, and thus that the development of the [[cortex]], of [[technology]], and of [[language]] all follow from the adoption of an upright stance.
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*[[Kenneth Oakley]] (1911-1981) is known for his work in the [[Dating methodology (archaeology)|relative dating]] of [[fossil]]s by [[fluorine]] content, which was instrumental in the exposure in the 1950s of the [[Piltdown Man]] hoax.<ref>Encyclopædia Britannica, Kenneth Oakley (2008). </ref>
  
 
==Notes==
 
==Notes==
 
<references/>
 
<references/>
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==References==
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* Bowman-Kruhm, Mary. ''The Leakeys: A Biography''. Westport, CT: Greenwood Press, 2005. ISBN 0313329850.
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* Cole, Sonia. ''Leakey's Luck: The Life of Louis Seymour Bazett Leakey, 1903-1972''. Harcourt, 1975. ISBN 0151494568.
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* Mayr, Ernst. ''What Evolution Is''. New York: Basic Books, 2001. ISBN 0465044255.
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* McKie, Robin. ''Dawn of Man: The Story of Human Evolution''. DK Publishing, 2000. ISBN 978-0789462626.
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* Shipman, Pat. ''The Man who Found the Missing Link: The Extraordinary Life of Eugene Dubois''. Diane Publishing Co, 2001. ISBN 075679160X.
 +
* Wells, Jonathan. ''Icons of Evolution''. Washington, DC: Regnery Publishing, 2000. ISBN 0895262762.
 +
* Wheelhouse, Frances. ''Dart: Scientist and Man of Grit''. Hornsby, Australia: Transpareon Press, 2001. ISBN 0908021216.
  
 
==External links==
 
==External links==
All links retrieved October 28, 2007.
+
All links retrieved November 18, 2022.
* [http://www.jqjacobs.net/anthro/paleo/index.html Paleoanthropology in the 1990's] - by James Q. Jacobs
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* [http://www.jqjacobs.net/anthro/paleo/index.html Paleoanthropology in the 1990's] by James Q. Jacobs
* [http://www.talkorigins.org/faqs/homs/index.html Fossil Hominids] - talkorigins.org
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* [http://www.talkorigins.org/faqs/homs/index.html Fossil Hominids] talkorigins.org
* [http://cogweb.ucla.edu/ep/Paleoanthropology.html#Introduction Aspects of Paleoanthropology] - ucla.edu
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* [http://cogweb.ucla.edu/ep/Paleoanthropology.html#Introduction Aspects of Paleoanthropology] ucla.edu
* [http://www.becominghuman.org/ Paleoanthropology, Evolution and Human Origins] - ''Becoming Human''
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* [http://www.becominghuman.org/ Paleoanthropology, Evolution and Human Origins] ''Becoming Human''
  
 
[[Category:Politics and social sciences]]
 
[[Category:Politics and social sciences]]
 
[[Category:Anthropology]]
 
[[Category:Anthropology]]
 
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[[Category:Paleontology]]
 
{{credits|Paleoanthropology|167238118}}
 
{{credits|Paleoanthropology|167238118}}

Latest revision as of 11:00, 11 March 2023

Human evolution scheme.svg

Paleoanthropology is a sub-discipline of anthropology and paleontology, and is also known as human anthropology. Studying hominid fossil evidence, such as petrified bones and footprints, as well as artifacts such as tools, and even incorporating knowledge of current primate species, paleoanthropologists essentially study the origin of human beings.

Of great interest to this field is the theory of human evolution, including the pattern and process of evolution, or the search for the "missing link" between the apes and humans. Many paleoanthropologists have made their name through finding fossils purported to be of such an intermediate species, such as Davidson Black, who discovered "Peking Man," Eugene Dubois with "Java Man," Richard Leakey and his "Turkana Boy," and Donald Johanson who discovered the 3.2 million year old Australopithecine fossil " Lucy." Beyond the excitement of finding such examples, these scientists also debate the geographical origins of humankind, with each find supporting or disproving the various theories. Historically, a major source of controversy has been the process by which humans have developed, whether by a force with a random component (natural selection) or by the creative force of a Creator God. Abrahamic religions believe in a single-point origin of modern humans, beginning with an "Adam and Eve."

Etymology

The word paleoanthropology is an academic creation that combines the Ancient Greek paleo, which refers to prehistoric time periods, with "anthropology," itself a combination of Greek words which mean "study of man."[1]

Paleoanthropology is actually a sub-division of two different larger fields of study: Anthropology and paleontology, and is sometimes known as human paleontology. Anthropology is concerned with the study of humankind's cultural and biological evolution through all time. Paleontology is the study of prehistoric life forms on Earth through the examination of fossils. Hence, paleoanthropology studies the prehistoric ancestors of humankind, referred to in a group as hominids. The discipline often overlaps with geology (the study of rocks and rock formations) as well as with botany, biology, zoology, and ecology—fields concerned with life forms and how they interact.

Origins

Reconstruction of a Neanderthal child, made using modern techniques of computer-assisted paleoanthropology from the Gibraltar 2 Neanderthal specimen.

The modern field of paleoanthropology (study of human origins) began in the nineteenth century with the discovery of "Neanderthal man" (the eponymous skeleton was found in 1856, but there had been finds elsewhere since 1830). However, fossils of Neanderthals were widely misinterpreted as skeletons of modern humans with deformation or disease.[2]

Research activities

Paleoanthropologists usually operate in one of two arenas: searching for physical remains and evidence in the field, or analyzing finds in a laboratory. In the field, discovering physical remains and other fossils follows painstaking procedures similar to those archaeologists use when uncovering cultural remains.

Areas where evidence is thought to be buried are systematically noted for geological data before layers of earth are removed slowly. Noting the condition and details of the location of the find is just as crucial as uncovering fossils. Once remains are discovered, they are usually sent to a laboratory or research center where they are carefully studied, using chemical and physical dating methods, X-Rays, MRIs, and other special tools. Paleoanthropologists are most interested in noting how the finds are similar and how they are different from already established ancestral lines.

Africa and Asia are two of the most popular sites for paleoanthropologists in the field, for they have historically yielded the oldest and most promising evidence. However, as recent finds in South America and Europe push the date of humankind's origin further back, paleoanthropologists can be found worldwide seeking to make a discovery of ancient human remains.

Theories and findings

Charles Darwin (1809-1882) in his later years.

The science of paleoanthropology is based upon the scientific theories of Human evolution. These involve both the pattern of evolution (descent with modification, the non-causal relations between ancestral and descendant species) and the process of evolution (various theories involving mechanisms and causes for the pattern observed, including fundamental concepts such as natural selection, punctuated equilibrium, and design, and specific scenarios, such as those involving movement from trees, use of tools, "out of Africa," and so forth).

The idea that humans are similar to certain great apes had been obvious to people for some time, but the idea of the biological evolution of species in general was not significantly advanced until after Charles Darwin published On the Origin of Species, in 1859. Though Darwin's first book on evolution did not address the specific question of human evolution—"light will be thrown on the origin of man and his history" was all Darwin wrote on the subject—the implications of evolutionary theory were clear to contemporary readers.

Debates between Thomas Huxley and Richard Owen focused on the idea of human evolution. Huxley convincingly illustrated many of the similarities and differences between humans and apes in his 1863 book, Evidence as to Man's Place in Nature. By the time Darwin published his own book on the subject, Descent of Man, it was already a well-known interpretation of his theory—and the interpretation helped make the theory of natural selection highly controversial. Even many of Darwin's original supporters (such as Alfred Russel Wallace and Charles Lyell) balked at the idea that human beings could have evolved their apparently boundless mental capacities and moral sensibilities through natural selection.

Substantial evidence has been marshaled for the fact that humans have descended from common ancestors by a process of branching (descent with modification) and for a primate origin of humans. However, proposals for the specific ancestral-descendant relationships and for the process leading to humans tend to be speculative. And, while the theory of natural selection typically is central to scientific explanations for the process, evidence for natural selection being the directive or creative force is limited to extrapolation from the microevolutionary level (changes within the level of species).

Since the time of Carolus Linnaeus in the eighteenth century, the great apes were considered the closest relatives of human beings, based on morphological similarity. In the nineteenth century, it was speculated that the closest living relatives to human beings were chimpanzees and gorillas. Based on the natural range of these creatures, it was surmised that humans share a common ancestor with other African apes and that fossils of these ancestors would ultimately be found in Africa.

It was not until the 1920s that hominid fossils were discovered in Africa. In 1924, Raymond Dart described Australopithecus africanus. The specimen was the Taung Child, an australopithecine infant discovered in a cave deposit at Taung, South Africa. The remains were a remarkably well-preserved tiny skull and an endocranial cast of the individual's brain. Although the brain was small (410 cm3), its shape was rounded, unlike that of chimpanzees and gorillas, and more like a modern human brain. Also, the specimen exhibited short canine teeth, and the position of the foramen magnum (the hole in the skull where the spine enters) was evidence of bipedal locomotion. All of these traits convinced Dart that the Taung baby was a bipedal human ancestor, a transitional form between apes and humans.[3] However, the prevailing view of the time was that a large brain evolved before bipedality, it being thought that intelligence on par with modern humans was a prerequisite to bipedalism. Another twenty years would pass before Dart's claims were taken seriously, following the discovery of more fossils that resembled his find.

Human evolution/Species chart

Debates

While there are many aspects of human evolution that paleoanthropologist agree upon, there are several puzzles in regards to the human evolutionary line. One of the most contested issues is from where humans originated. There are two dominant views on the issue of human origins, the Out of Africa position and the multiregional position. There are also various combinations of these ideas.

The Out of Africa (or Out of Africa II, or replacement) model holds that there was a migration of Homo erectus (or Homo ergaster) out of Africa and into Europe and Asia, but that these populations did not subsequently contribute significant amounts of genetic material (or, some say, contributed absolutely nothing) to later populations along the lineage to Homo sapiens.[4] Later, approximately 200,000 years ago, there was a second migration of hominids out of Africa, and this was modern Homo sapiens that replaced the populations that then occupied Europe and Asia.[4] This view maintains a specific speciation event that led to Homo sapiens in Africa, and this is the modern human.

The multiregional (or continuity) camp hold that since the origin of Homo erectus, there have been populations of hominids living in the Old World and that these all contributed to successive generations in their regions.[4] According to this view, hominids in China and Indonesia are the most direct ancestors of modern East Asians, and those in Africa are the most direct ancestors of modern Africans. The European populations either gave rise to modern Europeans or contributed significant genetic material to them, while their origins were in Africa or West Asia.[4] According to this model, there is genetic flow to allow for the maintenance of one species, but not enough to prevent racial differentiation.

Overall, the disagreement between these two camps has caused "fairly severe strife within the paleoanthropologist community."[4] Multiregionalism is often "portrayed as a racist theory," while Out of Africa II "has often been portrayed as a religiously motivated idea" that strives to align with the biblical story of Genesis.[4]

Other issues facing paleoanthropology deal with how current evidence is used to create theories. A major contention is that the fossil record remains fragmentary. No fossils of hominids have been found for the period between 6 and 13 million years ago (mya), the time when branching between the chimpanzee and human lineages is expected to have taken place.[5] Furthermore, as author Ernst Mayr notes "most hominid fossils are extremely incomplete. They may consist of part of a mandible, or the upper part of a skull without face and teeth, or only part of the extremities."[5] Even the famous "Lucy" finding (Australopithecus afarensis) was only a 40 percent complete female skeleton and lacked a head.[6]

Coupled with this is a recurrent problem that interpretation of fossil evidence is heavily influenced by personal beliefs and prejudices. Fossil evidence often allows a variety of interpretations, since the individual specimens may be reconstructed in a variety of ways.[7] As Mayr notes, "subjectivity is inevitable in the reconstruction of the missing parts," and virtually all hominid finds and interpretations are "somewhat controversial!"[5] Another author recounts several examples where the pieces of fossils found offered a variety of reconstructions that were sometimes dramatically different, such as long face versus a short face, a heavy brow, a missing forehead. Different interpretations of two sections of a fossil skull and how to place one of those pieces led Roger Lewin to recount, "How you held it really depended on your preconceptions. It was very interesting what people did with it."[7]

Renowned paleoanthropologists

  • Robert Ardrey (1908-1980), wrote African Genesis (1961), The Territorial Imperative (1966), The Social Contract (1970), and The Hunting Hypothesis (1976) detailing the mid-twentieth century transition in paleoanthropologist studies and methodology.
Zhoukoudian Peking Man Site—the Museum (taken in July 2004). At the center: What Peking Man looked like.
  • Davidson Black (1884-1934) discovered Sinanthropus pekinensis (now Homo erectus pekinensis) or the “Peking Man.” Despite the fact that Africa was later found to be the origin of humankind, proving Black's theory of an Asian origination wrong, Black's work greatly advanced our knowledge of the development of human beings in Asia.[8]
  • Carleton S. Coon (1904-1981), in such works as The Origin of Races (1962), The Story of Man (1954), and "The Races of Europe" (1939), concluded that sometimes different racial types had annihilated other types while in other cases warfare and/or settlement had only led to the partial displacement of racial types.
  • Eugene Dubois (1858-1940) discovered several fossils of seemingly hominid origin, and called his finds Pithecanthropus erectus, or Java Man—"a species in between humans and apes." Later, they were classified as Homo erectus. What Dubois found in 1890s was a set of teeth, a skullcap, and a left femur (thigh bone). The femur suggested that its owner had walked erect. From the teeth and the skull, Dubois argued that the specimen was exactly between humans and apes on the evolutionary timeline.[9]
  • Johann Carl Fuhlrott (1803-1877) is famous for the discovery of the Neanderthal 1, a Neanderthal specimen found during an archaeology dig in August 1856.
Louis Leakey examining skulls from Olduvai Gorge
  • Louis Leakey (1903-1972) was one of the most renowned paleoanthropologists of all time. Among Leakey’s many extraordinary finds was the 1959 unearthing of Zinjanthropus, a robust hominid that hinted at the great complexity of humankind's evolutionary roots. Leakey called it Zinjanthropus boisei, and believed that it belonged in the line of direct human ancestors. Later, however, it was classified as australopithecus. In 1964, he and his wife discovered the skull and hand of what was recognized as a new species—Homo habilis, or “the human who used tools.” Using the carbon-14 dating technique, researchers from the University of California at Berkeley have estimated that the site where they were found, and the bones themselves, were 1.75 million years old.[10]
  • Mary Leakey (1913-1996) is as famous as her husband Louis. Mary discovered a set of footprints discovered at the Laetoli site located 27 miles (45 kilometers) south of Olduvai Gorge. The site is Pliocene, dated by the Potassium-argon method to 3.7 million years ago. The footprints, preserved in powdery ash from an eruption of the 20 kilometers distant Sadiman volcano, demonstrate that these hominids walked upright habitually, as there are no knuckle-impressions. The feet do not have the mobile big toe of apes; instead, they have an arch (the bending of the sole of the foot) typical of modern humans. The discovery caused serious debate among scientists, requiring them to change their theories concerning the evolution of bipedalism.[11]
  • Richard Leakey (1944- ), the son of Mary and Louis Leakey, built upon the legacy of his parents. In 1969, his discovery of a cranium of Australopithecus boisei caused great excitement. A Homo habilis skull (ER 1470) and a Homo erectus skull (ER 3733), discovered in 1972 and 1975, respectively, were among the most significant finds of Leakey's early expeditions. In 1978, an intact cranium of Homo erectus (KNM-ER 3883) was discovered. In 1984, he made his most important discovery—"Turkana Boy"—the nearly complete skeleton of a young Homo erectus who died 1.6 million years ago. It was one of the first well-preserved skeletons of that origin ever found.[11]
  • André Leroi-Gourhan (1911-1986), created theories about human evolution involving the notion that the transition to bipedality freed the hands for grasping, and the face for gesturing and speaking, and thus that the development of the cortex, of technology, and of language all follow from the adoption of an upright stance.
  • Kenneth Oakley (1911-1981) is known for his work in the relative dating of fossils by fluorine content, which was instrumental in the exposure in the 1950s of the Piltdown Man hoax.[12]

Notes

  1. Dictionary.com, Paleoanthropology, Dictionary.com Unabridged (v 1.1). Retrieved May 05, 2008.
  2. S.J. Gould, "Men of the Thirty-third Division," Natural History (1990): 12-24.
  3. Frances Wheelhouse, Dart: Scientist and Man of Grit (Hornsby, Australia: Transpareon Press, 2001, ISBN 0908021216).
  4. 4.0 4.1 4.2 4.3 4.4 4.5 C.D. Kreger, Homo sapiens: Introduction, Archaeology.info (2005).
  5. 5.0 5.1 5.2 Ernst Mayr, What Evolution Is (New York: Basic Books, 2001, ISBN 0465044255).
  6. S.J. Gould, "Lucy on the Earth in stasis," Natural History, (1994): 12-20.
  7. 7.0 7.1 Jonathan Wells, Icons of Evolution. (Washington, DC: Regnery Publishing, 2000, ISBN 0895262762)
  8. Encyclopædia Britannica, Davidson Black (2008).
  9. Pat Shipman, The Man who Found the Missing Link: The Extraordinary Life of Eugene Dubois (Diane Publishing Co, 2001, ISBN 075679160X).
  10. Sonia Cole, Leakey's Luck: The Life of Louis Seymour Bazett Leakey, 1903-1972 (Harcourt, 1975, ISBN 0151494568).
  11. 11.0 11.1 Mary Bowman-Kruhm, The Leakeys: A Biography (Westport, CT: Greenwood Press, 2005, ISBN 0313329850).
  12. Encyclopædia Britannica, Kenneth Oakley (2008).

References
ISBN links support NWE through referral fees

  • Bowman-Kruhm, Mary. The Leakeys: A Biography. Westport, CT: Greenwood Press, 2005. ISBN 0313329850.
  • Cole, Sonia. Leakey's Luck: The Life of Louis Seymour Bazett Leakey, 1903-1972. Harcourt, 1975. ISBN 0151494568.
  • Mayr, Ernst. What Evolution Is. New York: Basic Books, 2001. ISBN 0465044255.
  • McKie, Robin. Dawn of Man: The Story of Human Evolution. DK Publishing, 2000. ISBN 978-0789462626.
  • Shipman, Pat. The Man who Found the Missing Link: The Extraordinary Life of Eugene Dubois. Diane Publishing Co, 2001. ISBN 075679160X.
  • Wells, Jonathan. Icons of Evolution. Washington, DC: Regnery Publishing, 2000. ISBN 0895262762.
  • Wheelhouse, Frances. Dart: Scientist and Man of Grit. Hornsby, Australia: Transpareon Press, 2001. ISBN 0908021216.

External links

All links retrieved November 18, 2022.

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