Difference between revisions of "Rhinoceros (genus)" - New World Encyclopedia

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The two extant species in the ''Rhinoceros'' genus are both found in Asia, the Indian rhino (''R. unicornis'') and the Javan rhinoceros (''R. rhinoceros''). The Sumatran rhinoceros also is found in Asia, but the other two extant rhinos are found in Africa.
 
The two extant species in the ''Rhinoceros'' genus are both found in Asia, the Indian rhino (''R. unicornis'') and the Javan rhinoceros (''R. rhinoceros''). The Sumatran rhinoceros also is found in Asia, but the other two extant rhinos are found in Africa.
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==Evolution==
  
 
The Indian and Javan rhinoceros first appear in the fossil record in Asia around 1.6 million to 3.3 million years ago. Molecular estimates, however, suggest the species may have diverged much earlier, around 11.7 million years ago (Rougard et al. 2001; Xu et al. 1996). Although belonging to the [[type genus]], the Indian and Javan rhinoceros are not believed to be closely related to other rhino species. A detailed [[clade|cladistic]] analysis of the Rhinocerotidae placed ''Rhinoceros'' and the extinct ''Punjabitherium'' in a clade with ''[[Dicerorhinus]]'', the Sumatran rhino. Other studies have suggested the Sumatran rhinoceros is more closely related to the two African species (Cerdeño 1995). The Sumatran Rhino may have diverged from the other Asian rhinos as far back as 15 million years ago (Lacombat 2005; Dinerstein 2003).  
 
The Indian and Javan rhinoceros first appear in the fossil record in Asia around 1.6 million to 3.3 million years ago. Molecular estimates, however, suggest the species may have diverged much earlier, around 11.7 million years ago (Rougard et al. 2001; Xu et al. 1996). Although belonging to the [[type genus]], the Indian and Javan rhinoceros are not believed to be closely related to other rhino species. A detailed [[clade|cladistic]] analysis of the Rhinocerotidae placed ''Rhinoceros'' and the extinct ''Punjabitherium'' in a clade with ''[[Dicerorhinus]]'', the Sumatran rhino. Other studies have suggested the Sumatran rhinoceros is more closely related to the two African species (Cerdeño 1995). The Sumatran Rhino may have diverged from the other Asian rhinos as far back as 15 million years ago (Lacombat 2005; Dinerstein 2003).  
  
 
Fossils of ''Rhinoceros unicornis'' appear in the [[Middle Pleistocene]].  In the [[Pleistocene]] (1,808,000 to 11,550 years BP), the ''Rhinoceros'' genus ranged throughout Southeast Asia and South Asia, with specimens located on [[Sri Lanka]].  Into the [[Holocene]], some rhinoceros lived as far west as [[Gujarat]] and [[Pakistan]] until as recently as 3,200 years ago (Laurie et al. 1983).  
 
Fossils of ''Rhinoceros unicornis'' appear in the [[Middle Pleistocene]].  In the [[Pleistocene]] (1,808,000 to 11,550 years BP), the ''Rhinoceros'' genus ranged throughout Southeast Asia and South Asia, with specimens located on [[Sri Lanka]].  Into the [[Holocene]], some rhinoceros lived as far west as [[Gujarat]] and [[Pakistan]] until as recently as 3,200 years ago (Laurie et al. 1983).  
 
Lacombat, Frédéric. The evolution of the rhinoceros.  In Fulconis 2005, pp. 46–49.
 
Fulconis, R. (ed.) (2005). Save the rhinos: EAZA Rhino Campaign 2005/6. London: European Association of Zoos and Aquaria.
 
 
Dinerstein, Eric (2003), The Return of the Unicorns; The Natural History and Conservation of the Greater One-Horned Rhinoceros, New York: Columbia University Press, ISBN 0-231-08450-1
 
 
j Laurie, W.A.; E.m. Lang, and C.P. Groves (1983). [http://www.jstor.org/pss/3504002 ''Rhinoceros unicornis'']. Mammalian Species (211): 1–6. doi:10.2307/3504002. http://www.science.smith.edu/departments/Biology/VHAYSSEN/msi/default.html.
 
 
  
 
==Indian rhinoceros==
 
==Indian rhinoceros==
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==References==
 
==References==
 +
Lacombat, Frédéric. The evolution of the rhinoceros.  In Fulconis 2005, pp. 46–49.
 +
Fulconis, R. (ed.) (2005). Save the rhinos: EAZA Rhino Campaign 2005/6. London: European Association of Zoos and Aquaria.
 +
 +
Dinerstein, Eric (2003), The Return of the Unicorns; The Natural History and Conservation of the Greater One-Horned Rhinoceros, New York: Columbia University Press, ISBN 0-231-08450-1
 +
 +
j Laurie, W.A.; E.m. Lang, and C.P. Groves (1983). [http://www.jstor.org/pss/3504002 ''Rhinoceros unicornis'']. Mammalian Species (211): 1–6. doi:10.2307/3504002. http://www.science.smith.edu/departments/Biology/VHAYSSEN/msi/default.html.
 +
 +
 
<ref name=Cerdeno>{{Cite journal | last = Cerdeño | first = Esperanza | url = http://digitallibrary.amnh.org/dspace/bitstream/2246/3566/1/N3143.pdf | journal = Novitates | publisher = [[American Museum of Natural History]] | year = 1995 | title = Cladistic Analysis of the Family Rhinocerotidae (Perissodactyla) | issn = 0003-0082 | number = 3143 | accessdate = 2007-11-04}}</ref>   
 
<ref name=Cerdeno>{{Cite journal | last = Cerdeño | first = Esperanza | url = http://digitallibrary.amnh.org/dspace/bitstream/2246/3566/1/N3143.pdf | journal = Novitates | publisher = [[American Museum of Natural History]] | year = 1995 | title = Cladistic Analysis of the Family Rhinocerotidae (Perissodactyla) | issn = 0003-0082 | number = 3143 | accessdate = 2007-11-04}}</ref>   
  

Revision as of 00:03, 9 February 2009

Rhinoceros
Indian rhinoceros (Rhinoceros unicornis)
Indian rhinoceros (Rhinoceros unicornis)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Rhinocerotidae
Genus: Rhinoceros
Linnaeus, 1758
Species

Rhinoceros unicornis
Rhinoceros sondaicus

The genus Rhinoceros of the family Rhinocerotidae includes the one-horned rhinoceroses, of which there are two extant species: the Indian rhinoceros (R. unicornis) and the Javan rhinoceros (R. sondaicus). These are characterized by one horn (actually thickly matted hair) on the upper snout.

The Javan rhinoceros is one of the most endangered large mammals in the world with only 60 individuals surviving, in only two known locations: Java (Indonesia) and Vietnam.

Overview and description

The Rhinoceros genus is one of four extant genera of rhinoceroses in the rhinoceros family (Rhinocerotidae), the others being Ceratotherium (white rhinocerosis, C. simum), Diceros (black rhinoceros, Diceros bicornis), and Dicerorhinus (Sumatran rhinoceros, Dicerorhinus sumatrensis). All extant rhinoceroses are large, thick-skinned herbivores, with three functional toes on each foot, and either one or two horns on the upper snout. Members of the Rhinoceros genus are characterized by just one horn on the upper snout, while the other extant genera are represented by species with two horns, although one of the Sumatran rhino's horns typically is a stub. These projections are not true horns, but rather are epidermal derivatives, composed of a solid mass of thickly matted hair that grows from the skull without skeletal support.

The word "rhinoceros" is of Greek origin; rhino, meaning "nose," and cero, meaning "horn."

The two extant species in the Rhinoceros genus are both found in Asia, the Indian rhino (R. unicornis) and the Javan rhinoceros (R. rhinoceros). The Sumatran rhinoceros also is found in Asia, but the other two extant rhinos are found in Africa.

Evolution

The Indian and Javan rhinoceros first appear in the fossil record in Asia around 1.6 million to 3.3 million years ago. Molecular estimates, however, suggest the species may have diverged much earlier, around 11.7 million years ago (Rougard et al. 2001; Xu et al. 1996). Although belonging to the type genus, the Indian and Javan rhinoceros are not believed to be closely related to other rhino species. A detailed cladistic analysis of the Rhinocerotidae placed Rhinoceros and the extinct Punjabitherium in a clade with Dicerorhinus, the Sumatran rhino. Other studies have suggested the Sumatran rhinoceros is more closely related to the two African species (Cerdeño 1995). The Sumatran Rhino may have diverged from the other Asian rhinos as far back as 15 million years ago (Lacombat 2005; Dinerstein 2003).

Fossils of Rhinoceros unicornis appear in the Middle Pleistocene. In the Pleistocene (1,808,000 to 11,550 years BP), the Rhinoceros genus ranged throughout Southeast Asia and South Asia, with specimens located on Sri Lanka. Into the Holocene, some rhinoceros lived as far west as Gujarat and Pakistan until as recently as 3,200 years ago (Laurie et al. 1983).

Indian rhinoceros

(R. unicornis) and

The Indian Rhinoceros or the Great One-horned Rhinoceros or the Asian One-horned Rhinoceros (Rhinoceros unicornis) is a large mammal found in Nepal, Bhutan, Pakistan and in Assam, India. It is confined to the tall grasslands and forests in the foothills of the Himalayas. The Indian Rhinoceros can run at speeds of up to 25 mph (40 km/h) for short periods of time and is also an excellent swimmer. It has excellent senses of hearing and smell, but relatively poor eyesight.

In size it is equal to that of the white rhino in Africa; together they are the largest of all rhino species. Fully grown males are larger than females in the wild, weighing from 2200- 3000 kg (4,800 - 6,600 lb). Female Indian rhinos weigh about 1600 kg. The Indian Rhino is from 1.7 to 2 m tall (5.7 to 6.7 feet) and can be up to 4m (13 ft) long. The record-sized specimen of this rhino was approximately 3500 kg.

The Great One-Horned Rhinoceros has a single horn; this is present in both males and females, but not on newborn young. The horn, like human fingernails, is pure keratin and starts to show after about 6 years. In most adults the horn reaches a length of about 25 centimeters,[1] but have been recorded up to 57.2 centimeters in length. The nasal horn curves backwards from the nose. Its horn is naturally black.


Javan rhinoceros

the Javan rhinoceros (R. sondaicus).


References
ISBN links support NWE through referral fees

Lacombat, Frédéric. The evolution of the rhinoceros. In Fulconis 2005, pp. 46–49. Fulconis, R. (ed.) (2005). Save the rhinos: EAZA Rhino Campaign 2005/6. London: European Association of Zoos and Aquaria.

Dinerstein, Eric (2003), The Return of the Unicorns; The Natural History and Conservation of the Greater One-Horned Rhinoceros, New York: Columbia University Press, ISBN 0-231-08450-1

j Laurie, W.A.; E.m. Lang, and C.P. Groves (1983). Rhinoceros unicornis. Mammalian Species (211): 1–6. doi:10.2307/3504002. http://www.science.smith.edu/departments/Biology/VHAYSSEN/msi/default.html.


[2]

[3][4]

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  1. Dinerstein 2003, pp. 272
  2. Cerdeño, Esperanza (1995). Cladistic Analysis of the Family Rhinocerotidae (Perissodactyla). Novitates.
  3. Tougard, C. and T. Delefosse, C. Hoenni, and C. Montgelard (2001). Phylogenetic relationships of the five extant rhinoceros species (Rhinocerotidae, Perissodactyla) based on mitochondrial cytochrome b and 12s rRNA genes. Molecular Phylogenetics and Evolution 19 (1): 34–44.
  4. Xu, Xiufeng and Axel Janke, and Ulfur Arnason. The Complete Mitochondrial DNA Sequence of the Greater Indian Rhinoceros, Rhinoceros unicornis, and the Phylogenetic Relationship Among Carnivora, Perissodactyla, and Artiodactyla (+ Cetacea). Molecular Biology and Evolution 1996 13 (9): 1167–1173.