Difference between revisions of "Oak" - New World Encyclopedia

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See [[List of Quercus species|List of ''Quercus'' species]]
 
 
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The term '''oak''' can be used as part of the common name of any of several hundred species of [[tree]]s and [[shrub]]s in the [[genus]] ''Quercus'' (from [[Latin]] "oak tree"), and some related genera, notably ''[[Cyclobalanopsis]]'' and ''[[Lithocarpus]]''. The genus is native to the northern hemisphere, and includes [[deciduous]] and [[evergreen]] species extending from cold latitudes to tropical [[Asia]] and the [[Americas]].  
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'''Oak''' is the common name for any of the [[tree#Deciduous versus evergreen|deciduous]] or evergreen [[tree]]s and [[shrub]]s comprising the [[flowering plant]] [[genus]] ''Quercus'' of the beech family [[Fagaceae]], characterized by a fruit that is a [[nut (fruit)|nut]] called an [[acorn]], simple, spirally arranged [[leaf|leaves]], and male flowers in the form of catkins. The term also is used for the durable wood. Oak also appears in the common name of some species in related genera, notably the quite similar acorn-producing ''[[Cyclobalanopsis]]'' and ''[[Lithocarpus]],'' both in the Fagaceae family, as well as some species in unrelated taxa, such as the poison oaks, which are shrubs in the genus Toxicodendron of the Anacardiaceae family.
  
Oaks have spirally arranged [[leaf|leaves]], with a lobed margin in many species; some have serrated leaves or entire leaves with a smooth margin. The [[flower]]s are [[catkin]]s, produced in spring. The [[fruit]] is a [[nut (fruit)|nut]] called an [[acorn]], borne in a cup-like structure known as a [[cupule]]; each acorn contains one seed (rarely two or three) and takes 6-18 months to mature, depending on species. The [[live oak|"live oaks"]] (oaks with [[evergreen]] leaves) are not a distinct group, instead with their members scattered among the sections below.
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There are several hundred [[species]] in the [[Quercus]] genus, many of which are commercially important as the source of hardwood lumber. Oaks also provide many cultural, symbolic, aesthetic, and ecological values. The strength, hardness, elasticity, durability, and attractive grain of the wood leads to its use for furniture, flooring, buildings, caskets, tool handles, and even aging red wines and other alcoholic beverages, among many other uses. The oak tree itself has long been considered sacred by many civilizations (Schonbeck and Frey 2005) and it is a common symbol of [[virtue]] and [[endurance]]. Oaks are widely planted as ornamental trees for shade and aesthetic purposes, and [[Ecology|ecologically]], the acorns and leaves provide food for many animals.  
  
==Classification==
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==Description==
The genus is divided into a number of sections:
 
*Sect. ''Quercus'' (synonyms ''Lepidobalanus'' and ''Leucobalanus''), the [[List of Quercus species#Section Quercus|oaks]] of [[Europe]], Asia and [[North America]]. Styles short; acorns mature in 6 months, sweet or slightly bitter, inside of acorn shell hairless. Leaves mostly lack a bristle on lobe tips, which are usually rounded.
 
*Sect. ''Mesobalanus'', the [[List of Quercus species#Section Mesobalanus|Hungarian oak and its relatives]] of Europe and Asia. Styles long; acorns mature in 6 months, bitter, inside of acorn shell hairless (closely related to sect. ''Quercus'' and sometimes included in it).
 
*Sect. ''Cerris'', the [[List of Quercus species#Section Cerris|Turkey oak and its relatives]] of Europe and Asia. Styles long; acorns mature in 18 months, very bitter, inside of acorn shell hairless. Leaves typically have sharp lobe tips, with bristles at the lobe tip.
 
*Sect. ''Protobalanus'', the [[List of Quercus species#Section Protobalanus|Canyon live oak and its relatives]], in southwest [[United States]] and northwest [[Mexico]]. Styles short, acorns mature in 18 months, very bitter, inside of acorn shell woolly. Leaves typically have sharp lobe tips, with bristles at the lobe tip.
 
*Sect. ''Lobatae'' (synonym ''Erythrobalanus''), the [[List of Quercus species#Section Lobatae|red oaks]] of North America, [[Central America]] and northern [[South America]]. Styles long, acorns mature in 18 months, very bitter, inside of acorn shell woolly. Leaves typically have sharp lobe tips, with spiny bristles at the lobe tip. 
 
 
 
  
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''Quercus'' is native to the northern hemisphere, and includes [[deciduous]] and [[evergreen]] species extending from cold latitudes to tropical [[Asia]] and the [[Americas]], as well as [[Europe]]. The more southern species tend to be [[tree#Deciduous versus evergreen|evergreen]], keeping their leaves all year round, and the more northern are deciduous, losing their leaves in autumn. Many deciduous species turn bright gold or scarlet in the autumn (Schonbeck and Frey 2005).
  
[[List of Quercus species|'''Full list of ''Quercus'' species''']]
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Oaks have spirally arranged [[leaf|leaves]], with a lobed margin in many species; some have serrated leaves or entire leaves with a smooth margin. Small flowers are produced in spring. The male [[flower]]s hang in [[catkin]]s, and profuse amounts of [[pollen]], while the female flowers are in spikes, both on the same tree (Schonbeck and Frey 2005). The female flowers are wind pollinated by the pollen.
  
[[Image:Quercus stellata.jpg|thumb|A hybrid white oak, possibly ''[[Post oak|Quercus stellata]]'' × ''[[Chinkapin oak|Q. muhlenbergii]]'']]
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The [[fruit]] is a [[nut (fruit)|nut]] called an [[acorn]], borne in a cup-like structure known as a [[cupule]]. Each acorn contains one seed (rarely two or three) and takes 6-18 months to mature, depending on species.  
[[Hybrid]]s are common in oaks but usually only between species within the same section; no verified inter-section hybrids are known, except between species of sections ''Quercus'' and ''Mesobalanus'', where several occur.
 
  
The genus ''[[Cyclobalanopsis]]'', here treated as a distinct genus following the [http://flora.huh.harvard.edu:8080/flora/browse.do?flora_id=2&taxon_id=108828 ''Flora of China''], is often included within ''Quercus'' as a distinct subgenus.
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Trees typically are long-lived, with the average life expectancy for most oaks between 200 and 400 years, with some known to be over 800 years old (Schonbeck and Frey 2005).  
  
==Uses==
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Most oaks are placed into the two categories of red (or black) oaks and the white oaks (Schonbeck and Frey 2005). The red oaks (section Quercus; synonym sect. Erythrobalanus), such as the scarlet oak ''(Quercus coccinea)'' have lobes that are sharp tipped, acorns that mature in 18 months, and the inside of the acorn shell is woolly. The white oaks (section Lobate; synonym sect. Lepidobalanus), such as the white oak ''(Quercus alba),'' have leaves that are smooth-lobed, acorns that mature in six months, and the inside of the acorn shell is hairless.
Oaks are [[hardwood]] trees, and the [[wood]] is commonly used in [[furniture]] and flooring. Oak wood has a density of about 0.75 g/cm³.
 
  
The bark of ''Quercus suber'', or [[Cork oak]], is used to produce [[wine]] [[stopper (plug)|stopper]]s (corks). This species grows in the [[Mediterranean Sea]] region, with [[Portugal]], [[Spain]], [[Algeria]] and [[Morocco]] producing most of the world's supply. Some European and American oak species are used to make [[Barrel (storage)|barrel]]s where wine and other spirits are [[Aging barrel|aged]]; the barrels, which are in some cases charred before use, contribute to the taste, aroma, and color of the contents.
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The [[live oak|"live oaks"]] (oaks with [[evergreen]] leaves) are not a distinct group, instead with their members scattered among the sections below.
  
Of the North American oaks, the [[Northern red oak]] ''Quercus rubra'' is the most prized of the red oak group for lumber, all of which is marketed as red oak regardless of the species of origin. The standard for the lumber of the white oak group, all of which is marketed as white oak, is the [[White Oak]] ''Quercus alba''. White Oak is often used to make wine barrels. The wood of [[Pedunculate Oak]] ''Quercus robur'' and [[Sessile Oak]] ''Quercus petraea'' are extensively used in [[Europe]].
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==Classification==
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The genus is divided into a number of sections:
 +
*Sect. ''Quercus'' (synonyms ''Lepidobalanus'' and ''Leucobalanus''), the oaks of [[Europe]], Asia and [[North America]]. Styles short; acorns mature in 6 months, sweet or slightly bitter, inside of acorn shell hairless. Leaves mostly lack a bristle on lobe tips, which are usually rounded.
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*Sect. ''Mesobalanus,'' the Hungarian oak and its relatives of Europe and Asia. Styles long; acorns mature in six months, bitter, inside of hairless acorn shell (closely related to sect. ''Quercus'' and sometimes included in it).
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*Sect. ''Cerris,'' the Turkey oak and its relatives of Europe and Asia. Styles long; acorns mature in 18 months, very bitter, inside of acorn shell hairless. Leaves typically have sharp lobe tips, with bristles at the lobe tip.
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*Sect. ''Protobalanus,'' the Canyon live oak and its relatives, in southwest [[United States]] and northwest [[Mexico]]. Styles short, acorns mature in 18 months, very bitter, inside of acorn shell woolly. Leaves typically have sharp lobe tips, with bristles at the lobe tip.
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*Sect. ''Lobatae'' (synonym ''Erythrobalanus''), the red oaks of North America, Central America and northern [[South America]]. Styles long, acorns mature in 18 months, very bitter, inside of acorn shell woolly. Leaves typically have sharp lobe tips, with spiny bristles at the lobe.
 +
 
 +
[[Image:Quercus stellata.jpg|thumb|Scrub Oak (''Quercus sinuata var. breviloba'') leaves and acorns'']]
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[[Hybrid]]s are common in oaks but usually only between species within the same section; no verified inter-section hybrids are known, except between species of sections ''Quercus'' and ''Mesobalanus,'' where several occur.
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 +
The genus ''[[Cyclobalanopsis]],'' here treated as a distinct genus following the ''Flora of China'' (eFloras), is often included within ''Quercus'' as a distinct subgenus.
  
The [[bark]] of the White Oak is dried and used in medical preparations. Oak bark is also rich in [[tannin]], and is used by tanners for [[tanning]] [[leather]]. Acorns are used for making flour or roasted for acorn coffee.  Oak galls were used for centuries as the main ingredient in manuscript ink, harvested at a specific time of year.
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==Uses==
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Oak is prized wood for construction because of its strength, elasticity, durability, attractive grain, and resistance to pest species. As such, it is used for furniture, flooring, buildings, mine timbers, caskets, tool handles, railroad ties, barrels, and in the past, ships.
  
Japanese oak is used in the making of professional drums from manufacturer [[Yamaha Drums]]. The rough, hard surface of oak gives the drum a brighter and louder tone compared to traditional drum materials such as [[maple]] and [[birch]].
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Oak wood has a density of about 0.75 g/cm³, great strength and hardness, and is very resistant to [[insect]] and [[fungus|fungal]] attack because of its high tannin content. It also has very attractive grain markings, particularly when quarter-sawn. Wide, [[quarter-sawn]] boards of oak have been prized since the [[Middle Ages]] for use in interior paneling of prestigious buildings such as the debating chamber of the [[British House of Commons]] in [[London]], [[England]], and in the construction of fine furniture. Oak wood, from ''Quercus robur'' and ''Q. petraea,'' was used in Europe for the construction of ships until the nineteenth century, and was the principal timber used in the construction of European [[timber framing|timber-framed]] buildings. Today oak[[wood]] is still commonly used for [[furniture]] making and flooring, timber frame buildings, and for veneer production. Oak wood chips are used for [[smoking (cooking)|smoking]] fish, meat, [[cheese]]s, and other foods.
  
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[[Image:Sherry cellar, Solera system, 2003.jpg|thumb|[[Sherry]] maturing in oak barrels]]
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[[Barrel (storage)|Barrel]]s in which red [[wine]]s, [[sherry]], [[brandy]], and spirits such as [[Scotch whisky]] and [[Bourbon whiskey]] are [[Aging barrel|aged]] are made from European and American oak. The use of [[oak (wine)|oak in wine]] can add many different dimensions to wine based on the type and style of the oak. Oak barrels, which may be charred before use, contribute to the color taste and aroma, of the contents, imparting a desirable oaky [[vanillin]] flavor to these drinks. The great dilemma for wine producers is to choose between French and American oakwoods. French oaks (''Quercus robur,'' ''Q. petraea'') give the wine greater refinement and are chosen for best wines, since they increase the price compared to those aged in American oak wood. American oak contributes greater texture and resistance to aging, but produces more violent wine bouquets. In addition to flavor, oak used for wine barrels increases the antioxidant activity of the wines (Schonbeck and Frey 2005.)
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{{readout||left|200px|Oak used for wine barrels increases the antioxidant activity of the wines}}
  
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[[Image:Cork oak trunk section.jpg|thumb|A section of the trunk of a cork oak, ''Quercus suber'']]
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The bark of ''Quercus suber,'' or [[cork oak]], is used to produce [[wine]] [[stopper (plug)|stopper]]s (corks). This species grows in the [[Mediterranean Sea]] region, with [[Portugal]], [[Spain]], [[Algeria]], and [[Morocco]] producing most of the world's supply.
  
==Diseases and pests==
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Of the North American oaks, the [[northern red oak]] ''Quercus rubra'' is the most prized of the red oak group for lumber, all of which is marketed as red oak regardless of the species of origin. The standard for the lumber of the white oak group, all of which is marketed as white oak, is the [[white oak]] ''Quercus alba.'' White oak is often used to make wine barrels. The wood of the [[deciduous]] [[pedunculate oak]] ''Quercus robur'' and [[sessile oak]] ''Quercus petraea'' account for most of the European oak production, but [[live oak|evergreen species]], such as [[holm oak]] ''Quercus ilex,'' and cork oak ''Quercus suber'' also produce valuable timber.
[[Sudden Oak Death]] (''Phytophthora ramorum'') is a [[water mould]] that can kill oaks within just a few weeks. [[Oak Wilt]], caused by the fungus ''Ceratocystis fagacearum'' (a fungus closely related to [[Dutch Elm Disease]]), is also a lethal disease of some oaks, particularly the red oaks (the white oaks can be infected but generally live longer). Other dangers include wood-boring [[beetle]]s, as well as root [[rot]] in older trees which may not be apparent on the outside, often only being discovered when the trees come down in a strong [[gale]]. Oaks are used as food plants by the [[larva]]e of [[Lepidoptera]] species.
 
*''See also [[list of Lepidoptera which feed on Oaks]]''
 
  
K4t13 + J355
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The [[bark]] of the White Oak is dried and used in medical preparations. Oak bark is also rich in [[tannin]], and is used by tanners for [[tanning]] [[leather]]. Acorns are used for making flour or roasted for acorn coffee. Oak galls were used for centuries as the main ingredient in manuscript ink, harvested at a specific time of year.
  
  '''[[8FF1!]]'''
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[[Ecology|Ecologically]], acorns are an important source of food for wildlife, and flour from ground acorns was eaten by Native Americans (Schonbeck and Frey 2005). The tannin has been used in [[leather]] production and some species are used for cork. Oak bark has been used medicinally to treat diarrhea and as an anti-inflammatory, whether as a gargle for sore throats or topically for skin inflammations such as dermatitis (Schonbeck and Frey 2005). 
  
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The leaves and acorns of the oak tree are poisonous to [[horse]]s in large amounts, due to the toxin [[tannic acid]], and causes [[kidney]] damage and [[gastroenteritis]]. Additionally, once horses have a taste for the leaves and acorns, they may seek them out. Therefore, horse owners are encouraged to fence out oak trees from their pasture, especially if forage is scarce. Symptoms of poisoning include lack of appetite, depression, constipation, diarrhea (which may contain blood), blood in urine, and [[equine colic|colic]].
  
G4881 + 1ZZY
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==Diseases and pests==
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[[Sudden oak death]] is a disease from a water mold ''(Phytophthora ramorum)'' that can kill oaks within just a few weeks. [[Oak wilt]], caused by the fungus ''Ceratocystis fagacearum'' (a fungus closely related to [[Dutch elm disease]]), is also a lethal disease of some oaks, particularly the red oaks. (The white oaks can be infected but generally live longer.) Other dangers include wood-boring [[beetle]]s, as well as root [[rot]] in older trees, which may not be apparent on the outside, often only being discovered when the trees come down in a strong [[gale]]. [[Oak apple]]s are [[gall]]s on oaks made by the [[gall wasp]]. The female [[kermes (genus)|kermes]] [[scale insect|scale]] causes galls to grow on [[kermes oak]]. Oaks are used as food plants by the [[larva]]e of [[Lepidoptera]] species.
  
'''[[8FF1!]]'''
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Also another pest would be the [[gypsy moth]]. The gypsy moth is dominant in North America and there are many concerns of the loss of economically critical and ecologically dominant oak species.
  
 
==Cultural significance==
 
==Cultural significance==
[[Image:5ec ger.png|right|thumb|An oak twig is depicted on the back of [[German euro coins|German 1-, 2- and 5-cent coins]]]]
 
The oak is a common [[symbol]] of [[Virtue|strength]] and [[endurance]] and has been chosen as the [[national tree]] of [[England]], [[France]], [[Germany]], the [[United States]] and [[Wales]].
 
  
[[Thor's Oak]] was a sacred tree of the Germanic [[Chatti]] tribe. Its destruction marked the [[Christianisation]] of the heathen tribes by the [[Franks]]{{Fact|date=April 2007}}.
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The oak has long been considered sacred by many civilizations (Schonbeck and Frey 2005). The Oak of Mamre, Abraham's Oak, is considered to be the location where Abraham pitched his tent, and legend states that anyone defacing this tree will lose his or her firstborn son (Schonbeck and Frey 2005). In the [[Bible]], the oak tree at [[Shechem]] is the site where Jacob buries the foreign gods of his people (Gen. 35.4). In addition, Joshua erects a stone under an oak tree as the first covenant of the Lord (Josh. 24.25-7).
  
In the [[Bible]], the oak tree at [[Shechem]] is the site where Jacob buries the foreign gods of his people (Gen. 35.4) . In addition, Joshua erects a stone under an oak tree as the first covenant of the Lord (Josh. 24.25-7). See other [http://www.biblegateway.com/quicksearch/?quicksearch=oak&x=0&y=0 examples] from the Bible.
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In [[Celtic mythology]], the oak is the tree of doors, believed to be a gateway between worlds, or a place where portals could be erected. [[Thor's Oak]] was a sacred tree of the Germanic [[Chatti]] tribe. Its destruction marked the [[Christianization]] of the heathen tribes by the [[Franks]]. In [[Classical mythology]] the oak was a symbol of [[Zeus]] and his sacred tree. An example is the oracle of [[Dodona]], which in prehistory consisted solely of a holy oak. The Druids considered the oak to have both mystical and medicinal significance (Schonbeck and Frey 2005). According to legend, King Arthur's round table was made from one huge slice of an ancient oak tree (Schonbeck and Frey 2005).  
  
In [[Classical mythology]] the oak was a symbol of [[Zeus]] and his sacred tree.
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[[Image:Coat of arms of Estonia.svg|right|thumb|Oak leaves on the [[coat of arms of Estonia]]]]
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The Oak tree is traditionally sacred to [[Serbs]] and is widely used throughout [[Serbia]] on national and regional symbols both old and new. The oak is a common [[symbol]] of [[Virtue|strength]] and [[endurance]].  
  
Several individual oak trees, such as the [[Royal Oak (tree)|Royal Oak]] in Britain and the [[Charter Oak]] in the United States, are of great historical or cultural importance; for a list of important oaks, see [[:Category:Individual oak trees|Individual oak trees]].
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There is a [[proverb]], "Mighty oaks from little [[acorn]]s grow." Another is: "Every majestic oak tree was once a nut who stood his ground."
  
[[Iowa]] has designated the oak as its official [[list of U.S. state trees|state tree]] in 1961, and the White Oak is the state tree of [[Connecticut]], [[Illinois]] and [[Maryland]].
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Oak leaves symbolize rank in the [[United States Armed Forces]]. A gold oak leaf indicates an O-4 ([[Major]] or [[LCDR|Lt. Commander]]), whereas a silver oak leaf indicates an O-5 ([[Lt. Colonel]] or [[Commander]]). Arrangements of oak leaves, acorns and sprigs indicate different branches of the [[United States Navy]] officers.
  
"Ambrosian Oaks" set to the [[Finlandia Hymn]] is the school song of [[St. Ambrose University]] in [[Davenport, Iowa]].
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==Historical note on Linnaean species==
 
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[[Carolus Linnaeus|Linnaeus]] described only five species of oak from eastern North America, based on general leaf form. These were [[White oak]]- ''Q. alba,'' [[Chestnut oak]]- ''Q. Montana,'' [[Red oak]]- ''Q. rubra,'' [[Willow oak]]- ''Q. phellos,'' and [[Water oak]]- ''Q. nigra.'' Because he was dealing with confusing leaf forms, the ''Q. prinus'' and ''Q. rubra'' specimens actually included mixed foliage of more than one species. For that reason, some taxonomists in the past proposed different names for these two species (''Q. Montana'' and ''Q. borealis,'' respectively), but the original Linnaean names have now been lectotypified with only the specimens in Linnaeus' herbarium that refer to the species the names are applied to now.
The oak is the emblem of [[County Londonderry]] in [[Northern Ireland]], as  a vast amount of the county was covered in forests of the tree until relatively recently. The name of the county comes from the city of [[Derry]], which originally in [[Irish language|Irish]] was known as ''Doire'' meaning ''oak''.
 
 
 
There is a [[proverb]], "Mighty oaks from little [[acorn]]s grow."  Another is:  "Every majestic oak tree was once a nut who stood his ground."
 
  
Many woods are connected to certain birth months, according to the Irish, and oak is the wood of June/July. However, in some variations, rosewood has been known to be June's wood {{Fact|date=February 2007}}.
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==References==
  
[[Raleigh, NC|Raleigh]], [[North Carolina]] has been nicknamed the "City of Oaks."
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* Byfield, L. 1990. ''An oak tree.'' London: Collins Educational. ISBN 0003135268.
 +
 
 +
* eFloras. n.d. [http://flora.huh.harvard.edu:8080/flora/browse.do?flora_id=2&taxon_id=108828 Cyclobalanopsis]. ''Flora of China'' 4: 380. Retrieved November 19, 2007.  
  
The [[Romania national rugby union team]] is nicknamed ''The Oaks.''
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* Logan, W. B. 2005. ''Oak: The Frame of Civilization.'' New York: W. W. Norton. ISBN 0393047733.
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* Paterson, R. T. 1993. ''Use of Trees by Livestock. 5. Quercus.'' Chatham: Natural Resources Institute. ISBN 085954365X.
  
In [[Celtic mythology]] it is the tree of doors, believed to be a gateway between worlds, or a place where portals could be erected.
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* Royston, A. 2000. ''Life Cycle of an Oak Tree.'' Oxford: Heinemann Library. ISBN 0431083916.  
  
Oak leaves are used to symbolize rank in the [[United States Armed Forces]]. A gold oak leaf indicates an O-4 ([[Major]] or [[LCDR|Lt. Commander]]), whereas a silver oak leaf indicates an O-5 ([[Lt. Colonel]] or [[Commander]]). Arrangements of oak leaves, acorns and sprigs indicate different branches of the [[United States Navy]] [[List of United States Navy staff corps|Staff corps]] officers.
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* Savage, S. 1994. ''Oak Tree.'' Hove: Wayland. ISBN 0750211962.
  
==Historical note on Linnaean species==
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* Schonbeck, J., and R. J. Frey. 2005. Oak. In J. L. Longe, ''The Gale Encyclopedia of Alternative Medicine.'' Farmington Hills, Mich: Thomson/Gale. ISBN 0787693960.
[[Carolus Linnaeus|Linnaeus]] described only five species of oak from eastern North America, based on general leaf form. These were White oak, ''Q. alba'', Chestnut oak, ''Q. montana'', Red oak, ''Q. rubra'', Willow oak, ''Q. phellos'', and Water oak, ''Q. nigra''. Because he was dealing with confusing leaf forms, the ''Q. prinus'' and ''Q. rubra'' specimens actually included mixed foliage of more than one species. For that reason, some taxonomists in the past proposed different names for these two species (''Q. montana'' and ''Q. borealis'', respectively), but the original Linnaean names have now been lectotypified with only the specimens in Linnaeus' herbarium that refer to the species the names are applied to now.
 
  
==References and notes==
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* Tansley, A. G. 1952. ''Oaks and Oak Woods.'' London: Methuen.
<References/>
 
  
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==Gallery==
 
<gallery>
 
<gallery>
 
Image:Raunkiaer.jpg|A [[Pedunculate oak]] in [[Denmark]]
 
Image:Raunkiaer.jpg|A [[Pedunculate oak]] in [[Denmark]]
Line 106: Line 113:
 
Image:Oakbark.jpg|Bark of ''[[Quercus robur]]''
 
Image:Oakbark.jpg|Bark of ''[[Quercus robur]]''
 
Image:Spanish-moss-tree.jpg|[[Southern live oak]] with [[spanish moss]]
 
Image:Spanish-moss-tree.jpg|[[Southern live oak]] with [[spanish moss]]
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Image:Quercus mongolica mongolian oak MN 2007.JPG|''Quercus mongolica'' Mongolian Oak in [[Minnesota Landscape Arboretum]]
 
</gallery>
 
</gallery>
  
==External links==
 
{{commonscat|Quercus}}
 
* [http://flora.huh.harvard.edu:8080/flora/browse.do?flora_id=1&taxon_id=127839 Flora of North America: ''Quercus'']
 
* [http://flora.huh.harvard.edu:8080/flora/browse.do?flora_id=2&taxon_id=127839 Flora of China: ''Quercus'']
 
*[http://rbg-web2.rbge.org.uk/cgi-bin/nph-readbtree.pl/feout?FAMILY_XREF=&GENUS_XREF=Quercus&SPECIES_XREF=&TAXON_NAME_XREF=&RANK= Flora Europaea: ''Quercus'']
 
*[http://www.bcgov.net/bftlib/oaktrees.htm Oak Trees of the Lowcountry (South Carolina) — Beaufort County Library]
 
[[Category:Fagaceae]]
 
  
{{credit|133233424}}
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[[Category:Life sciences]]
 
[[Category:Life sciences]]
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[[Category:Plants]]
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{{credit|Oak|172505839|List_of_Quercus_species|171707429}}

Latest revision as of 02:37, 8 December 2009

Oaks
Foliage and acorns of Quercus robur
Foliage and acorns of Quercus robur
Scientific classification
Kingdom: Plantae
Phylum: Magnoliophyta
Class: Magnoliopsida
Order: Fagales
Family: Fagaceae
Genus: Quercus
L.

Oak is the common name for any of the deciduous or evergreen trees and shrubs comprising the flowering plant genus Quercus of the beech family Fagaceae, characterized by a fruit that is a nut called an acorn, simple, spirally arranged leaves, and male flowers in the form of catkins. The term also is used for the durable wood. Oak also appears in the common name of some species in related genera, notably the quite similar acorn-producing Cyclobalanopsis and Lithocarpus, both in the Fagaceae family, as well as some species in unrelated taxa, such as the poison oaks, which are shrubs in the genus Toxicodendron of the Anacardiaceae family.

There are several hundred species in the Quercus genus, many of which are commercially important as the source of hardwood lumber. Oaks also provide many cultural, symbolic, aesthetic, and ecological values. The strength, hardness, elasticity, durability, and attractive grain of the wood leads to its use for furniture, flooring, buildings, caskets, tool handles, and even aging red wines and other alcoholic beverages, among many other uses. The oak tree itself has long been considered sacred by many civilizations (Schonbeck and Frey 2005) and it is a common symbol of virtue and endurance. Oaks are widely planted as ornamental trees for shade and aesthetic purposes, and ecologically, the acorns and leaves provide food for many animals.

Description

Quercus is native to the northern hemisphere, and includes deciduous and evergreen species extending from cold latitudes to tropical Asia and the Americas, as well as Europe. The more southern species tend to be evergreen, keeping their leaves all year round, and the more northern are deciduous, losing their leaves in autumn. Many deciduous species turn bright gold or scarlet in the autumn (Schonbeck and Frey 2005).

Oaks have spirally arranged leaves, with a lobed margin in many species; some have serrated leaves or entire leaves with a smooth margin. Small flowers are produced in spring. The male flowers hang in catkins, and profuse amounts of pollen, while the female flowers are in spikes, both on the same tree (Schonbeck and Frey 2005). The female flowers are wind pollinated by the pollen.

The fruit is a nut called an acorn, borne in a cup-like structure known as a cupule. Each acorn contains one seed (rarely two or three) and takes 6-18 months to mature, depending on species.

Trees typically are long-lived, with the average life expectancy for most oaks between 200 and 400 years, with some known to be over 800 years old (Schonbeck and Frey 2005).

Most oaks are placed into the two categories of red (or black) oaks and the white oaks (Schonbeck and Frey 2005). The red oaks (section Quercus; synonym sect. Erythrobalanus), such as the scarlet oak (Quercus coccinea) have lobes that are sharp tipped, acorns that mature in 18 months, and the inside of the acorn shell is woolly. The white oaks (section Lobate; synonym sect. Lepidobalanus), such as the white oak (Quercus alba), have leaves that are smooth-lobed, acorns that mature in six months, and the inside of the acorn shell is hairless.

The "live oaks" (oaks with evergreen leaves) are not a distinct group, instead with their members scattered among the sections below.

Classification

The genus is divided into a number of sections:

  • Sect. Quercus (synonyms Lepidobalanus and Leucobalanus), the oaks of Europe, Asia and North America. Styles short; acorns mature in 6 months, sweet or slightly bitter, inside of acorn shell hairless. Leaves mostly lack a bristle on lobe tips, which are usually rounded.
  • Sect. Mesobalanus, the Hungarian oak and its relatives of Europe and Asia. Styles long; acorns mature in six months, bitter, inside of hairless acorn shell (closely related to sect. Quercus and sometimes included in it).
  • Sect. Cerris, the Turkey oak and its relatives of Europe and Asia. Styles long; acorns mature in 18 months, very bitter, inside of acorn shell hairless. Leaves typically have sharp lobe tips, with bristles at the lobe tip.
  • Sect. Protobalanus, the Canyon live oak and its relatives, in southwest United States and northwest Mexico. Styles short, acorns mature in 18 months, very bitter, inside of acorn shell woolly. Leaves typically have sharp lobe tips, with bristles at the lobe tip.
  • Sect. Lobatae (synonym Erythrobalanus), the red oaks of North America, Central America and northern South America. Styles long, acorns mature in 18 months, very bitter, inside of acorn shell woolly. Leaves typically have sharp lobe tips, with spiny bristles at the lobe.
Scrub Oak (Quercus sinuata var. breviloba) leaves and acorns

Hybrids are common in oaks but usually only between species within the same section; no verified inter-section hybrids are known, except between species of sections Quercus and Mesobalanus, where several occur.

The genus Cyclobalanopsis, here treated as a distinct genus following the Flora of China (eFloras), is often included within Quercus as a distinct subgenus.

Uses

Oak is prized wood for construction because of its strength, elasticity, durability, attractive grain, and resistance to pest species. As such, it is used for furniture, flooring, buildings, mine timbers, caskets, tool handles, railroad ties, barrels, and in the past, ships.

Oak wood has a density of about 0.75 g/cm³, great strength and hardness, and is very resistant to insect and fungal attack because of its high tannin content. It also has very attractive grain markings, particularly when quarter-sawn. Wide, quarter-sawn boards of oak have been prized since the Middle Ages for use in interior paneling of prestigious buildings such as the debating chamber of the British House of Commons in London, England, and in the construction of fine furniture. Oak wood, from Quercus robur and Q. petraea, was used in Europe for the construction of ships until the nineteenth century, and was the principal timber used in the construction of European timber-framed buildings. Today oakwood is still commonly used for furniture making and flooring, timber frame buildings, and for veneer production. Oak wood chips are used for smoking fish, meat, cheeses, and other foods.

Sherry maturing in oak barrels

Barrels in which red wines, sherry, brandy, and spirits such as Scotch whisky and Bourbon whiskey are aged are made from European and American oak. The use of oak in wine can add many different dimensions to wine based on the type and style of the oak. Oak barrels, which may be charred before use, contribute to the color taste and aroma, of the contents, imparting a desirable oaky vanillin flavor to these drinks. The great dilemma for wine producers is to choose between French and American oakwoods. French oaks (Quercus robur, Q. petraea) give the wine greater refinement and are chosen for best wines, since they increase the price compared to those aged in American oak wood. American oak contributes greater texture and resistance to aging, but produces more violent wine bouquets. In addition to flavor, oak used for wine barrels increases the antioxidant activity of the wines (Schonbeck and Frey 2005.)

Did you know?
Oak used for wine barrels increases the antioxidant activity of the wines
A section of the trunk of a cork oak, Quercus suber

The bark of Quercus suber, or cork oak, is used to produce wine stoppers (corks). This species grows in the Mediterranean Sea region, with Portugal, Spain, Algeria, and Morocco producing most of the world's supply.

Of the North American oaks, the northern red oak Quercus rubra is the most prized of the red oak group for lumber, all of which is marketed as red oak regardless of the species of origin. The standard for the lumber of the white oak group, all of which is marketed as white oak, is the white oak Quercus alba. White oak is often used to make wine barrels. The wood of the deciduous pedunculate oak Quercus robur and sessile oak Quercus petraea account for most of the European oak production, but evergreen species, such as holm oak Quercus ilex, and cork oak Quercus suber also produce valuable timber.

The bark of the White Oak is dried and used in medical preparations. Oak bark is also rich in tannin, and is used by tanners for tanning leather. Acorns are used for making flour or roasted for acorn coffee. Oak galls were used for centuries as the main ingredient in manuscript ink, harvested at a specific time of year.

Ecologically, acorns are an important source of food for wildlife, and flour from ground acorns was eaten by Native Americans (Schonbeck and Frey 2005). The tannin has been used in leather production and some species are used for cork. Oak bark has been used medicinally to treat diarrhea and as an anti-inflammatory, whether as a gargle for sore throats or topically for skin inflammations such as dermatitis (Schonbeck and Frey 2005).

The leaves and acorns of the oak tree are poisonous to horses in large amounts, due to the toxin tannic acid, and causes kidney damage and gastroenteritis. Additionally, once horses have a taste for the leaves and acorns, they may seek them out. Therefore, horse owners are encouraged to fence out oak trees from their pasture, especially if forage is scarce. Symptoms of poisoning include lack of appetite, depression, constipation, diarrhea (which may contain blood), blood in urine, and colic.

Diseases and pests

Sudden oak death is a disease from a water mold (Phytophthora ramorum) that can kill oaks within just a few weeks. Oak wilt, caused by the fungus Ceratocystis fagacearum (a fungus closely related to Dutch elm disease), is also a lethal disease of some oaks, particularly the red oaks. (The white oaks can be infected but generally live longer.) Other dangers include wood-boring beetles, as well as root rot in older trees, which may not be apparent on the outside, often only being discovered when the trees come down in a strong gale. Oak apples are galls on oaks made by the gall wasp. The female kermes scale causes galls to grow on kermes oak. Oaks are used as food plants by the larvae of Lepidoptera species.

Also another pest would be the gypsy moth. The gypsy moth is dominant in North America and there are many concerns of the loss of economically critical and ecologically dominant oak species.

Cultural significance

The oak has long been considered sacred by many civilizations (Schonbeck and Frey 2005). The Oak of Mamre, Abraham's Oak, is considered to be the location where Abraham pitched his tent, and legend states that anyone defacing this tree will lose his or her firstborn son (Schonbeck and Frey 2005). In the Bible, the oak tree at Shechem is the site where Jacob buries the foreign gods of his people (Gen. 35.4). In addition, Joshua erects a stone under an oak tree as the first covenant of the Lord (Josh. 24.25-7).

In Celtic mythology, the oak is the tree of doors, believed to be a gateway between worlds, or a place where portals could be erected. Thor's Oak was a sacred tree of the Germanic Chatti tribe. Its destruction marked the Christianization of the heathen tribes by the Franks. In Classical mythology the oak was a symbol of Zeus and his sacred tree. An example is the oracle of Dodona, which in prehistory consisted solely of a holy oak. The Druids considered the oak to have both mystical and medicinal significance (Schonbeck and Frey 2005). According to legend, King Arthur's round table was made from one huge slice of an ancient oak tree (Schonbeck and Frey 2005).

Oak leaves on the coat of arms of Estonia

The Oak tree is traditionally sacred to Serbs and is widely used throughout Serbia on national and regional symbols both old and new. The oak is a common symbol of strength and endurance.

There is a proverb, "Mighty oaks from little acorns grow." Another is: "Every majestic oak tree was once a nut who stood his ground."

Oak leaves symbolize rank in the United States Armed Forces. A gold oak leaf indicates an O-4 (Major or Lt. Commander), whereas a silver oak leaf indicates an O-5 (Lt. Colonel or Commander). Arrangements of oak leaves, acorns and sprigs indicate different branches of the United States Navy officers.

Historical note on Linnaean species

Linnaeus described only five species of oak from eastern North America, based on general leaf form. These were White oak- Q. alba, Chestnut oak- Q. Montana, Red oak- Q. rubra, Willow oak- Q. phellos, and Water oak- Q. nigra. Because he was dealing with confusing leaf forms, the Q. prinus and Q. rubra specimens actually included mixed foliage of more than one species. For that reason, some taxonomists in the past proposed different names for these two species (Q. Montana and Q. borealis, respectively), but the original Linnaean names have now been lectotypified with only the specimens in Linnaeus' herbarium that refer to the species the names are applied to now.

References
ISBN links support NWE through referral fees

  • Byfield, L. 1990. An oak tree. London: Collins Educational. ISBN 0003135268.
  • eFloras. n.d. Cyclobalanopsis. Flora of China 4: 380. Retrieved November 19, 2007.
  • Logan, W. B. 2005. Oak: The Frame of Civilization. New York: W. W. Norton. ISBN 0393047733.
  • Paterson, R. T. 1993. Use of Trees by Livestock. 5. Quercus. Chatham: Natural Resources Institute. ISBN 085954365X.
  • Royston, A. 2000. Life Cycle of an Oak Tree. Oxford: Heinemann Library. ISBN 0431083916.
  • Schonbeck, J., and R. J. Frey. 2005. Oak. In J. L. Longe, The Gale Encyclopedia of Alternative Medicine. Farmington Hills, Mich: Thomson/Gale. ISBN 0787693960.
  • Tansley, A. G. 1952. Oaks and Oak Woods. London: Methuen.

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