Difference between revisions of "Toucan" - New World Encyclopedia

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{{Taxobox
 
{{Taxobox
| color = pink
 
 
| name = Toucan
 
| name = Toucan
| image = PortraitOfAToucan.jpg
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| image = Pteroglossus-torquatus-001.jpg
| image_caption = [[Keel-billed Toucan]]
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| image_caption = [[Collared Aracari]] (Pteroglossus torquatus)
| image_width = 220px
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| image_width = 250px
 
| regnum = [[Animal]]ia
 
| regnum = [[Animal]]ia
 
| phylum = [[chordate|Chordata]]
 
| phylum = [[chordate|Chordata]]
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| familia = '''Ramphastidae'''
 
| familia = '''Ramphastidae'''
 
| familia_authority = [[Nicholas Aylward Vigors|Vigors]], 1825
 
| familia_authority = [[Nicholas Aylward Vigors|Vigors]], 1825
| subdivision_ranks = [[Genera]]
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| subdivision_ranks = [[Genus|Genera]]
 
| subdivision =  
 
| subdivision =  
 
''[[Andigena]]''<br />
 
''[[Andigena]]''<br />
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}}
 
}}
  
'''Toucans''' are [[near passerine]] [[bird]]s from the [[neotropics]]. They are brightly marked and have large, colorful [[beak|bill]]s. The family includes five [[genus|genera]] and about forty different [[species]].
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'''Toucans''' are a family, '''Ramphastidae''', of [[near-passerine]] [[bird]]s from the [[neotropics]] (i.e. Southern Mexico, Central, South American, and Caribbean region).  The family is most closely related to the [[Capitonidae|American barbets]]. They are brightly marked and have large, colorful bills. The family includes five [[genus|genera]] and about forty different [[species]]. The name of this bird group is derived from [[Tupi language|Tupi]] ''tucana'', via [[French language|French]].
  
Toucans range in size from the Lettered Aracari ''(Pteroglossus inscriptus)'', at 130 g (4.6 oz) and 29 cm (11.5 inches), to the [[Toco Toucan]] ''(Ramphastos toco)'', at 680 g (1.5 lb) and 63 cm (25 inches). Their bodies are short (of comparable size to a [[crow]]'s) and thick. The tail is rounded, and varies in length from half the length to the whole length of the body. The neck is short and thick, and at the base of the head is a huge, brightly-colored beak that measures, in some large species, more than half the length of the body. A toucan's tongue is long, narrow, grey, and singularly frayed on each side, adding to its sensitivity as an organ of taste.  
+
==Morphology==
 +
Toucans range in size from the [[Lettered Aracari]] (''Pteroglossus inscriptus''), at 130 g (4.6 oz) and 29 cm (11.5 inches), to the [[Toco Toucan]] (''Ramphastos toco''), at 680 g (1.5 lb) and 63 cm (29 inches). Their bodies are short (of comparable size to a [[crow|crow's]]) and compact. The tail is rounded and varies in length, from half the length to the whole length of the body. The neck is short and thick. The wings are small, as they are forest-dwelling birds who only need to travel short distances, and are often of about the same span as the bill-tip-to-tail-tip measurements of the bird.
  
The legs of a toucan are strong and rather short. Their toes are arranged in pairs with the first and fourth toes turned backward. Males and females are the same color. The feathers in the genus containing the largest toucans are generally black, with touches of white, yellow, and scarlet. The underparts of the [[araçari]]s (smaller toucans) are yellow, crossed by one or more black or red bands, and the edges of the beak are saw-toothed. The [[toucanet]]s have mostly green plumage with blue markings.  
+
The legs of a toucan are strong and rather short. Their toes are arranged in pairs with the first and fourth toes turned backward. The majority of toucans do not show any [[sexual dimorphism]] in their coloration, the genus ''[[Selenidera]]'' being the most notable exception to this rule (hence their common name, "dichromatic toucanets"). However, the bills of female toucans are usually shorter, deeper and sometimes straighter, giving more of a "blocky" impression compared to male bills. The feathers in the genus containing the largest toucans are generally black, with touches of white, yellow, and scarlet. The underparts of the [[araçari]]s (smaller toucans) are yellow, crossed by one or more black or red bands. The [[toucanet]]s have mostly green plumage with blue markings.
  
Toucans are [[frugivorous]] (fruit-eating), but will take [[prey]] such as [[insect]]s and small [[lizards]]. However, the function of the beak in feeding is not known, since many other birds consume these foods without the giant bill to help them. One likely use is to specialize on prey such as [[nestling]]s and [[bat]]s in tree holes. In this view, the beak allows the bird to reach deep into the tree hole to access food unavailable to other birds.
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[[Image:Keel-billed Toucan-27527.jpg|thumb|right|[[Keel-billed Toucan]] (Ramphastos sulfuratus)]]
 +
The colorful, giant [[beak|bill]], which in some large species measure more than half the length of the body, is the hallmark of toucans. Despite its size it is very light, being composed of bone struts with little solid material between them. The bill has forward-facing [[serration]]s resembling teeth, which historically led naturalists to believe that toucans captured fish and were primarily carnivorous, but today we know that they eat mostly fruit. Why the bill is so large and brightly colored is still debated and may be complex. As there is no sexual dimorphism in coloration it is unlikely to be a sexual [[Signalling theory|signal]]; It does aid in their feeding behavior (as they sit in one spot and reach for all fruit in range, thereby reducing energy expenditure). It has also been theorised that the bill may intimidate smaller birds, so that the toucan may plunder nests undisturbed (see Behaviour). Also, the beak allows the bird to reach deep into treeholes to access food unavailable to other birds, and also to depredate suspended nests built by smaller birds.
  
They are [[arboreal]] and nest in tree holes laying 2–4 white [[egg (biology)|egg]]s. The young hatch completely naked, without any [[down feathers|down]]. Toucans are resident breeders and do not [[bird migration|migrate]]. Toucans are usually found in pairs or small flocks.  
+
A toucan's tongue is long (up to 14-15 cm, or 6 inches), narrow, grey, and singularly frayed on each side, adding to its sensitivity as an organ of taste.
  
The name of this bird group is derived from [[Tupi language|Tupi]] ''tucana'', via [[French language|French]].
+
A structural complex probably unique to toucans involves the modification of several tail [[vertebra]]e. The rear three vertebrae are fused and attached to the spine by a ball-and-socket joint. Because of this, toucans may snap their tail forwards until it touches the head.<ref>Reynolds, J. (2002) "Book Review: Handbook of the Birds of the World, Vol. 7. Jacamars to Woodpeckers Edited by Josep del Hoyo, Andrew Elliott and Jordi Sargatal. Lynx Edicions, Barcelona, 2002. ISBN 84-87334-377. 613 pages." Biological Conservation'' '''111''' (2): 280-281 [http://www.sciencedirect.com/science?_ob=ArticleURL&_udi=B6V5X-47RBJXH-1&_user=1495406&_rdoc=1&_fmt=&_orig=search&_sort=d&view=c&_acct=C000053190&_version=1&_urlVersion=0&_userid=1495406&md5=7aace646b0bd410653fa02a5f7aa6e64] </ref> This is the posture in which they sleep, often appearing simply as a ball of feathers, with the tip of the tail sticking out over the head.
  
[[Image:TOUCAN_LOVE.jpg|right|thumbnail|242px|Two [[Toco Toucan]]s]]
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==Behavior==
 +
[[Image:Ramphastos dicolorus nesting.jpg|Right|200px|thumb|Toucans, like this [[Red-breasted Toucan]] (Ramphastos dicolorus), nest in hollows in trees]]
 +
 
 +
Toucans are primarily [[frugivorous]] (fruit eating), but are opportunistically [[omnivorous]] and will take [[prey]] such as [[insect]]s and small [[lizards]].<ref>Remsen, J.V. Remsen, Jr.; Hyde M.A. & A. Chapman. (1993) "The Diets of Neotropical Trogons, Motmots, Barbets and Toucans" ''The Condor'' '''95''' (1): 178-192</ref> Captive toucans have been reported to actively hunt insects in their cages, and it is possible to keep toucans on an insect-only diet. They also plunder nests of smaller birds, taking eggs and nestlings.<ref>Robinson, S.K. (1985) "Coloniality in the Yellow-Rumped Cacique as a Defense against Nest Predators" ''Auk'' '''10''' (3): 506-519</ref> This probably provides a crucial addition of protein to their diet. However, in their range, toucans are the dominant frugivores, and as such play an extremely important ecological role as vectors for seed dispersal of fruiting trees.<ref>Pizo, M.A.; Donatti, C.I.; Guedes, N.M.R. &  M. Galetti (2008) "Conservation puzzle: Endangered hyacinth macaw depends on its nest predator for reproduction" ''Biological Conservation'' '''141''' (3): 792-796 <small>{{doi|10.1016/j.biocon.2007.12.023}}</small></ref>
 +
 
 +
Toucans are [[arboreal]] and typically lay 2&ndash;4 white [[egg (biology)|eggs]] in their nests. They make their nests in already-existing treeholes like natural cavities and holes excavated by other animals such as [[woodpecker]]s - the toucan bill has very limited use as an excavation tool. When the eggs hatch, the young emerge completely naked, without any [[down feathers|down]]. Toucans are resident breeders and do not [[bird migration|migrate]]. Toucans are usually found in pairs or small flocks. They sometimes fence with their bills and wrestle, which scientists hypothesize they do to establish dominance hierarchies.
  
 
==Systematic list==
 
==Systematic list==
{{seealso|Visual gallery of toucans}}
 
  
 
* Genus ''[[Aulacorhynchus]]'' - green toucanets (6-13 species, depending on [[taxonomy]])<!-- Zootaxa918:1 —>
 
* Genus ''[[Aulacorhynchus]]'' - green toucanets (6-13 species, depending on [[taxonomy]])<!-- Zootaxa918:1 —>
* Genus ''[[Selenidera]]'' - dichromatic toucanets
+
* Genus ''[[Selenidera]]'' - dichromatic toucanets (6 species)
**[[Guianan Toucanet]], ''Selenidera culik''
+
* Genus ''[[Andigena]]'' - mountain toucans (4 species) <!-- Zootaxa918:1 —>
**[[Tawny-tufted Toucanet]], ''Selenidera nattereri''
+
* Genus ''[[Pteroglossus]]'' - araçaris (14 species, incl. [[Saffron Toucanet]])
**[[Gold-collared Toucanet]], ''Selenidera reinwardtii''
+
* Genus ''[[Ramphastos]]'' - typical toucans (about 8 species)
***Langsdorff's Toucanet or Green-billed Toucanet, ''Selenidera (reinwardtii) langsdorffii''
 
**[[Gould's Toucanet]], ''Selenidera gouldii'' - sometimes included in ''S. maculirostris'' <!-- Zootaxa918:1 —>
 
**[[Spot-billed Toucanet]], ''Selenidera maculirostris''
 
**[[Yellow-eared Toucanet]], ''Selenidera spectabilis'' <!-- Zootaxa918:1 —>
 
 
 
* Genus ''[[Andigena]]'' - mountain-toucans <!-- Zootaxa918:1 —>
 
** [[Grey-breasted Mountain-toucan]] ''Andigena hypoglauca''
 
<!-- ** ''Andigena hypoglauca hypoglauca''
 
** ''Andigena hypoglauca lateralis'' —>
 
** [[Hooded Mountain-toucan]], ''Andigena cucullata''
 
** [[Plate-billed Mountain-toucan]], ''Andigena laminirostris''
 
** [[Black-billed Mountain-toucan]], ''Andigena nigrirostris''
 
<!-- **''Andigena nigrirostris nigrirostris''
 
**''Andigena nigrirostris occidentalis''
 
**''Andigena nigrirostris spylorhynchus'' —>
 
 
 
* Genus ''[[Pteroglossus]]'' - aracaris (14 species, incl. [[Saffron Toucanet]])
 
  
* Genus ''[[Ramphastos]]'' - typical toucans (about 10 species)
+
==Toucans in Popular Culture==
 +
*[[Toucan Sam]], mascot for Kellogg's [[Froot Loops]] cereal
 +
*The [[Alcohol advertising#Guinness|Guinness beer toucan]]
 +
*In the UK, street crossings for both cyclists and pedestrians are called [[toucan crossing|toucan crossings]].
  
==Toucans in fiction==
+
==References==
* Toucan Sam, mascot of [[Froot Loops]] cereal.
+
{{Reflist}}
* Tookie Tookie, George's pet Toucan from [[George of the Jungle]].
 
* A humorous derivation of the name of the bird features in David McKee's children's book "Two Can Toucan."
 
* Señor Tucán, the Spanish-speaking toucan from the American cartoon [[Dora the Explorer]].
 
* [[Wulffmorgenthaler]] comic strip features a character named 'Toucan kid', who is an offspring of a human and a toucan.
 
* The character [[Lupe Toucan]] from [[My Gym Partner's a Monkey]]
 
*The character [[Poco]] from the American cartoon [[Maya and Miguel]].
 
* In [[The Legend of Zelda: The Wind Waker]], the [[Wizzrobe]], a magician-like monster that dates back to the first game for the NES, highly resembles a slightly anthropomorphic toucan, possessing the brightly colored beak, and wing-like sleeves on its robe.
 
* A joke in the Emperor's New School. "If you can speak toucan, can you also speak threecan and fourcan?"
 
  
 
== External links ==
 
== External links ==
All links retrieved November 21, 2007
 
 
{{commonscat|Ramphastidae}}
 
{{commonscat|Ramphastidae}}
*[http://www.emeraldforestbirds.com/EmeraldGallery.htm Emerald Photo Gallery]
+
*[http://www.emeraldforestbirds.com/EmeraldGallery.htm Extensive Gallery on Toucans]
*[http://www.mangoverde.com/birdsound/fam/fam102.html Toucans] List of Species
+
*[http://www.mangoverde.com/birdsound/fam/fam102.html List of Toucans]
*[http://www.toucans.net/Gallery/ToucanBird.html The Toucan Bird]
+
*[http://www.toucans.net/Gallery/ToucanBird.html Gallery of Toucans]
*[http://ibc.hbw.com/ibc/phtml/familia.phtml?idFamilia=101 The Internet Bird Collection] Toucan Videos
+
*[http://ibc.hbw.com/ibc/phtml/familia.phtml?idFamilia=101 Toucan videos] on the Internet Bird Collection
  
 
[[Category:Life sciences]]
 
[[Category:Life sciences]]
 
[[Category:Animals]]
 
[[Category:Animals]]
 +
[[Category:Birds]]
  
{{Credits|172618901}}
+
{{Credits|Toucan|254193031}}

Revision as of 22:51, 26 November 2008

For other uses, see Toucan (disambiguation).
Toucan
Collared Aracari (Pteroglossus torquatus)
Collared Aracari (Pteroglossus torquatus)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Piciformes
Family: Ramphastidae
Vigors, 1825
Genera

Andigena
Aulacorhynchus
Pteroglossus
Ramphastos
Selenidera

Toucans are a family, Ramphastidae, of near-passerine birds from the neotropics (i.e. Southern Mexico, Central, South American, and Caribbean region). The family is most closely related to the American barbets. They are brightly marked and have large, colorful bills. The family includes five genera and about forty different species. The name of this bird group is derived from Tupi tucana, via French.

Morphology

Toucans range in size from the Lettered Aracari (Pteroglossus inscriptus), at 130 g (4.6 oz) and 29 cm (11.5 inches), to the Toco Toucan (Ramphastos toco), at 680 g (1.5 lb) and 63 cm (29 inches). Their bodies are short (of comparable size to a crow's) and compact. The tail is rounded and varies in length, from half the length to the whole length of the body. The neck is short and thick. The wings are small, as they are forest-dwelling birds who only need to travel short distances, and are often of about the same span as the bill-tip-to-tail-tip measurements of the bird.

The legs of a toucan are strong and rather short. Their toes are arranged in pairs with the first and fourth toes turned backward. The majority of toucans do not show any sexual dimorphism in their coloration, the genus Selenidera being the most notable exception to this rule (hence their common name, "dichromatic toucanets"). However, the bills of female toucans are usually shorter, deeper and sometimes straighter, giving more of a "blocky" impression compared to male bills. The feathers in the genus containing the largest toucans are generally black, with touches of white, yellow, and scarlet. The underparts of the araçaris (smaller toucans) are yellow, crossed by one or more black or red bands. The toucanets have mostly green plumage with blue markings.

Keel-billed Toucan (Ramphastos sulfuratus)

The colorful, giant bill, which in some large species measure more than half the length of the body, is the hallmark of toucans. Despite its size it is very light, being composed of bone struts with little solid material between them. The bill has forward-facing serrations resembling teeth, which historically led naturalists to believe that toucans captured fish and were primarily carnivorous, but today we know that they eat mostly fruit. Why the bill is so large and brightly colored is still debated and may be complex. As there is no sexual dimorphism in coloration it is unlikely to be a sexual signal; It does aid in their feeding behavior (as they sit in one spot and reach for all fruit in range, thereby reducing energy expenditure). It has also been theorised that the bill may intimidate smaller birds, so that the toucan may plunder nests undisturbed (see Behaviour). Also, the beak allows the bird to reach deep into treeholes to access food unavailable to other birds, and also to depredate suspended nests built by smaller birds.

A toucan's tongue is long (up to 14-15 cm, or 6 inches), narrow, grey, and singularly frayed on each side, adding to its sensitivity as an organ of taste.

A structural complex probably unique to toucans involves the modification of several tail vertebrae. The rear three vertebrae are fused and attached to the spine by a ball-and-socket joint. Because of this, toucans may snap their tail forwards until it touches the head.[1] This is the posture in which they sleep, often appearing simply as a ball of feathers, with the tip of the tail sticking out over the head.

Behavior

Toucans, like this Red-breasted Toucan (Ramphastos dicolorus), nest in hollows in trees

Toucans are primarily frugivorous (fruit eating), but are opportunistically omnivorous and will take prey such as insects and small lizards.[2] Captive toucans have been reported to actively hunt insects in their cages, and it is possible to keep toucans on an insect-only diet. They also plunder nests of smaller birds, taking eggs and nestlings.[3] This probably provides a crucial addition of protein to their diet. However, in their range, toucans are the dominant frugivores, and as such play an extremely important ecological role as vectors for seed dispersal of fruiting trees.[4]

Toucans are arboreal and typically lay 2–4 white eggs in their nests. They make their nests in already-existing treeholes like natural cavities and holes excavated by other animals such as woodpeckers - the toucan bill has very limited use as an excavation tool. When the eggs hatch, the young emerge completely naked, without any down. Toucans are resident breeders and do not migrate. Toucans are usually found in pairs or small flocks. They sometimes fence with their bills and wrestle, which scientists hypothesize they do to establish dominance hierarchies.

Systematic list

  • Genus Aulacorhynchus - green toucanets (6-13 species, depending on taxonomy)
  • Genus Selenidera - dichromatic toucanets (6 species)
  • Genus Andigena - mountain toucans (4 species)
  • Genus Pteroglossus - araçaris (14 species, incl. Saffron Toucanet)
  • Genus Ramphastos - typical toucans (about 8 species)

Toucans in Popular Culture

  • Toucan Sam, mascot for Kellogg's Froot Loops cereal
  • The Guinness beer toucan
  • In the UK, street crossings for both cyclists and pedestrians are called toucan crossings.

References
ISBN links support NWE through referral fees

  1. Reynolds, J. (2002) "Book Review: Handbook of the Birds of the World, Vol. 7. Jacamars to Woodpeckers Edited by Josep del Hoyo, Andrew Elliott and Jordi Sargatal. Lynx Edicions, Barcelona, 2002. ISBN 84-87334-377. 613 pages." Biological Conservation 111 (2): 280-281 [1]
  2. Remsen, J.V. Remsen, Jr.; Hyde M.A. & A. Chapman. (1993) "The Diets of Neotropical Trogons, Motmots, Barbets and Toucans" The Condor 95 (1): 178-192
  3. Robinson, S.K. (1985) "Coloniality in the Yellow-Rumped Cacique as a Defense against Nest Predators" Auk 10 (3): 506-519
  4. Pizo, M.A.; Donatti, C.I.; Guedes, N.M.R. & M. Galetti (2008) "Conservation puzzle: Endangered hyacinth macaw depends on its nest predator for reproduction" Biological Conservation 141 (3): 792-796 Digital object identifier (DOI): 10.1016/j.biocon.2007.12.023

External links

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