Neanderthal

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For other uses, see Neanderthal (disambiguation).

Neanderthals
H. neanderthalensis La Ferrassie 1 H. neanderthalensis La Ferrassie 1
Conservation status
Prehistoric
Scientific classification
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Primates Family: Hominidae Genus: Homo Species: H. neanderthalensis
Binomial name
†Homo neanderthalensis King, 1864

The Neanderthal (Homo neanderthalensis) or Neandertal was a species of the Homo genus that inhabited Europe and parts of western Asia from about 230,000 to 29,000 years ago, during the Middle Paleolithic period.

Neanderthals were adapted to the cold, as shown by their large braincases, short but robust builds, and large noses — traits selected by nature in cold climates, as observed in modern sub-arctic populations. Their brain sizes have been estimated as larger than "modern" humans, but their brains may in fact have been approximately the same as those of modern humans. On average, Neanderthal males stood about 1.65m tall (just under 5' 6") and were heavily built, and muscular due to their physical activity. Females were about 1.53 to 1.57m tall (about 5'-5'2").

The characteristic style of stone tools in the Middle Paleolithic is called the Mousterian Culture, after a prominent archaeological site where the tools were first found. The Mousterian culture is typified by the wide use of the Levallois technique. Mousterian tools were often produced using soft hammer percussion, such as bones, antlers, and wood, rather than hard hammer percussion, using stone. Near the end of the time of the Neanderthals, they created the Chatelperronian tool style, considered more "advanced" than that of the Mousterian. They either invented the Chatelperronian themselves or "borrowed" elements from the incoming modern humans who are thought to have created the Aurignacian.

Etymology and classification

The term "Neanderthal Man" was coined in 1863 by Irish anatomist William King. Neanderthal is now spelled two ways: The spelling of the German word Thal, meaning "valley or dale", was changed to Tal in the early 20th century, but the former spelling is often retained in English and always in scientific names, while the modern spelling is used in German.

The Neanderthal or "Neander valley" was named after theologian Joachim Neander, who lived there in the late seventeenth century.

The original German pronunciation (regardless of spelling) is with the sound /t/, rather than the sound /θ/ which is typical of the digraph th in English. When used in English, the term may get an anglicised /θ/ or an original /t/, depending on the speaker.

For many years, professionals vigorously debated about whether Neanderthals should be classified as Homo neanderthalensis or as Homo sapiens neanderthalensis, the latter placing Neanderthals as a subspecies of Homo sapiens. However, recent evidence from mitochondrial DNA studies have been interpreted as evidence that Neanderthals were not a subspecies of H. sapiensTemplate:Cite needed. Still, some scientists argue that fossil evidence suggests that the two species interbred, and hence were the same biological species.

Discovery

File:Neandertals48.jpg

Reconstruction of a Neandertal man made in 1948

A Neanderthal skull was first discovered in Forbes' Quarry, Gibraltar, in 1848, eight years prior to the "original" discovery in a limestone quarry of the Neander Valley (near Düsseldorf) in August, 1856, three years before Charles Darwin's On the Origin of Species was published.

The type specimen, dubbed Neanderthal 1, consisted of a skull cap, two femora, three bones from the right arm, two from the left arm, part of the left ilium, fragments of a scapula, and ribs. The workers who recovered this material originally thought it to be the remains of a bear. They gave the material to amateur naturalist Johann Karl Fuhlrott, who turned the fossils over to anatomist Hermann Schaafhausen. The discovery was jointly announced in 1857.

That discovery is now considered the beginning of paleoanthropology. These and other discoveries led to the idea that these remains were from ancient Europeans who had played an important role in modern human origins. The bones of over 400 Neanderthals have been found since.

Physical traits

The following is a list of physical traits that distinguish Neanderthals from modern humans; however, not all of them can be used to distinguish specific Neanderthal populations, from various geographic areas or periods of evolution, from other extinct humans. Also, many of these traits occasionally manifest in modern humans, particularly among certain ethnic groups. Nothing is known about the skin color, the hair, or the shape of soft parts such as eyes, ears, and lips of Neanderthals.

Compared to modern humans, Neanderthals were larger in size and had distinct morphological features, especially of the cranium, which gradually accumulated more derived aspects, particularly in certain relatively isolated geographic regions. Their relatively robust stature is thought to be an adaptation to the cold climate of Europe during the Pleistocene epoch.

File:Neanderthaler.JPG

Reconstruction of a Neandertal man shown in the Neandertal museum in Mettmann

• Cranial
  • Suprainiac fossa, a groove above the inion
  • Occipital bun, a protuberance of the occipital bone that looks like a hair knot
  • Projecting mid-face
  • Globe-shaped skull (from rear)
  • Low, flat, elongated skull
  • Supraorbital torus, a prominent browridge
  • 1200-1700 cm³ skull capacity (10% greater than modern human average)
  • Receding chin
  • Crest on the mastoid process behind the ear opening
  • No groove on canine teeth
  • A space behind the last molars
  • A broad, projecting nose
  • Bony projections on the sides of the nasal opening
  • Distinctive shape of the bony labyrinth in the ear
  • Larger foram in skull for facial blood supply.
• Post-Cranial
  • Considerably more muscular
  • Large round finger tips
  • Barrel-shaped rib cage
  • Long, gracile pelvic pubis (superior pubic ramus);
  • Large kneecaps
  • Long collar bones
  • Short, bowed shoulder blades
  • Thick, bowed shaft of the thigh bones
  • Short shinbones and calf bones

Based on a 2001 study, some commentators speculated that Neanderthals exhibited rufosity, and that some red-headed and freckled humans today share some heritage with Neanderthals. [1] However, other researchers disagree. [2]

Pathology

Within the west Asian and European record there are five broad groups of pathology or injury noted in Neanderthal skeletons.

Fractures

Neanderthals seemed to suffer a high frequency of fractures, especially common on the ribs (Shanidar IV, La Chapelle-aux-Saints ‘Old Man’), the femur (La Ferrassie 1), fibulae (La Ferrassie 2 and Tabun 1), spine (Kebara 2) and skull (Shanidar I, Krapina, Sala 1). These fractures are often healed and show little or no sign of infection, suggesting that they were taken care of during times of incapacitation.

Trauma

Particularly related to fractures are cases of trauma seen on many skeletons of Neanderthals. These usually taking the form of stab wounds, as seen on Shanidar III, whose lung was probably punctured by a stab wound to the chest between the 8-9th ribs. This may have been an intentional attack or merely a hunting accident, either way the man survived for some weeks after his injury before being killed by a rock fall in the Shanidar cave. Other signs of trauma include blows to the head (Shanidar I and IV, Krapina), all of which seemed to have healed, although traces of the scalp wounds are visible on the surface of the skulls.

Degenerative Disease

Arthritis is particularly common in the older Neanderthal population, specifically targeting areas of articulation such as the ankle (Shanidar III), spine and hips (La Chapelle-aux-Saints ‘Old Man’), arms (La Quina 5, Krapina, Feldhofer) knees, fingers and toes. This is closely related to degenerative joint disease, which can range from normal, use-related degeneration to painful, debilitating restriction of movement and deformity and is seen in varying degree in the Shanidar skeletons (I-IV).

Hypoplastic Disease

Dental enamel hypoplasia is an indicator of stress during the development of teeth and records in the striations and grooves in the enamel periods of food scarcity, trauma or disease. A study of 669 Neanderthal dental crowns showed that 75% of individuals suffered some degree of hypoplasia and the nutritional deficiencies were the main cause of hypoplasia and eventual tooth loss. All particularly aged skeletons show evidence of hypoplasia and it is especially evident in the Old Man of La Chapelle-aux-Saints and La Ferrassie 1 teeth.

Infection

Evidence of infections on Neanderthal skeletons is usually visible in the form of lesions on the bone, which are created by systematic infection on areas closest to the bone. Shanidar I has evidence of the degenerative lesions as does La Ferrassie 1, whose lesions on both femurs, tibiae and fibulae are indicative of a systematic infection or carcinoma (malignant tumour/cancer).

Language

The theory that Neanderthals lacked complex language was widespread until 1983, when a Neanderthal hyoid bone was found at the Kebara Cave in Israel. The hyoid is a small bone that holds the root of the tongue in place, a requirement to human speech and, therefore, its presence seems to imply ability to speak. The bone that was found is virtually identical to that of modern humans.

Many people believe that even without the hyoid bone evidence, it is obvious that tools as advanced as those of the Mousterian Era, attributed to Neanderthals, could not have been developed without cognitive skills encompassing some form of spoken language.

A recent study conducted on the Neanderthal hyoid found that due to the physical characteristics of Neanderthals and the fact that their larynx would have been stouter than that of the modern human, the average note emitted by Neanderthals would have been high pitched and sharper than that of modern man, contrary to the media stereotype of Neanderthals having ape-like grunts.

The base of the Neanderthal tongue was positioned higher in the throat, crowding the mouth somewhat. As a result, Neanderthal speech would most likely have been nasalized.

Tools

File:Neanderthal 2D.jpg

Neanderthal Hunter, (American Mus. Nat. Hist.) Template:3d alt

Neanderthal (Middle Paleolithic) archaeological sites show a smaller and different toolkit than have been found in Upper Paleolithic sites, which were perhaps occupied by modern humans that superceded them. Fossil evidence indicating who may have made the tools found in Early Upper Paleolithic sites is still missing.

There is little evidence that Neanderthals used antlers, shell, or other bone materials to make tools; their bone industry was relatively simple. However, there is good evidence that they routinely constructed a variety of stone implements. The Neanderthal (Mousterian) tool kits consisted of sophisticated stone-flakes, task-specific hand axes, and spears. Many of these tools were very sharp. There is also good evidence that they used a lot of wood, objects which are unlikely to have been preserved until today.

Also, while they had weapons, none have yet been found that were used as projectile weapons. They had spears, in the sense of a long wooden shaft with a spearhead firmly attached to it, but these were not spears specifically crafted for flight (perhaps better described as a lance). However, a number of 400,000 year old wooden projectile spears were found at Schöningen in northern Germany. These are thought to have been made by the Neanderthal's ancestors, Homo erectus or Homo heidelbergensis. Generally, projectile weapons are more commonly associated with H. sapiens. The lack of projectile weaponry is an indication of different sustenance methods, rather than inferior technology or abilities.

Although much has been made of the Neanderthal's burial of their dead, their burials were less elaborate than those of anatomically modern humans. The interpretation of the Shanidar IV burials as including flowers, and therefore being a form of ritual burial (Ralph Solecki 1975), has been questioned (Sommer 1999). On the other hand, five of the six flower pollens found with Shanidar IV are known to have had 'traditional' medical uses, even among relatively recent 'modern' populations. In some cases Neanderthal burials include grave goods, such as bison and aurochs bones, tools, and the pigment ochre.

Neanderthals performed a sophisticated set of tasks normally associated with humans alone. For example, they constructed complex shelters, controlled fire, and skinned animals. Particularly intriguing is a hollowed-out bear femur with four holes spaced like four holes in the diatonic scale, claimed by many to have been deliberately bored into it. This flute was found in western Slovenia in 1995, near a Mousterian Era fireplace used by Neanderthals, but its significance is still a matter of dispute.

See also: prehistoric music and Divje Babe.

Ritual defleshing or cannibalism

Intentional burial and the inclusion of grave goods is the most typical representation of ritual behaviour in the Neanderthals and denote a developing ideology. However, another much debated and controversial manifestation of this ritual treatment of the dead comes from the evidence of cut-marks on the bone which has historically been viewed as evidence of cannibalism. Neanderthal bones from various sites (Combe-Grenal and Abri Moula in France, Krapina in Croatia and Grotta Guattari in Italy) have all been cited as bearing cut marks made by stone tools. However, re-evaluation of these marks using high-powered microscopes, comparisons to contemporary butchered animal remains and recent ethnographic cases of excarnation mortuary practises have shown that perhaps this was a case of ritual defleshing. Fragments of bones from Krapina bear marks that are similar to those seen on bones from secondary burials at a Michigan ossuary (14th century AD) and are indicative of removing the flesh of a partially decomposed body. At Grotta Guattari, the apparently purposefully widened base of the skull (for access to the brains) has been shown to be caused by carnivore action, with hyena tooth marks found on the skull and mandible.

Extinction

No one knows why the Neanderthals became extinct and was replaced by Homo sapiens sapiens (Cro-magnon man). There was likely contact between Neanderthal and Cro-magnon populations, and the absence of mitochondrial DNA traceable to the Neanderthals in modern humans has been taken to suggest a species barrier between Neanderthals and Cro-magnon.

Key dates

• 1848: Skull of ancient human found in Forbe's Quarry, Gibraltar. Its significance is not realised at the time.
• 1856: Johann Karl Fuhlrott first recognizes the fossil called “Neanderthal man.”
• 1880: The mandible of a Neanderthal child was found in a secure context, associated with cultural debris, including hearths, Mousterian tools, and bones of extinct animals.
• 1899: Hundreds of Neanderthal bones were described in stratigraphic position in association with cultural remains and extinct animal bones.
• 1908: A nearly complete Neanderthal skeleton in association with Mousterian tools and bones of extinct animals discovered.
• 1953-1957: Shanidar Cave, Northern Iraq: Ralph Solecki uncovers nine Neanderthal skeletons.
• 1975: Erik Trinkaus’s study of Neanderthal feet confirms they walked like modern humans.
• 1987: New thermoluminescence dates in the Levant place Neanderthal levels at Kebara at ca. 60,000 BP and modern humans at Qafzeh to 90,000 BP. These dates are confirmed by ESR dates for Qafzeh (90,000 BP) and Skhul (80,000 BP).
• 1991: New Electron Spin Resonance (ESR) dates for Near Eastern remains show Tabun Neanderthal to be contemporaneous with modern humans from Skhul and Qafzeh.
• 2000: Igor Ovchinnikov, Kirsten Liden, William Goodman et al. retrieve DNA from a late (29,000 BP) Neanderthal infant from Mezmaikaya Cave in the Caucausus.
• 2005: The Max Planck Institute for Evolutionary Anthropology launches a project to reconstruct the Neanderthal genome.

Popular culture

Popular Literature has tended to greatly exaggerate the ape-like gait and related characteristics of the Neanderthals. It has been determined that some of the earliest specimens found in fact suffered from severe arthritis. The Neanderthals were fully bipedal and had a slightly larger average brain capacity than a typical modern human, though it is thought the brain structure was organised differently.

In popular idiom the word Neanderthal is sometimes used as an insult, to suggest that a person combines a deficiency of intelligence and an attachment to brute force, as well as perhaps implying the person is old fashioned or attached to outdated ideas, much in the same way as "dinosaur" is also used. Counterbalancing this are sympathetic literary portrayals of Neanderthals, as in the novel The Inheritors by William Golding and Jean M. Auel's Earth's Children series, or the more serious treatment by palaeontologist Björn Kurtén, in several works including Dance of the Tiger, and British psychologist Stan Gooch in his hybrid-origin theory of humans.

Science fiction has depicted Neanderthals in several ways:

• In The Ugly Little Boy by Isaac Asimov, a Neanderthal child is brought into the present via time travel.
• Michael Crichton's 1976 novel Eaters of the Dead places a small Neanderthal population in Europe as the source of the battles recorded in Beowulf.
• In the Riverworld series, Philip José Farmer introduces a prominent Neanderthal character named Kazz, who interacts with modern humans. Jose Farmer's novella The Alley Man concerns a Neanderthal whose family has survived into modern times.
• Robert Sawyer's Neanderthal Parallax trilogy portrays contact with an alternate world where Neanderthals, not Homo sapiens, became the dominant species. The first book in this series, Hominids, won the Hugo Award in 2003. (Sawyer's 1997 novel Frameshift used Neanderthal DNA as an element of a plot set in modern-day America.)
• In John Darnton's 1996 novel Neanderthal a group of surviving Neanderthals is discovered in the mountains of Afghanistan. In the novel Neanderthals are said to possess the ability to read minds due to their larger cranial capacity.
• In Darwin's Radio by Greg Bear (winner of 2003 Nebula Award), a phenomenon which caused the Neanderthals to die off now threatens modern humans.
• In the 1989 Doctor Who serial Ghost Light a Neanderthal named Nimrod is a butler. He shows good intelligence throughout the serial. Neanderthals also appear in the 2005 Ninth Doctor Adventures spin-off novel Only Human where they also show good intelligence but struggle with concepts such as fiction and lies, and they appear to not understand why humans 'are always making things up'
• Neanderthals are portrayed as having been brought back from extinction to fill low paying jobs in Jasper Fforde's novel Lost in a Good Book, as well as later novels in the series.
• Hindu Epic Ramayana mentions human-like apes and bears resembling Neanderthals.
• A character in Sergio Bonelli's comic book, "Martin Mystère", is a Neanderthal called Java. He is a part of the last surviving Neanderthal population, living in a hidden city in Mongolia.
• A Neanderthal named Java also appeared as a supporting character in the adventures of Metamorpho the Element Man by DC Comics. The name of this character and the 'Java' from Martin Mystère are both inspired by the Java Man remains.
• Down in the Bottomlands by Harry Turtledove is set in an alternate timeline where Europe is still inhabited by Homo neanderthalensis.
• Neanderthals appear as characters in Jean M. Auel's "Earth's Children" Series

See also

• Caveman
• List of neanderthal sites
• Neandertal interaction with Cro-Magnons
• Physical anthropology

External links

Wikibooks has a book on the topic of

Introduction to Paleoanthropology

• BBC.co.uk - 'Neanderthals "mated with modern humans": A hybrid skeleton showing features of both Neanderthal and early modern humans has been discovered, challenging the theory that our ancestors drove Neanderthals to extinction', BBC (April 21, 1999)

• BBC.co.uk - 'Neanderthals "had hands like ours": The popular image of Neanderthals as clumsy, backward creatures has been dealt another blow', Helen Briggs, BBC (March 27, 2003)

• GeoCities.com - 'The Neanderthal Sites at Veldwezelt-Hezerwater, Belgium'
• Greenwych.ca - 'Neanderthal Flute: Oldest Musical Instrument's 4 Notes Matches 4 of Do, Re, Mi Scale - Evidence of Natural Foundation to Diatonic Scale (oldest known musical instrument), Greenwich Publishing

• Greenwych.ca - 'Chewed or Chipped? Who Made the Neanderthal Flute? Humans or Carnivores?' Bob Fink, Greenwich Publishing (March, 2003)

• Humans Edged Out Neanderthals Earlier, Study Says
• IndState.edu - 'Neanderthals: A Cyber Perspective', Kharlena María Ramanan, Indiana State University (1997)
• Krapina.com - 'Krapina: The World's Largest Neanderthal Finding Site'
• Neanderthal.de - 'Neanderthal Museum'
• UniZH.ch - 'Comparing Neanderthals and modern humans: Neanderthals differ from anatomically modern Homo sapiens in a suite of cranial features' (cranio-facial reconstructions), Institut für Informatik der Universität Zürich
• WebShots.com - 'IMG_6922 The Neandertal foot prints' (photo of fossilized footprints discovered in 1970 on volcanic layers near Demirkopru Dam Reservoir, Manisa, Turkey)

References

ISBN links support NWE through referral fees

• C. David Kreger 6/30/00 Homo Neanderthalensis (archive link, was dead)
• Dennis O'Neil 12/6/04 Evolution of Modern Humans Neandertals retrieved 12/26/2004
• Fink, Bob (1997) The Neanderthal Flute... (Greenwich, Canada) ISBN 0912424125
• Hickmann, Kilmer, Eichmann (ed.) (2003) Studies in Music Archaeology III International Study Group on Music Archaeology's 2000 symposium. ISBN 3896466402
• Serre et al. (2004). No evidence of Neandertal mtDNA contribution to early modern humans. PLoS Biology 2 (3): 313–7. PMID 15024415.
• Solecki, R. S. (1975) Shanidar IV, a Neanderthal flower burial in N. Iraq Science 190 (28) 880
• Sommer, J.D. (1999) The Shanidar IV 'Flower Burial': A Reevaluation of Neanderthal Burial Ritual, Cambridge Archaeological Journal, 9 127-129.
• Eva M. Wild, Maria Teschler-Nicola, Walter Kutschera, Peter Steier, Erik Trinkaus & Wolfgang Wanek (19 May 2005) Direct dating of Early Upper Palaeolithic human remains from Mladeč, Nature 435:332–335. link for Nature subscribers

Part of the series on Human evolution

Hominini

Sahelanthropus tchadensis • Orrorin tugenensis • Ardipithecus

Australopithecines

Australopithecus: A. anamensis • A. afarensis • A. bahrelghazali • A. africanus • A. garhi

Paranthropus: P. aethiopicus • P. boisei • P. robustus

Humans and Proto-humans

Kenyanthropus platyops

Homo: H. habilis • H. rudolfensis • H. georgicus • H. ergaster • H. erectus (H. e. lantianensis • H. e. palaeojavanicus • H. e. pekinensis • H. e. soloensis) • H. cepranensis • H. antecessor • H. heidelbergensis • H. neanderthalensis • H. rhodesiensis • H. floresiensis • Homo sapiens (H. s. idaltu • H. s. sapiens)

Topics: Timeline of human evolution • List of human fossils • Human evolutionary genetics

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