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| − | {{Taxobox
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| − | | color=pink
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| − | | name=Lion
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| − | | status=VU
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| − | | status_system=iucn3.1
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| − | | trend=down
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| − | | status_ref =<ref name=IUCN>{{IUCN2006|assessors=Nowell & Bauer|year=2004|id=15951|title=Panthera leo|downloaded=11 May 2006}} Database entry includes a lengthy justification of why this species is vulnerable </ref>
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| − | | image=Lion waiting in Nambia.jpg
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| − | | image_width=250px
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| − | | image_caption=Male
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| − | | image2=Tree lion 2.jpg
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| − | | image2_width=250px
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| − | | image2_caption=Female (Lioness)
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| − | | regnum=[[Animal]]ia
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| − | | phylum=[[Chordate|Chordata]]
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| − | | classis=[[Mammalia]]
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| − | | ordo=[[Carnivora]]
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| − | | familia=[[Felidae]]
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| − | | genus=''[[Panthera]]''
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| − | | species='''''P. leo'''''
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| − | | binomial=''Panthera leo''
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| − | | binomial_authority=([[Carolus Linnaeus|Linnaeus]], 1758)
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| − | | synonyms=<center>'''''Felis leo '''''<br /><small>([[Carolus Linnaeus|Linnaeus]], 1758)</small></center>
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| − | | range_map=Lion_distribution.svg
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| − | | range_map_width=250px
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| − | | range_map_caption=Distribution of lions in Africa
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| − | }}
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| − | The '''lion''' ''(Panthera leo)'' is a member of the family [[Felidae]] and one of four "[[big cat]]s" in the [[genus]] ''[[Panthera]]''. Reaching 272 [[kilogram|kg]] (600 [[pound (mass)|lb]]), it is the second-largest cat after the [[tiger]]. They currently exist in the wild in [[sub-Saharan Africa]] and in [[Asia]] with a [[critically endangered]] remnant population in northwest [[India]], having disappeared from [[North Africa]], the Middle East and western Asia in historic times. Until the late [[Pleistocene]] (about 10,000 years ago), the lion was the most widespread large land mammal beside man. They were found in most of Africa, much of Eurasia from western Europe to India and the [[Bering land bridge]] and in the Americas from [[Yukon]] to Peru.
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| − | Lions live for approximately 10–14 years in the wild, while in captivity they can live over 20 years. They typically inhabit [[savanna]] and [[grassland]], although they may take to bush and [[forest]]. Lions are unusually [[social animal|social]] compared to other cats. A lion ''pride'' consists of related females and offspring and a small number of dominant males. Groups of female lions typically hunt together, preying mostly on large [[ungulate]]s. The lion is an [[apex predator|apex]] and [[keystone predator]], though will resort to scavenging if the opportunity arises. While lions, in general, do not selectively hunt humans, some have been known to become man-eaters and seek human prey.
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| − | The lion is a [[vulnerable species]], having seen a possibly irreversible population decline of 30 to 50% over the past two decades in its African range;<ref name=IUCN/> populations are untenable outside designated reserves and national parks. Although the cause of the decline is not well-understood, habitat loss and conflicts with humans are currently the greatest causes of concern. Lions have been kept in [[menagerie]]s since [[Roman era|Roman times]] and have been a key species sought after and exhibited in [[zoo]]s the world over since the late 18th century. Zoos are cooperating worldwide in breeding programs for the endangered [[Asiatic Lion|Asiatic subspecies]].
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| − | The male lion is highly distinctive and is easily recognized by its [[mane]]. The lion, particularly the face of the male, is one of the most widely recognized animal symbols in human [[culture]]. Depictions have existed from the [[Upper Paleolithic]] period, with carvings and paintings from the [[Lascaux]] and [[Chauvet Cave]]s, through virtually all ancient and medieval cultures where they historically occurred. It has been extensively depicted in literature, in [[sculpture]]s, in [[painting]]s, on national [[flag]]s, and in contemporary [[film]]s and [[literature]].
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| − | ==Naming and etymology==
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| − | The lion's name, similar in many languages, derives from the [[Latin]] ''leo'',<ref>{{cite book | author=Simpson DP | title=Cassell's Latin Dictionary | publisher=Cassell Ltd. | date=1979 | edition=5th edition | location=London | pages=p. 883 | isbn=0-304-52257-0}}</ref> and before that the [[Ancient Greek]] ''leōn''/λεων.<ref name="Liddell 1980">{{cite book | author=[[Henry George Liddell|Liddell, Henry George]] and [[Robert Scott (philologist)|Robert Scott]] | year=1980 | title=[[A Greek-English Lexicon]] (Abridged Edition) | publisher=[[Oxford University Press]] | location=United Kingdom | isbn=0-19-910207-4}}</ref> The [[Hebrew]] word ''lavi'' (לָבִיא) may also be related,<ref>{{Cite encyclopedia| title=Lion|encyclopedia=Oxford English Dictionary|editor=Simpson, J., Weiner, E. (eds)| year=1989 |edition= 2nd edition| location=Oxford |publisher=Clarendon Press|id=ISBN 0-19-861186-2}}</ref> as well as the [[Ancient Egyptian]] ''rw''.<ref>[http://www.yourdictionary.com/ahd/l/l0190400.html yourdictionary.com]. As in other ancient scripts, in Ancient Egyptian only the consonants are written. No distinction was made between 'l' and 'r'.</ref> It was one of the many species originally described, as ''Felis leo'', by [[Carolus Linnaeus|Linnaeus]] in his 18th century work, ''[[Systema Naturae]]''.<ref>{{la icon}} {{cite book | last=Linnaeus | first=C. | authorlink=Carolus Linnaeus | title=Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. | publisher=Holmiae. (Laurentii Salvii). | date=1758| pages=p. 824 | url=http://dz1.gdz-cms.de/index.php?id=img&no_cache=1&IDDOC=265100}}</ref> The generic component of its scientific designation, ''Panthera leo'', is often presumed to derive from Greek ''pan-'' ("all") and ''ther'' ("beast"), but this may be a [[folk etymology]]. Although it came into English through the classical languages, ''panthera'' is probably of [[East Asian]] origin, meaning "the yellowish animal," or "whitish-yellow".<ref>{{cite web | url=http://www.etymonline.com/index.php?term=panther | title="Panther" | work=Online Etymology Dictionary | publisher=[[Douglas Harper]] | accessdate=December 10, 2007}}</ref>
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| − | ==Taxonomy and evolution==
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| − | [[Image:Panthera leo Kruger Skull.jpg|thumb|left|Skull of a modern lion at Kruger National Park]]
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| − | The oldest lion-like [[fossil]] is known from [[Laetoli]] in [[Tanzania]] and is perhaps 3.5 million years old; some scientists have identified the material as ''Panthera leo''. These records are not well-substantiated, and all that can be said is that they pertain to a ''Panthera''-like felid. The oldest confirmed records of ''Panthera leo'' in Africa are about 2 million years younger.<ref>{{cite journal |last=Werdelin |first=Lars |coauthors=Lewis, Margaret E. |year=2005 |month=June |title=Plio-Pleistocene Carnivora of eastern Africa: species richness and turnover patterns |journal=Zoological Journal of the Linnean Society |volume=144|issue=2 |pages=121–144 |publisher=The Linnean Society of London |url=http://www.ingentaconnect.com/content/bsc/zoj/2005/00000144/00000002/art00001 |accessdate=December 10, 2007}}</ref>
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| − | The closest relatives of the lion are the other ''[[Panthera]]'' species: the [[tiger]], the [[jaguar]] and the [[leopard]]. Morphological and genetic studies reveal that the tiger was the first of these recent species to diverge. About 1.9 million years ago the Jaguar branched off the remaining group, which contained ancestors of the leopard and lion. The Lion and leopard subsequently separated about 1 to 1.25 million years ago from each other.<ref>{{cite journal |last=Yu |first=Li |coauthors=Ya-ping Zhang |month=May |year=2003 |title=Phylogenetic studies of pantherine cats (Felidae) based on multiple genes, with novel application of nuclear β-fibrinogen intron 7 to carnivores |journal=Molecular Phylogenetics and Evolution |volume=35 |issue=2 |pages=483–495 |doi=10.1016/j.ympev.2005.01.017}}</ref>
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| − | ''Panthera leo'' itself evolved in Africa between 1 million and 800,000 years ago before spreading throughout the [[Holarctic]] region;<ref>{{cite journal |last= Yamaguchi |first=Nobuyuki| coauthors=Alan Cooper, Lars Werdelin and David W. Macdonald|year=2004 |month=August |title= Evolution of the mane and group-living in the lion (Panthera leo): a review|journal=Journal of Zoology |issn=1469-7998 |volume=263 |issue=4 |pages=329–42 |doi=10.1017/S0952836904005242 }}</ref> It appeared in Europe for the first time 700,000 years ago with the subspecies ''[[Panthera leo fossilis]]'' at [[Isernia]] in [[Italy]]. From this lion derived the later [[Cave lion]] ''(Panthera leo spelaea)'', which appeared about 300,000 years ago. During the upper [[Pleistocene]] the lion spread to North and South America, and developed into ''Panthera leo atrox'', the [[American lion]].<ref>{{cite book |last=Turner |first=Allen |year=1997 |title=The big cats and their fossil relatives : an illustrated guide to their evolution and natural history |location=New York |publisher=Columbia University Press |isbn=0-231-10229-1 |pages=}}</ref> Lions died out in northern [[Eurasia]] and America at the end of the last [[glaciation]], about 10,000 years ago;<ref name="BurgerJ-Molecular-phylogeny"/> this may have been secondary to the extinction of [[megafauna]].<ref>{{cite web | last = Harington | first = CR | title = American Lion | work = Yukon Beringia Interpretive Centre website | publisher = Yukon Beringia Interpretive Centre | date = 1996 | url = http://www.beringia.com/02/02maina5.html | accessdate = December 10, 2007}}</ref>
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| − | ===Subspecies===
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| − | [[Image:P l Bleyenberghi.jpg|thumb|upright|Southwest African lion ''(Panthera leo bleyenberghi)'']]
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| − | Traditionally 12 recent subspecies of lion were recognized, the largest of which has been recognized as the [[Barbary Lion]].<ref>[http://www.tigerhomes.org/animal/barbary-lion.cfm Barbary Lion - Panthera leo leo - Largest Lion Subspecies] Retrieved December 10, 2007.</ref> The major differences between these subspecies are location, mane appearance, size and distribution. Because these characteristics are very insignificant and show a high individual variability, most of these forms were debatable and probably invalid; additionally, they were often based upon zoo material of unknown original who may have had "striking, but abnormal" morphological characteristics.<ref name="zoos_encyclopedia">{{cite encyclopedia |last=Grisham |first=Jack |editor=Catherine E. Bell |title=Lion |encyclopedia=Encyclopedia of the World's Zoos |volume=Volume 2: G–P |year=2001 |publisher=Fitzroy Dearborn |location=Chofago |id=ISBN1-57958-174-9 |pages=pp.733–739 }}</ref> Today only eight subspecies are usually accepted,<ref name="Conservation-Genetics:Preserving-Genetic-Diversity" /><ref name="BurgerJ-Molecular-phylogeny">{{cite journal |author=Burger, Joachim ''et al.'' |month=March |year=2004 |title=Molecular phylogeny of the extinct cave lion ''Panthera leo spelaea'' |journal=[[Molecular Phylogenetics and Evolution]] |volume=30 |issue=3 |pages=841–849 |doi=10.1016/j.ympev.2003.07.020| url= http://www.uni-mainz.de/FB/Biologie/Anthropologie/MolA/Download/Burger%202004.pdf| accessdate=December 10, 2007 }}</ref> but one of these, the Cape lion formerly described as ''Panthera leo melanochaita'' is probably invalid.<ref name="Conservation-Genetics:Preserving-Genetic-Diversity" />
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| − | Even the remaining seven subspecies might be too much; [[mitochondria]]l variation in recent African lions is modest, which suggests that all sub-Saharan lions could be considered a single subspecies, possibly divided in two main clades: one to the west of the [[Great Rift Valley]] and the other to the east. Lions from [[Tsavo]] in Eastern Kenya are much closer genetically to lions in [[Transvaal]] (South Africa), than to those in the [[Aberdare Range]] in Western Kenya.<ref>{{cite journal |last=Barnett |first=Ross |coauthors=Nobuyuki Yamaguchi, Ian Barnes and Alan Cooper |year=2006 |title=The origin, current diversity and future conservation of the modern lion ''(Panthera leo)'' |journal= Proceedings of the Royal Society B: Biological Sciences|volume=273 |issue=1598 |pages=2119–2125 |doi=10.1098/rspb.2006.3555 |url= http://www.adelaide.edu.au/acad/publications/papers/Barnett%20PRS%20lions.pdf |format=PDF|accessdate=December 10, 2007}}</ref><ref>{{cite journal |last=Dubach |first=Jean| coauthors=et ''al'' |year=2005 |month=January |title=Molecular genetic variation across the southern and eastern geographic ranges of the African lion, ''Panthera leo'' |journal=Conservation Genetics |volume=6 |issue=1 |pages=15–24 |doi=10.1007/s10592-004-7729-6}}</ref>
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| − | ====Recent====
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| − | Eight recent subspecies are recognized today:
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| − | * '''''P. l. persica''''', known as the '''[[Asiatic lion|Asiatic-]]''' or '''South Asian, Persian or Indian lion''', was once widespread from [[Turkey]], across the [[Middle East]], to [[Pakistan]], [[India]] and even [[Bangladesh]]. However, large prides and daylight activity made it easier to poach than tigers or leopards; now around 300 exist in and near the [[Gir Forest]] of India.<ref name ="Asiaticweb">{{cite web | author=Wildlife Conservation Trust of India | title=Asiatic Lion - History | work=Asiatic Lion Information Centre | publisher=Wildlife Conservation Trust of India | date=2006 | url=http://www.asiaticlion.org/asiatic-lion-history.htm | accessdate=December 10, 2007}}</ref>
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| − | * '''''P. l. leo''''', known as the '''[[Barbary lion]]''', is extinct in the wild due to excessive hunting, although captive individuals may still exist. This was the largest of the lion subspecies, at 3–3.5m approx., and weighing over 150 kilograms and more. They ranged from Morocco to Egypt. The last wild Barbary lion was killed in Morocco in 1922.<ref name=CAP/>
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| − | * '''''P. l. senegalensis''''', known as the '''West African lion''', is found in Western Africa, from Senegal to Nigeria.
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| − | * '''''P. l. azandica''''', known as the '''North East Congo lion''', is found in the Northeastern parts of the [[Democratic Republic of the Congo|Congo]].
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| − | * '''''P. l. nubica''''', known as the '''East African-''' or '''Massai lion''', is found in East Africa, from [[Ethiopia]] and Kenya to [[Tanzania]] and [[Mozambique]].
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| − | * '''''P. l. bleyenberghi''''', known as the '''Southwest African-''' or '''Katanga lion'''. It is found in southwestern Africa, [[Zimbabwe]], [[Angola]], [[Katanga]] ([[Democratic Republic of the Congo|Zaire]]).
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| − | * '''''P. l. krugeri''''', known as the '''Southeast African-''' or '''Transvaal lion''', is found in the [[Transvaal]] region of South eastern Africa, including [[Kruger National Park]].
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| − | * '''''P. l. melanochaita''''', known as the '''[[Cape lion]]''', became extinct in the wild around 1860. Results of [[mitochondrial DNA]] research do not support the status as a distinct subspecies. It seems probable that the Cape lion was only the southernmost population of the extant southern African lion.<ref name="Conservation-Genetics:Preserving-Genetic-Diversity" />
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| − | ====Prehistoric====
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| − | Several additional subspecies of lion existed in prehistoric times:
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| − | <!-- prose form please —>
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| − | * '''''P. l. atrox''''', known as the '''[[American lion]]''' or '''American cave lion''', was abundant in the Americas from [[Alaska]] to [[Peru]] in the Pleistocene Epoch until about 10,000 years ago. This form as well as the cave lion are sometimes considered to represent separate species, but recent phylogenetic studies lead to suggest, that they are in fact subspecies of the lion ''(Panthera leo)''.<ref name="BurgerJ-Molecular-phylogeny" /> One of the largest lion subspecies to have existed, its body length is estimated to have been 1.6–2.5 m (5–8 ft).<ref name="Quaternary extinctions">{{cite book |last=Martin |first=P.S. |title=Quaternary Extinctions |year=1984 |publisher= University of Arizona Press|location= Tucson, Arizona|isbn=0-8165-1100-4 }}</ref>
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| − | * '''''P. l. fossilis''''', known as the '''[[Panthera leo fossilis|Early Middle Pleistocene European cave lion]]''', flourished about 500,000 years ago; fossils have been recovered from [[Germany]] and [[Italy]]. <!--(commented out until I can find a ref!!) It was larger than today's African lions, with a maximum head and body length of 2.40 m (8 ft).{{Fact|date=July 2007}}—>
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| − | [[Image:Lascaux-diverticule-félins.jpg|thumb|right|[[Cave lion]]s, Chamber of Felines, [[Lascaux]] caves]]
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| − | * '''''P. l. spelaea''''', known as the '''[[Cave lion|European cave lion]]''', Eurasian cave lion or Upper Pleistocene European cave lion, occurred in Eurasia 300,000 to 10,000 years ago.<ref name="BurgerJ-Molecular-phylogeny" /> It is known from [[Paleolithic]] [[cave paintings]], [[ivory]] carvings, and clay busts,<ref name="Packer00">{{cite journal | last=Packer | first=Craig | coauthors=Jean Clottes | title=When Lions Ruled France | journal=Natural History | volume=| issue=| pages=52–57 | date=November 2000 | url=http://www.lionresearch.org/current_docs/m_pdf/36.pdf | format=PDF | accessdate=December 10, 2007}}</ref> indicating it had protruding ears, tufted tails, faint tiger-like stripes, and that at least some males had a "ruff" or primitive mane around their necks.<ref name ="Koenigswald02"/>
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| − | * '''''P. l. vereshchagini''''', known as the '''East Siberian-''' or '''[[Beringian cave lion]]''', was found in [[Yakutia]] ([[Russia]]), [[Alaska]] ([[USA]]), and the [[Yukon Territory]] ([[Canada]]). Analysis of skulls and mandibles of this lion demonstrate that it is distinct—larger than the European cave lion and smaller than the American cave lion with differing [[skull]] proportions.<ref>{{cite journal |last= Baryshnikov|first=G.F. |coauthors=G. Boeskorov |month= |year=2001 |title=The Pleistocene cave lion, ''Panthera spelaea'' (Carnivora, Felidae) from Yakutia, Russia |journal=Cranium |volume=18 |issue=1 |pages=7–24 |doi=}}</ref><ref name="BurgerJ-Molecular-phylogeny" />
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| − | ====Dubious====
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| − | * '''''P. l. sinhaleyus''''', known as the '''[[Sri Lanka lion]]''', appears to have become extinct [[circa|around]] 39,000 years ago. It is only known from two teeth found in deposits at [[Kuruwita]]. Based on these teeth, P. Deraniyagala erected this subspecies in 1939.<ref>{{cite journal |author=Kelum Manamendra-Arachchi, Rohan Pethiyagoda, Rajith Dissanayake, Madhava Meegaskumbura |year=2005 |month= |title=A second extinct big cat from the late Quaternary of Sri Lanka. |journal=The Raffles Bulletin of Zoology Supplement |volume=12 |pages=423–434 |id= |url=http://rmbr.nus.edu.sg/rbz/biblio/s12/s12rbz423-434.pdf|publisher=National University of Singapore |accessdate= December 10, 2007}}</ref>
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| − | * '''''P. l. europaea''''', known as the '''[[European lion]]''', was probably identical with ''Panthera leo persica'' or ''Panthera leo spelea''; its status as subspecies is unconfirmed. It became extinct around 100 C.E. due to persecution and over-exploitation. Inhabited the [[Balkans]], the [[Italian Peninsula]], southern [[France]] and the [[Iberian Peninsula]]. It was a very popular object of hunting among [[Ancient Rome|Romans]], [[ancient Greece|Greeks]] and [[Macedonians (ethnic group)|Macedonians]].
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| − | * '''''P. l. youngi''''' or ''Panthera youngi'' , known as the '''[[North-Eastern Pleistocene China cave lion]]''', flourished 350,000 years ago.<ref>{{cite journal |last=Harington |first=CR |year=1969 |title=Pleistocene remains of the lion-like cat ''(Panthera atrox)'' from the Yukon Territory and northern Alaska |journal=Canadian Journal Earth Sciences |volume=6 |issue=5 |pages=1277–88 }}</ref> Its relationship to the extant lion subspecies is obscure, and probably represents a distinct species.
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| − | *'''''P. l. maculatus''''', known as the '''[[Marozi]]''' or '''Spotted lion''', is sometimes believed to be a distinct subspecies, but may be an adult lion that has retained its juvenile spotted pattern. If it was a subspecies in its own right, rather than a small number of aberrantly colored individuals, it has been extinct since 1931. A less likely identity is a natural leopard/lion [[Panthera hybrid|hybrid]] commonly known as a [[leopon]].<ref name="shuker">{{cite book | last=Shuker | first=Karl P.N. | authorlink=Karl Shuker | title=Mystery Cats of the World | publisher=Robert Hale | year=1989 | isbn=0-7090-3706-6}}</ref>
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| − | ==Physical characteristics==
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| − | [[Image:HansomeLion 002.jpg|thumb|left|During confrontations with others, the mane makes the lion look bigger than he really is.]]
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| − | The lion is the second largest feline after the [[tiger]]. With powerful legs, a strong [[jaw]], and long canine [[teeth]], the lion can bring down and kill large prey.<ref name=Honolulu>{{cite web |url=http://www.honoluluzoo.org/lion.htm |title=Lion |accessdate=December 10, 2007 |publisher=Honolulu Zoo}}</ref> Lion coloration varies from light buff to yellowish, reddish or dark ochraceous brown. The underparts are generally lighter and the tail tuft is black. The color of the mane varies from blond to black.
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| − | Average listed weights for the lions are between 150–225 kg (330–500 lb) for males, and 120–150 kg (260–330 lb) for females.<ref>{{cite web|url=http://www.bbc.co.uk/nature/wildfacts/factfiles/14.shtml|title=BBC Wildfacts – Lion}}</ref> Nowell and Jackson report average weights of 181 kg for males and 126 kg for females; one male shot near [[Mount Kenya]] was weighed at 272 kg (600 lb).<ref name=CAP>{{cite book |author=Nowell K, Jackson P |title= Wild Cats: Status Survey and Conservation Action
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| − | Plan|url=http://carnivoractionplans1.free.fr/wildcats.pdf |format=PDF |year=1996 |publisher=IUCN/SSC Cat Specialist Group |location= Gland, Switzerland |isbn=2-8317-0045-0 |pages= 17–21|chapter= Panthera Leo}}</ref> Head and body length is 170–250 cm (5 ft 7 in–8 ft 2 in) in males and 140–175 cm (4 ft 7 in–5 ft 9 in) in females; shoulder height is about 123 cm (4 ft) in males and 100 cm (3 ft 3 in) in females. The tail length is 70–100 cm (2 ft 3 in–3 ft 3 in).<ref name="nowak">{{cite book |last=Nowak |first= Ronald M. |year=1999 |title=Walker's Mammals of the World |location=Baltimore |publisher=Johns Hopkins University Press |isbn=0-8018-5789-9}}</ref> The tail ends in a hairy tuft. The tuft conceals a spine, approximately 5 mm long, formed of the final sections of tail bone fused together. The lion is the only felid to have a tufted tail and the function of the tuft and spine are unknown. Absent at birth, the tuft develops around 5½ months of age and readily identifiable at 7 months.<ref name ="Schaller28">Schaller, p. 28</ref>
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| − | ====Mane====
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| − | [[Image:Maneless lion from Tsavo East National Park.png|thumb|right|A maneless male lion from Tsavo East National Park, Kenya]]
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| − | [[Image:wiki lion.jpg|thumb|left|250px|Thermal image of a lion, showing the insulating manes]]
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| − | The mane of the male lion, unique amongst cats, is one of the most distinctive characteristics of the species. It makes the lion appear larger, providing an excellent intimidation display; this aids the lion during confrontations with other lions and with the species' chief competitor in Africa, the [[spotted hyena]].<ref>
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| − | {{cite web | last=Trivedi | first=Bijal P. | title=Are Maneless Tsavo Lions Prone to Male Pattern Baldness? | publisher=National Geographic | date=2005 | url=http://news.nationalgeographic.com/news/2002/04/0412_020412_TVtsavolions.html | accessdate=December 10, 2007}}</ref> The presence, absence, color, and size of the mane is associated with genetic precondition, sexual maturity, climate and [[testosterone]] production; the rule of thumb is the darker and fuller the mane, the healthier the lion.<ref name="Trivedi02">{{cite web | last=Trivedi | first =Bijal P. | title=Female Lions Prefer Dark-Maned Males, Study Finds | work=National Geographic News | publisher=National Geographic | date=Aug 22, 2002 | url=http://news.nationalgeographic.com/news/2002/08/0822_020822_TVlion.html| accessdate= December 10, 2007}}</ref> Research in [[Tanzania]] also suggests mane length signals fighting success in male-male relationships. Darker-maned individuals may have longer reproductive lives and higher offspring survival, although they suffer in the hottest months of the year.<ref name=West>{{cite journal |last=West |first=Peyton M. |coauthors=Craig Parker|year=2002 |month=August |title=Sexual Selection, Temperature, and the Lion's Mane |journal=[[Science (journal)|Science]] |volume=297 |issue=5585 |pages= 1339–1343 |doi=10.1126/science.1073257}}</ref> In prides led by a coalition of two or three males, it is possible that lionesses solicit mating more actively with heavily maned lions.<ref name="Trivedi02"/>
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| − | Scientists once believed that the distinct status of some subspecies could be justified by [[morphology (biology)|morphology]], including the size of the mane. Morphology was used to identify subspecies such as the [[Barbary lion]] and [[Cape Lion]]. Research has suggested, however, that environmental factors influence the color and size of a lion's mane, such as the [[ambient temperature]].<ref name=West/> The cooler ambient temperature in European and North American [[zoo]]s, for example, can result in a heavy mane. Thus the mane is an inappropriate marker for identifying subspecies.<ref name="Conservation-Genetics:Preserving-Genetic-Diversity">{{cite journal |last=Barnett |first=Ross |coauthors=Nobuyuki Yamaguchi, Ian Barnes and Alan Cooper |month=August |year=2006 |title=Lost populations and preserving genetic diversity in the lion ''Panthera leo'': Implications for its ''ex situ'' conservation |journal=Conservation Genetics |volume=7 |issue=4 |pages=507–514 }} <!-- url not accessible at the moment http://abc.zoo.ox.ac.uk/Papers/consgen06_lion.pdf—></ref><ref name="yamaguchi-haddane">{{cite journal |last=Yamaguchi |first=Nobuyuki |coauthors=B. Haddane |month= |year=2002 |title=The North African Barbary lion and the Atlas Lion Project |journal=International Zoo News |volume=49 |pages=465–481}}</ref> However the males of the Asiatic subspecies are characterized by sparser manes than average African lions.<ref name="Menon">{{cite book |last=Menon |first=Vivek |year=2003 |title=A Field Guide to Indian Mammals |location=Delhi |publisher=Dorling Kindersley India |isbn=0-14-302998-3| pages=}}</ref>
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| − | [[Image:White Lion.jpg|thumb|right|[[White lion]]s owe their coloring to a recessive gene. They are rare forms of the subspecies ''Panthera leo krugeri''.]]
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| − | Maneless lions have been reported in [[Senegal]] and [[Tsavo East National Park]] in Kenya, and the original male white lion from Timbavati was also maneless. [[castration|Castrated]] lions have minimal manes. The lack of a mane is found in inbred lion populations; inbreeding also results in poor fertility.<ref>{{cite web | last=Trivedi | first =Bijal P. | title=To Boost Gene Pool, Lions Artificially Inseminated | work=National Geographic News | publisher=National Geographic | date=12 June 2002 | url=http://news.nationalgeographic.com/news/2002/06/0612_020612_TVlion.html| accessdate= December 10, 2007}}</ref> Cave paintings of European [[cave lion]]s show exclusively animals with no mane or just the hint of a mane, suggesting they were more or less maneless.<ref name ="Koenigswald02">{{de icon}} {{cite book |last=Koenigswald |first=Wighart von |year=2002 |title=Lebendige Eiszeit: Klima und Tierwelt im Wandel |location=Stuttgart |publisher=Theiss |isbn=3-8062-1734-3| pages=}}</ref>
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| − | ====White lions====
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| − | The [[white lion]] is not a distinct subspecies, but a special [[Polymorphism (biology)|morph]] with a genetic condition, [[leucism]],<ref name="zoos_encyclopedia"/> that causes paler coloration akin to that of the [[white tiger]]; the condition is similar to [[melanism]], which causes [[black panther]]s. White animals of the Transvaal lion ''(Panthera leo krugeri)'' have been occasionally encountered in and around the [[Kruger National Park]] and the adjacent [[Timbavati Private Game Reserve]] in eastern South Africa, but are more commonly found in [[captive animal|captivity]], where breeders deliberately select them. The unusual cream color of their coats is due to a [[recessive gene]].<ref>{{cite book |last=McBride |first=Chris |title=The White Lions of Timbavati |year=1977 |publisher=E. Stanton |location=Johannesburg |isbn= 0-949997-32-3 }}</ref> They have been reportedly bred in camps in South Africa for use as trophies for canned hunts.<ref>{{cite book |last=Tucker|first=Linda |title=Mystery of the White Lions - Children of the Sun God|year=2003 |publisher=Npenvu Press|location=Mapumulanga |isbn= 0-620-31409-5}}</ref>
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| − | Confirmation of the actual existence of the White lion only came in the late 20th century. For hundreds of years prior, the White lion had been a figment of legend circulating through South Africa, the white [[pelage]] of the animal said to represent the goodness in all creatures. Claimed sightings were first reported in the early 1900s, and continued, infrequently, for almost 50 years until, in 1975, a litter of white lion cubs were found at Timbavati Game Reserve.<ref name="whitelion">[http://www.lairweb.org.nz/tiger/lions.html The rare white lions] Retrieved December 10, 2007.</ref>
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| − | ===Hybrids===
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| − | [[Image:bertramliger.jpg|thumb|left|A [[liger]] is the offspring of a male lion and female [[tiger]].]]
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| − | {{further|[[Panthera hybrid]], [[liger]] and [[tigon]]}}
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| − | Lions have also been known to [[biological reproduction|breed]] with [[tiger]]s (most often the [[Amur Tiger|Amur]] and [[Bengal Tiger|Bengal]] subspecies) to create hybrids called [[liger]]s and [[tigon]]s.<ref>{{cite book |last=Guggisberg |first=C. A. W. |title=Wild Cats of the World |year=1975 |publisher=Taplinger Publishing |location=New York |isbn=0-8008-8324-1 }}</ref> They have also been crossed with [[leopards]] to produce [[leopon]]s,<ref>{{cite book |author=Doi H, Reynolds B |title=The Story of Leopons |year=1967 |publisher=Putnam |location=New York |oclc=469041 <!--Too old for ISBN—>}}</ref> and [[jaguar]]s to produce [[jaglion]]s. The [[marozi]] is reputedly a spotted lion or a naturally occurring leopon, while the [[Congolese spotted lion]] is a complex lion/jaguar/leopard hybrid called a [[lijagulep]]. Such hybrids were once commonly bred in zoos, but this is now discouraged due to the emphasis on conserving species and subspecies. Hybrids are still bred in private menageries and in zoos in [[China]].
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| − | The liger is a cross between a male lion and a tigress.<ref name="liger">{{cite book| last=Markel |first=Scott| coauthors= Darryl León| year= 2003 |title=Sequence Analysis in a Nutshell: a guide to common tools and databases | publisher= O'Reily |location=Sebastopol, California |isbn=0-596-00494-X}}</ref> Because the lion sire passes on a growth-promoting gene, but the corresponding growth-inhibiting gene from the female lion is absent, ligers grow far larger than either parent. They share physical and behavioral qualities of both parent species (spots and stripes on a sandy background). Male ligers are sterile, but female ligers are often fertile. Males have about a 50% chance of having a mane, but if they grow one their manes will be modest: around 50% of a pure lion mane. Ligers are typically between 10 to 12 feet in length, and can be between 800 and 1,000 pounds or more.<ref name="liger" /> The less common tigon is a cross between the lioness and the male tiger.<ref>{{citeweb|url=http://www.britannica.com/eb/article-9001344/tigon|title= tigon - Encyclopædia Britannica Article|accessdate=December 10, 2007}}</ref> <!--(commented out - not really necessary - can reinsert when ref arises) Because the male tiger does not pass on a growth-promoting gene and the lioness passes on a growth inhibiting gene, tigons are often relatively small, only weighing up to 150 kilograms (350 lb),{{Fact|date=August 2007}} which is about 20% smaller than lions. Like ligers, they have physical and behavioral traits from both parental species and males are sterile.—>
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| − | ==Biology and behavior==
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| − | Lions spend much of their time resting and are inactive for about 20 hours per day.<ref name="Schaller122">Schaller, p. 122</ref> Although lions can be active at any time, their activity generally peaks after dusk with a period of socializing, grooming and defecating. Intermittent bursts of activity follow through the night hours to dawn, when hunting most often takes place. They spend an average two hours a day walking and 50 minutes eating.<ref name="Schaller120">Schaller, p. 120–1</ref>
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| − | ===Hunting and diet===
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| − | [[Image:Female Lion.JPG|thumb|left|While a lion's teeth are sharp, prey is usually killed by strangulation.]]
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| − | Lions are powerful animals that usually hunt in groups and stalk their chosen prey. They can reach speeds of {{convert|59|km:h|mi:h|-1|abbr=on}},<ref name="Schaller233">Schaller, p. 233</ref> though only for short bursts,<ref name="Schaller2478">Schaller, p. 247–8</ref> so they have to be close to their prey before starting the attack. Lions take advantage of factors that reduce visibility; many kills take place near some form of cover or at night.<ref name="Schaller237">Schaller, p. 237</ref> They sneak up to the victim until they reach a distance of about 30 [[metre|m]] (98 [[foot (unit of length)|feet]]) or less. Typically, several female lions work together and encircle the herd from different points. Once they have closed with a herd, they usually target the closest prey. The attack is short and powerful, the lion attempting to catch the victim with a fast rush and final leap. The prey is usually killed by strangulation.<ref>{{cite web |title=About lions - Ecology and behavior |author=Dr Gus Mills |publisher=African Lion Working Group |url=http://www.african-lion.org/lions_e.htm |accessdate=December 10, 2007 }}</ref>
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| − | The lion's [[predation|prey]] consists mainly of large mammals, with a preference for [[wildebeest]], [[impala]]s, [[zebra]]s, [[African buffalo|buffalo]] and [[warthog]]s in Africa and [[nilgai]], [[wild boar]]s and several deer species in India. Many other species are hunted based on availability, mainly [[ungulate]]s weighing between 50 and 300 kg such as [[kudu]], [[hartebeest]], [[gemsbok]] and [[eland]].<ref name="nowak">Ronald M. Nowak: ''Walker's Mammals of the World''.</ref> Occasionally, lions take relatively small species such as [[Thomson's gazelle]] or [[Springbok Antelope|springbok]]. Lions, hunting in groups, are capable of taking down most animals, even healthy adults, but they rarely attack very large prey such as buffalo bulls or fully grown male giraffes, due to the danger of injury.<ref name="home">{{citeweb|url=http://home.intekom.com/ecotravel/Guides/Wildlife/Vertebrates/Mammals/Big_5/Lion/african-lion-hunting-habits.htm |title=African Lion Hunting Habits in South Africa and Southern Africa|accessdate=December 10, 2007}}</ref> They normally feed on mammals no larger than 550 [[kg]], which excludes most adult [[hippopotamus]]es, [[rhinoceros]]es, [[elephant]]s, [[giraffe]]s and buffalos.<ref>{{cite journal|last=Hayward |first= Matt W. |coauthors=Graham Kerley| year=2005| title=Prey preferences of the lion ''(Panthera leo)'' |journal=Journal of Zoology |volume=267 |issue=3 |pages=309–322 |doi=10.1017/S0952836905007508}}</ref> In some areas, lions specialize in hunting atypical prey-species; this is the case at the [[Chobe National Park#Geography and ecosystems|Savuti]] river, where they prey on young elephants.<ref>{{cite web|last=Kemp| first=Leigh| url=http://www.go2africa.com/botswana/chobe/savuti/legendary-elephant-eaters.asp| title=The Elephant Eaters of the Savuti| publisher=go2africa |accessdate=December 10, 2007}}</ref> Park guides in the area reported that the lions, driven by extreme hunger, started taking down baby elephants, and then moved on to adolescents and, occasionally, fully grown adults.<ref>{{cite news |title=King of the jungle defies nature with new quarry |first=Damien |last=Whitworth |date=October 9, 2006 |publisher=The Australian |url=http://www.news.com.au/story/0,23599,20547955-38195,00.html |accessdate=December 10, 2007 }}</ref> In the Kruger National Park, giraffes are regularly hunted.<ref name ="Pienaar69">{{cite journal |author=Pienaar U de V |year=1969 |title=Predator-prey relationships amongst the larger mammals of the Kruger National Park |journal=Koedoe |volume=12 |pages=108-76 |id= }}</ref> Lions also attack domestic livestock; in India cattle contribute significantly to their diet.<ref name=Menon>Vivek Menon: ''A Field Guide to Indian Mammals''.</ref> They are capable of killing other predators such as [[leopard]]s, [[cheetah]]s, [[hyena]]s and [[African Wild Dog|wild dog]]s, as well as scavenging animals either dead from natural causes or killed by other predators.<ref name="Schaller213">Schaller, p. 213</ref> A lion may gorge itself and eat up to 30 kg (66 lb) in one sitting;<ref name="simba">{{cite book |last= Guggisberg|first=C. A. W. |title=Simba: the life of the lion. |year=1961 |publisher=Howard Timmins |location=Cape Town |isbn= }}</ref> if it is unable to consume all the kill it will rest for a few hours before consuming more. On a hot day, the pride may retreat to shade leaving a male or two to stand guard.<ref name="Schaller2706">Schaller, p. 270–76</ref> An adult female lion requires an average of about 5 kg (11 lb) of meat per day, a male about 7 kg (15.4 lb).<ref>{{cite web |title=Lions |publisher=Honolulu Zoo |url=http://www.honoluluzoo.org/lion.htm |accessdate=December 10, 2007 }}</ref>
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| − | [[Image:Male Lion and Cub Chitwa South Africa Luca Galuzzi 2004.JPG|thumb|250px|right|Lion and cub eating a [[Cape Buffalo]]]]
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| − | Because lions hunt in open spaces where they are easily seen by their prey, cooperative hunting increases the likelihood of a successful hunt; this is especially true with larger species. Teamwork also enables them to defend their prey more easily against other large predators such as hyenas, which can be attracted by [[vulture]]s over kilometers in open savannas. Lionesses do most of the hunting. Males attached to prides do not usually participate, except when hunting large animals such as buffalo and giraffe. In group hunts, each lioness has a favored position in the group, either stalking prey on the "wing" then attacking, or moving a smaller distance in the centre of the group and capturing prey in flight from other lionesses.<ref>{{cite journal|author=Stander, P. E.|title=Cooperative hunting in lions: the role of the individual|journal=Behavioral Ecology and Sociobiology|volume=29|issue=6|pages=445—454|year=1992|doi=10.1007/BF00170175}}</ref> Young lions first display stalking behavior around three months of age, although they do not participate in actual hunting until they are almost a year old. They begin to hunt effectively when nearing the age of two.<ref name="Schaller153">Schaller, p. 153</ref>
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| − | ===Reproduction===
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| − | Most lionesses will have reproduced by the time they are four years of age.<ref name ="Schaller29">Schaller, p. 29</ref> Lions do not [[mating|mate]] at any specific time of year, and the females are [[Estrous cycle#Frequency|polyestrous]].<ref name ="Schaller174">Schaller, p. 174</ref> Like other cats, the male lion's penis has spines which point backwards. Upon withdrawal of the penis, the spines rake the walls of the female's vagina, which may cause ovulation.<ref>{{cite book |last=Asdell |first=Sydney A. |title=Patterns of mammalian reproduction |year=1993|origyear= 1964 |publisher=Cornell University Press|location=Ithaca |isbn=978-0-8014-1753-5}}</ref> A female may mate with more than one male when she is in heat;<ref name ="Schaller142">Schaller, p. 142</ref> during a mating bout, which could last several days, the couple copulates twenty to forty times a day and are likely to forgo hunting. Lions reproduce very well in captivity.
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| − | [[Image:Lion pair2.jpg|thumb|upright|left|During a mating bout, a couple may copulate twenty to forty times a day for several days.]]
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| − | The average gestation period is around 110 days,<ref name ="Schaller174"/> the female giving birth to a litter of one to four cubs. Lionesses in a pride will synchronize their reproductive cycles so that they cooperate in the raising and suckling of the young, who suckle indiscriminately from any or all of the nursing females in the pride. Cubs are usually born and initially kept hidden from view in thickets or sheltered areas. They weigh 1.2–2.1 kg at birth and are almost helpless, beginning to crawl a day or two after birth and walking around three weeks of age.<ref name ="Schaller143">Schaller, p. 143</ref> Weaning occurs after six to seven months. In the wild, competition for food is fierce, and as many as 80% of the cubs will die before the age of two.<ref>{{cite book |last=Macdonald |first=David |year=1984 |title=The Encyclopedia of Mammals|publisher=Facts on File|location=New York |pages=31|isbn= 0-87196-871-1}}</ref>
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| − | When one or more new males take over a pride and oust the previous master(s), the conquerors often kill any remaining cubs;<ref name="Packpus83">{{cite journal |author=Packer, C., Pusey, A. E. |year=1983 |month=May |title= Adaptations of female lions to infanticide by incoming males|journal= American Naturalist |volume=121 |issue=5 |pages=716–728 |url=http://www.lionresearch.org/current_docs/6.pdf |accessdate=December 10, 2007}}</ref> females do not again become fertile and receptive until the cubs grow up or die. The male lions reach maturity at about 3 years of age and are capable of taking over another pride at 4–5 years old. They begin to age and weaken between 10 and 15 years of age at the latest.<ref>{{cite book |last=Crandall |first=Lee S. |title=The management of wild animals in captivity |year=1964 |publisher=University of Chicago Press |location=Chicago |oclc=557916 }}</ref> This leaves a short window for their own offspring to be born and mature—the fathers have to procreate as soon as they take over the pride. The lioness will often attempt to defend her cubs fiercely from a usurping male, but such actions are rarely successful, as he usually kills all the previous top male's cubs that are less than two years old and the female is much lighter and has less strength than the male. However, success is more likely when a group of 3 or 4 mothers within the pride join forces against one male.<ref name="Packpus83"/>
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| − | One scientific study reports that both males and females may interact [[animal sexuality|homosexually]].<ref>{{cite book |last=Bagemihl |first=Bruce |year=1999 |title=Biological Exuberance: Animal Homosexuality and Natural Diversity |location=New York |publisher=St. Martin's Press |isbn=0-312-19239-8| pages=pp. 302–305}}</ref><ref>
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| − | {{cite web | last =Srivastav | first =Suvira | title =Lion, Without Lioness | work =TerraGreen: News to Save the Earth | publisher=Terragreen | date=15–31 December 2001 | url=http://www.teri.res.in/teriin/terragreen/issue3/feature.htm | accessdate =December 10, 2007}}</ref> Male lions pair-bond for a number of days and initiate homosexual activity with affectionate nuzzling and caressing, leading to mounting and thrusting. A study found that about 8% of mountings have been observed to occur with other males, while female pairings are held to be fairly common in captivity but have not been observed in the wild.
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| − | ===Health===
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| − | Though adult lions have no natural predators, evidence suggests that the majority die violently from humans or other lions.<ref name ="Schaller183">Schaller, p. 183</ref> Various species of [[tick]] commonly infest the ears, neck and groin regions of most lions.<ref name ="Schaller184">Schaller, p. 184</ref><ref>{{cite book |last=Yeoman |first=G.|coauthors= Jane B. Walker |title=The ixodid ticks of Tanzania|year=1967 |publisher=Commonwealth Institute of Entomology |location=London |oclc=955970}}</ref> Adult forms of several species of the tapeworm genus ''[[Taenia (tapeworm)|Taenia]]'' have been isolated from intestines, the lions having ingested larval forms from antelope meat.<ref>{{de icon}}{{cite journal |last=Sachs |first=R |year=1969|title=Untersuchungen zur Artbestimmung und Differenzierung der Muskelfinnen ostafrikanischer Wildtiere [Differentiation and species determination of muscle-cysticerci in East African game animals] |journal=Zeitschrift für tropenmedizin und Parasitologie |volume=20 |issue=1 |pages=39–50 |pmid=5393325}}</ref> Lions in the [[Ngorongoro Crater]] were afflicted by an outbreak of stable fly ''([[Stomoxys calcitrans]])'' in 1962; this resulted in lions becoming covered in bloody bare patches and emaciated. Lions sought unsuccessfully to evade the biting flies by climbing trees or crawling into hyena burrows; many perished or emigrated as the population dropped from 70 to 15 individuals.<ref>{{cite journal |last=Fosbrooke |first=Henry |year=1963 |title=The stomoxys plague in Ngorongoro |journal=East African Wildlife Journal |volume=1|pages=124–6}}</ref> A more recent outbreak in 2001 killed six lions.<ref name ="Nkwame06">{{cite news | last=Nkwame|first= Valentine M | title=King of the jungle in jeopardy | publisher=The Arusha Times | date=September 9, 2006 | url=http://www.arushatimes.co.tz/2006/36/features_10.htm | accessdate=December 10, 2007}}</ref> Lions, especially in captivity, are vulnerable to the [[Canine distemper]] [[virus]] (CDV), [[feline immunodeficiency virus]] (FIV) and [[feline infectious peritonitis]] (FIP).<ref name="zoos_encyclopedia"/> CDV is spread through domestic dogs and other [[Carnivora|carnivores]]; a 1994 outbreak in [[Serengeti National Park]] resulted in many lions developing neurological symptoms such as seizures. During the outbreak, several lions died from pneumonia and [[encephalitis]].<ref>{{cite journal |author=M.E. Roelke-Parker ''et al.'' |year=1996 |month=February |title=A canine distemper epidemic in Serengeti lions ''(Panthera leo)'' |journal= Nature|volume=379 |issue= |pages=441–445 |url=http://www.lionresearch.org/current_docs/17.pdf |format=PDF|accessdate=December 10, 2007 |doi=10.1038/379441a0|pmid=8559247}}</ref> FIV, which is similar to [[HIV]] while not known to adversely affect lions, is worrisome enough in its effect in domestic cats that the Species Survival Plan recommends systematic testing in captive lions. It occurs with high to endemic frequency in several wild lion populations, but is mostly absent from Asiatic and Namibian lions.<ref name="zoos_encyclopedia"/>''
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| − | ===Group organization===
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| − | Lions are predatory [[carnivore]]s who manifest two types of social organization. Some are ''residents,'' living in groups, called ''prides''.<ref name ="Schaller33">Schaller, p. 33</ref> The pride consists of usually around five or six related females, their cubs of both sexes, and a group of one to four males known as a ''coalition'' who mate with the adult females. Others are ''nomads'', ranging widely and moving sporadically, either singularly or in pairs.<ref name ="Schaller33"/> Note that a lion may switch lifestyles; nomads may become residents and vice versa. The area a pride occupies is called a ''pride area'', whereas that by a nomad is a ''range''.<ref name ="Schaller33"/> Why [[Social behavior|sociality]]—the most pronounced in any cat species—has developed in lions is the subject of much debate. Increased hunting success appears an obvious reason, but this is less than sure upon examination: coordinated hunting does allow for more successful predation, but also ensures that non-hunting "cheaters" reduce per capita caloric intake. Other benefits include possible [[kin selection]] (better to share food with a related lion than with a stranger), protection of the young, maintenance of territory, and individual insurance against injury and hunger.<ref name=CAP/>
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| − | Being smaller and more agile than males, and lacking the conspicuous mane, lionesses do the pride's hunting, while the stronger males patrol the territory and protect the pride. There is no clear hierarchy with food: male lions often eat animals killed by lionesses but will never share food they have killed themselves; they will take food from cubs but are more likely to share with cubs than lionesses, which are more likely to share with each other. There is more sharing with larger kills.<ref name ="Schaller133">Schaller, p. 133</ref>
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| − | Both males and females defend the pride against intruders. Some individual lions consistently lead the defense against intruders, while others lag behind.<ref>{{cite journal |last=Heinsohn|first= R. |coauthors= C. Packer|year=1995 |title=Complex cooperative strategies in group-territorial African lions |journal=Science |volume=269 |issue=5228 |pages=1260–1262 |doi=10.1126/science.7652573 |pmid=7652573}}</ref> These "laggards" are not punished by leaders. Possibly laggards provide other services to the group so that leaders forgive them.<ref>{{cite journal |last=Morell|first= V. |year=1995 |title=Cowardly lions confound cooperation theory |journal=Science |volume=269 |issue=5228 |pages=1216–1217|doi=10.1126/science.7652566}}</ref> An alternative hypothesis is that there is some reward associated with being a leader who fends off intruders.<ref>{{cite journal |last=Jahn|first= GAry C. |year=1996 |title= Lioness Leadership|journal=Science |volume=271 |issue=5253 |pages=1215}}</ref> The leading male or males often have to defend against outside males attempting to take over the pride. Females form a stable social unit in a pride and will not tolerate outside females; <ref name ="Schaller37">Schaller, p. 37</ref> membership only changes with the births and deaths of lionesses,<ref name ="Schaller39">Schaller, p. 39</ref> though some females do leave and become nomadic.<ref name ="Schaller44">Schaller, p. 44</ref> Subadult males on the other hand, leave the pride when they reach maturity at around 2–3 years of age.<ref name ="Schaller44"/>
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| − | ===Communication===
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| − | When resting, lion socialization occurs through a number of behaviors, and the animal's expressive movements are highly developed. The most common peaceful tactile gestures are head rubbing and social licking,<ref name ="Schaller85">Schaller, p. 85</ref> which have been compared with grooming in primates.<ref>{{cite book |editor=Desmond Morris|last=Sparks |first=J |title=Primate Ethology |chapter=Allogrooming in primates:a review |year=1967 |publisher=Aldine |location=Chicago |isbn=0-297-74828-9}} (2007 edition: 0-202-30826-X)</ref> Head rubbing—nuzzling one's forehead, face and neck against another lion—appears to be a form of greeting,<ref>{{de icon}}{{cite book |last=Leyhausen |first=Paul |title=Verhaltensstudien an Katzen|edition= 2nd ed. |year=1960 |publisher=Paul Parey |location=Berlin |isbn=3-489-71836-4 }}</ref> as it is seen often after an animal has been apart from others, or after a fight or confrontation. Males tend to rub other males, while cubs and females rub females.<ref name ="Schaller858">Schaller, p. 85–8</ref> Social licking often occurs in tandem with head rubbing; it is generally mutual and the recipient appears to express pleasure. The head and neck are the most common parts of the body licked, which may have arisen out of utility, as a lion cannot lick these areas individually.<ref name ="Schaller8891">Schaller, p. 88–91</ref>
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| − | Lions have an array of facial expressions and body postures that serve as visual gestures.<ref name ="Schaller92102">Schaller, p. 92–102</ref> Their repertoire of vocalizations is also large; variations in intensity and pitch, rather than discrete signals, appear central to communication. Lion sounds include snarling, purring, hissing, coughing, miaowing, woofing and roaring. Lions most often roar at night; the sound, which can be heard from a distance of 8 km, is used to advertise the animal's presence.<ref name ="Schaller10313">Schaller, p. 103-13</ref>
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| − | ==Distribution and habitat ==
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| − | [[Image:Map Guj Nat Parks Sanctuary.png|thumb|left|The Gir Forest in the State of Gujarat, India is the last natural range of the 300-odd wild [[Asiatic Lion]]s. Plans are afoot to re-introduce some to [[Palpur-Kuno Wildlife Sanctuary]] in the neighboring State of [[Madhya Pradesh]] in India.]]
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| − | In Africa, lions can be found in savannah grasslands with scattered ''[[Acacia]]'' trees which serve as shade;<ref>{{cite book |last=Rudnai|first= Judith A. |title=The social life of the lion |year=1973 |publisher=s.n. |location=Wallingford |isbn=0-85200-053-7 }}</ref> their habitat in India is a mixture of dry savannah forest and very dry deciduous scrub forest.<ref>{{cite web | title=The Gir - Floristic | work=Asiatic Lion Information Centre | publisher=Wildlife Conservation Trust of India | date=2006 | url=http://www.asiaticlion.org/gir-floristic.htm | accessdate=December 10, 2007}}</ref> In relatively recent times the habitat of lions spanned the southern parts of [[Eurasia]], ranging from [[Greece]] to [[India]], and most of [[Africa]] except the central [[rainforest]]-zone and the [[Sahara]] desert. [[Herodotus]] reported that lions had been common in Greece around 480 B.C.E.; they attacked the baggage camels of the Persian king [[Xerxes I of Persia|Xerxes]] on his march through the country. [[Aristotle]] considered them rare by 300 B.C.E. and by 100 C.E. extirpated.<ref name="Schaller5">Schaller, p. 5</ref> A population of the [[Asiatic lion]] survived until the 10th century in the [[Caucasus]], their last [[Europe]]an outpost.<ref>{{cite book| last=Heptner| first= V.G.| coauthors=A. A. Sludskii| year=1989| title=Mammals of the Soviet Union: Volume 1, Part 2: Carnivora (Hyaenas and Cats)|publisher=Amerind| location=New York| isbn=9004088768}}</ref>
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| − | The species was eradicated from [[Palestine]] by the [[Middle Ages]] and from most of the rest of Asia after the arrival of readily available firearms in the 18th century. Between the late 19th and early 20th century they became extinct in [[North Africa]] and the [[Middle East]]. By the late 19th century the lion had disappeared from [[Turkey]] and most of northern India,<ref name=zoos_encyclopedia /><ref>{{cite web |title=Past and present distribution of the lion in North Africa and Southwest Asia. |date=2001 |publisher=Asiatic Lion Information Centre |url=http://www.asiatic-lion.org/distrib.html |accessdate=December 10, 2007 }}</ref> while the last sighting of a live Asiatic Lion in [[Iran]] was in 1941 (between [[Shiraz]] and Jahrom, [[Fars]] province), though the corpse of a lioness was found on the banks of [[Karun]] river, [[Khuzestan]] province in 1944. There are no subsequent reliable reports from Iran.<ref name="simba"/> The subspecies now survives only in and around the [[Gir Forest]] of northwestern India.<ref name ="Asiaticweb"/> About 300 lions live in a 1,412 km² (558 square miles) sanctuary in the state of [[Gujarat]], which covers most of the forest. Their numbers are slowly increasing.<ref>{{cite web | author=Wildlife Conservation Trust of India | title=Asiatic Lion - Population | work=Asiatic Lion Information Centre | publisher=Wildlife Conservation Trust of India | date=2006 | url=http://www.asiaticlion.org/population-gir-forests.htm | accessdate=December 10, 2007}}</ref>
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| − | Until the late [[Pleistocene]] (about 10,000 years ago), the lion was the most widespread land mammal aside from man. They were found in most of Africa, much of Eurasia from western Europe to India and the [[Bering land bridge]], and in the Americas from [[Yukon]] to Peru. Parts of this range were occupied by subspecies that are extinct today.
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| − | ==Population and conservation status==
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| − | Most lions now live in eastern and southern Africa, and their numbers there are rapidly decreasing, with an estimated 30–50% decline over the last two decades.<ref name=IUCN/> Currently, estimates of the African lion population range between 16,500 and 47,000 living in the wild in 2002–2004,<ref>{{cite journal |author=Bauer H, Van Der Merwe S |year= 2002|title=The African lion database |journal=Cat news |volume=36 |issue= |pages=41–53 }}</ref><ref>{{citation |author=Chardonnet P |year=2002|title=Conservation of African lion|publisher =International Foundation for the Conservation of Wildlife|location=Paris, France}}</ref> down from early 1990s estimates that ranged as high as 100,000 and perhaps 400,000 in 1950. The cause of the decline is not well-understood, and may not be reversible.<ref name=IUCN/> Currently, habitat loss and conflicts with humans are considered the most significant threats to the species.<ref name="awf">{{cite web|title=AWF Wildlife: Lion|url=http://www.awf.org/content/wildlife/detail/lion|publisher=African Wildlife Foundation|retrieved=December 10, 2007}}</ref><ref>{{cite web |title=NATURE. The Vanishing Lions |publisher=PBS |url=http://www.pbs.org/wnet/nature/vanishinglions/index.html |accessdate=December 10, 2007 }}</ref> The remaining populations are often geographically isolated from each other, which can lead to [[inbreeding]], and consequently, a lack of [[genetic diversity]]. Therefore the lion is considered a [[vulnerable]] species by the [[International Union for Conservation of Nature and Natural Resources]], while the Asiatic subspecies is [[critically endangered]]. The lion population in the region of West Africa is isolated from lion populations of Central Africa, with little or no exchange of breeding individuals. The number of mature individuals in West Africa is estimated by two separate recent surveys at 850–1,160 (2002/2004). There is disagreement over the size of the largest individual population in West Africa: the estimates range from 100 to 400 lions in [[Burkina Faso]]'s [[Arly-Singou]] ecosystem.<ref name=IUCN/>
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| − | [[Image:Asiatic.lioness.arp.jpg|thumb|left|An [[Asiatic Lion]]ess ''Panthera leo persica'', named MOTI, was born in Helsinki Zoo (Finland) in October 1994; she arrived at Bristol Zoo (England) in January 1996. The [[Gir Forest]] in [[India]] is the natural home of the Asiatic lion but this animal was born in captivity.]]
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| − | Conservation of both African and Asian lions has required the setup and maintenance of national parks and game reserves; among the best known are [[Etosha National Park]] in [[Namibia]], [[Serengeti National Park]] in Tanzania and [[Kruger National Park]] in eastern [[South Africa]]. Outside these areas, the issues arising from lions' interaction with livestock and people usually results in the elimination of the former.<ref>{{cite web | last=Roach | first=John | title=Lions Vs. Farmers: Peace Possible? | work=National Geographic News | publisher=National Geographic | date=July 16, 2003 | url=http://news.nationalgeographic.com/news/2003/07/0716_030716_lions.html | accessdate=December 10, 2007}}</ref> In India, the last refuge of the Asiatic Lion is the 1,412 km² (558 square miles) [[Gir Forest National Park]] in western [[India]] which had about 359 lions (as of April 2006). As in Africa, numerous human habitations are close by with the resultant problems between lions, livestock, locals and wildlife officials.<ref>
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| − | {{cite journal |last= Saberwal|first=Vasant K|coauthors= James P. Gibbs, Ravi Chellam and A. J. T. Johnsingh |year=1994 |month=June |title= Lion-Human Conflict in the Gir Forest, India|journal=Conservation Biology |volume=8 |issue=2 |pages=501–507 |doi=10.1046/j.1523-1739.1994.08020501.x}}</ref> The [[Asiatic Lion Reintroduction Project]] plans to establish a second independent population of [[Asiatic Lions]] at the [[Kuno Wildlife Sanctuary]] in the Indian state of [[Madhya Pradesh]].<ref>{{cite journal |last=Johnsingh |first=A.J.T. |year=2004 |title=WII in the Field: Is Kuno Wildlife Sanctuary ready to play second home to Asiatic lions? |journal=Wildlife Institute of India Newsletter |volume=11 |issue=4 |url=http://www.wii.gov.in/publications/newsletter/winter04/wii%20in%20field.htm |accessdate=December 10, 2007}}</ref> It is important to start a second population to serve as a [[life insurance]] for the last surviving Asiatic Lions and to help develop and maintain [[genetic diversity]] enabling the species to survive.
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| − | The former popularity of the Barbary Lion as a zoo animal has meant that scattered lions in captivity are likely to be descended from Barbary Lion stock. This includes twelve lions at [[Port Lympne Zoo|Port Lympne Wild Animal Park]] in [[Kent]], [[England]] that are descended from animals owned by the [[King of Morocco]].<ref>{{cite web |url=http://www.bigcatrescue.org/barbary_lion_news.htm |title=Barbary Lion News |accessdate=December 10, 2007}}</ref> Another eleven animals believed to be Barbary lions were found in [[Addis Ababa]] zoo, descendants of animals owned by [[Haile Selassie I of Ethiopia|Emperor Haile Selassie]]. WildLink International, in collaboration with [[Oxford University]], launched their ambitious International [[Barbary Lion#The Barbary Lion Project|Barbary Lion Project]] with the aim of identifying and breeding Barbary lions in captivity for eventual reintroduction into a national park in the [[Atlas Mountains]] of [[Morocco]].<ref name="yamaguchi-haddane">{{cite journal |author=Yamaguchi N, Haddane B|year=2002 |title=The North African Barbary lion and the Atlas Lion Project |journal=International Zoo News |volume=49 |issue= |pages=465–81}}</ref>
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| − | Following the discovery of the decline of lion population in Africa, several co-ordinated efforts involving lion [[Wildlife conservation|conservation]] have been organised in an attempt to stem this decline.
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| − | Lions are one species included in the [[Species Survival Plan]], a coordinated attempt by the [[Association of Zoos and Aquariums]] to increase its chances of survival. The plan was originally started in 1982 for the Asiatic Lion, but was suspended when it was found that most North American lions were not [[Genetic pollution|genetically pure]], having been hybridized with African lions. The African lion plan started in 1993, focusing especially on the South African subspecies, although there are difficulties in assessing the genetic diversity of captive lions, since most individuals are of unknown origins, making maintenance of genetic diversity a problem.<ref name="zoos_encyclopedia"/>
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| − | ==Relation with humans==
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| − | ===Man-eaters===
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| − | While lions do not usually hunt people, some (usually males) seem to seek out human prey; well-publicized cases include the [[Tsavo maneaters]], where 28 railway workers were taken by lions over nine months during the construction of a railway bridge over the [[Tsavo River]] in [[Kenya]] in 1898, and the 1991 [[Mfuwe]] man-eater, which killed six people in the Laungwa River Valley in [[Zambia]].<ref>{{cite web | coauthors=Wayne Hosek | title=Man-eaters of the Field Museum: Lion of Mfuwe | work=Field Museum of Natural History | publisher=Field Museum of Natural History | date=2007| url =http://www.fieldmuseum.org/exhibits/exhibit_sites/tsavo/mfuwe.html | accessdate =December 10, 2007}}</ref> In both, the hunters who killed the lions wrote books detailing the animals' predatory behavior. The Mfuwe and Tsavo incidents bear similarities: the lions in both incidents were larger than normal, lacked manes, and seemed to suffer from [[tooth decay]]. The infirmity theory, including tooth decay, is not favored by all researchers. An analysis of teeth and jaws of man-eating lions in museum collections suggests that, while tooth decay may explain some incidents, prey depletion in human-dominated areas is a more likely cause of lion predation on humans.<ref>{{cite journal |last=Patterson |first=Bruce D. |coauthors=Ellis J. Neiburger, Ellis J.; Kasiki, Samuel M. |year=2003 |month=February |title=Tooth Breakage and Dental Disease as Causes of Carnivore-Human Conflicts |journal=Journal of Mammalogy |volume=84 |issue=1 |pages=190–196 |id= |url=http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1644%2F1545-1542(2003)084%3C0190%3ATBADDA%3E2.0.CO%3B2 |accessdate= December 10, 2007}}</ref> In their analysis of Tsavo and man-eating generally, Peterhans and Gnoske acknowledge that sick or injured animals may be more prone to man-eating, but that the behavior is "not unusual, nor necessarily 'aberrant'" where the opportunity exists; if inducements such as access to livestock or human corpses are present, lions will regularly prey upon human beings. The authors note that the relationship is well-attested amongst other pantherines and primates in the paleontological record.<ref>{{cite journal |url=http://www.man-eater.info/gpage6.html |accessdate=December 10, 2007 |last=Peterhans |first=Julian C. Kerbis |coauthors=Thomas Patrick Gnoske |title=The Science of Man-eating | journal = Journal of East African Natural History | issn = 0012-8317 | volume = 90 | issue = 1&2 | year = 2001 | pages = 1-40
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| − | }}</ref>
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| − | [[Image:Tsavo Maneaters Field Museum.jpeg|right|thumb|The [[Tsavo maneaters]] on display in the [[Field Museum of Natural History]]]]
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| − | The lion's proclivity for man-eating has been systematically examined. American and Tanzanian scientists report that man-eating behavior in rural areas of Tanzania increased greatly from 1990 to 2005. At least 563 villagers were attacked and many eaten over this period—a number far exceeding the more famed "Tsavo" incidents of a century earlier. The incidents occurred near [[Selous Game Reserve|Selous National Park]] in [[Rufiji River|Rufiji District]] and in [[Lindi Region|Lindi Province]] near the [[Mozambique|Mozambican]] border. While the expansion of villagers into bush country is one concern, the authors argue that conservation policy must mitigate the danger because, in this case, conservation contributes directly to human deaths. Cases in Lindi have been documented where lions seize humans from the center of substantial villages.<ref name="Packer05">{{cite journal |last=Packer |first=C. |coauthors=Ikanda, D.; Kissui, B.; Kushnir, H. |year=2005 |month=August |title=Conservation biology: lion attacks on humans in Tanzania |journal=[[Nature (journal)|Nature]] |volume=436 |issue=7053 |pages=927–928 |doi=10.1038/436927a| pmid=16107828}}</ref>
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| − | Author Robert R. Frump wrote in ''The Man-eaters of Eden'' that Mozambican refugees regularly crossing Kruger National Park at night in [[South Africa]] are attacked and eaten by the lions; park officials have conceded that man-eating is a problem there. Frump believes thousands may have been killed in the decades after [[apartheid]] sealed the park and forced the refugees to cross the park at night. Mozambicans had for nearly a century before the border was sealed regularly walked across the park in daytime with little harm.<ref>{{cite book |last=Frump |first=RR |title=The Man-Eaters of Eden: Life and Death in Kruger National Park |year=2006 |publisher=The Lyons Press |location= |isbn=1-592288-92-9}}</ref>
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| − | Packer estimates more than 200 Tanzanians are killed each year by lions, [[crocodile]]s, elephants, hippos and [[snake]]s, and that the numbers could be double that amount, with lions thought to kill at least 70 of those. Packer and Ikanda are among the few [[conservation movement|conservationists]] who believe western conservation efforts must take account of these matters not just because of ethical concerns about human life, but also for the long term success of conservation efforts and lion preservation.<ref name="Packer05"/>
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| − | A man-eating lion was killed by game scouts in Southern Tanzania in April 2004. It is believed to have killed and eaten at least 35 people in a series of incidents covering several villages in the Rufiji Delta coastal region.<ref>{{cite web |title=Toothache 'made lion eat humans' |author=Daniel Dickinson |date=October 19, 2004 |publisher=BBC News |url=http://news.bbc.co.uk/1/hi/world/africa/3756180.stm |accessdate=December 10, 2007 }}</ref><ref>{{cite web |title=Terror from man-eating lions increasing in Tanzania |author=Ludger Kasumuni |date=August 28, 2006 |publisher=IPPmedia.com |url=http://www.ippmedia.com/ipp/guardian/2006/08/28/73305.html |accessdate=December 10, 2007 }}</ref> Dr Rolf D. Baldus, the GTZ wildlife program coordinator, commented that it was likely that the lion preyed on humans because it had a large [[abscess]] underneath a [[molar (tooth)|molar]] which was cracked in several places. He further commented that "This lion probably experienced a lot of pain, particularly when it was chewing."<ref>{{cite journal |last=Baldus |first=R |year=2006|month=march |title=A man-eating lion ''(Panthera leo)'' from Tanzania with a toothache |journal=European Journal of Wildlife Research |volume=52 |issue= 1 |pages= 59–62|doi=10.1007/s10344-005-0008-0}}</ref> GTZ is the German development cooperation agency and has been working with the Tanzanian government on wildlife conservation for nearly two decades. Like in other cases this lion was large, lacked a mane, and had a tooth problem.
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| − | The "All-Africa" record of man-eating generally is considered to be not Tsavo, but the lesser-known incidents in the late 1930s through the late 1940s in what was then [[Tanganyika]] (now Tanzania). George Rushby, game warden and professional hunter, eventually dispatched the pride, which over three generations is thought to have killed and eaten 1,500 to 2,000 in what is now [[Njombe]] district.<ref>{{cite book |author=Rushby GG |title= No More the Tusker |year= 1965 |publisher=W. H. Allen |location=London}}</ref>
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| − | ===In captivity===
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| − | Widely seen in captivity,<ref>{{cite web | title=Givskud Zoo Lion Park | url=http://www.goodzoos.com/Denmark/Givskud.htm|accessdate=December 10, 2007}}</ref> lions are part of a group of exotic animals which have formed the core of [[zoo]] exhibits the world over since the late 18th century; members of this group are invariably large vertebrates and include elephants, rhinoceroses, hippopotamuses and larger primates as well as other big cats; zoos sought to gather as many of these species as possible.<ref name ="dc81">de Courcy, p. 81</ref> Though many modern zoos are more selective about their exhibits,<ref>de Courcy, p. 82</ref> there are over 1000 African and 100 Asiatic lions in zoos and wildlife parks around the world. They are considered an ambassador species and are kept for tourism, education and conservation purposes.<ref name ="WAZA">{{cite web | author=Dollinger P, Geser S | title=Animals: WAZA'S virtual zoo - lion | work=WAZA'S virtual zoo| publisher=WAZA (World Association of Zoos and Aquariums) | url=http://www.waza.org/virtualzoo/factsheet.php?id=112-007-002-001&view=Cats |accessdate=December 10, 2007}}</ref> Lions can reach an age of over 20 years in captivity; Apollo, a resident lion of [[Honolulu Zoo]] in [[Honolulu, Hawaii|Honolulu]], [[Hawaii]], died at age 22 in August 2007. His two sisters, born in 1986, are still living.<ref>{{cite journal |last=Aguiar |first=Eloise|year=2007 |month=August |title=Honolulu zoo's old lion roars no more |journal=Honolulu Advertiser|url=http://the.honoluluadvertiser.com/article/2007/Aug/08/ln/hawaii708080394.html |accessdate= December 10, 2007}}</ref> A zoo-based lion breeding program usually involves the consideration of several matters, such as the separation of the various lion subspecies, while, at the same time, attempting to prevent the often negative effects of [[inbred|inbreeding]], which can often occur when lions are divided in this manner.<ref>[http://www.catsurvivaltrust.org/lion.htm Captive Breeding and Lions in Captivity]. Retrieved December 10, 2007.</ref>
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| − | Lions were kept and bred by Assyrian kings as early as 850 B.C.E.,<ref name="Schaller5"/> and [[Alexander the Great]] was said to have been presented with tame lions by the [[Malhi]] of northern India.<ref>{{cite book |author=Smith, Vincent Arthur |authorlink= |title=The Early History of India |year=1924 |pages=97|publisher=Clarendon Press |location=Oxford |isbn= }}</ref> Later in [[Roman era|Roman times]], lions were kept by emperors to take part in the gladiator arenas. Roman notables, including [[Lucius Cornelius Sulla|Sulla]], [[Pompey]], and [[Julius Caesar]], often ordered the mass slaughter of hundreds of lions at a time.<ref>Thomas Wiedemann, ''Emperors and Gladiators'', Routledge, 1995, p. 60. ISBN 0415121647.</ref> In the East, Lions were tamed by Indian princes, and [[Marco Polo]] reported that [[Kublai Khan]] kept lions inside.<ref>Baratay & Hardouin-Fugier, p. 17.</ref> The first European "zoos" spread amongst noble and royal families in the 13th century, and until the 17th century were called [[seraglio]]s; at that time, they came to be called [[menagerie]]s, an extension of the [[cabinet of curiosities]]. They spread from France and Italy during the [[Renaissance]] to the rest of Europe.<ref>Baratay & Hardouin-Fugier, pp. 19–21, 42.</ref> In England, although the seraglio tradition was less developed, Lions were [[Tower of London#Menagerie|kept at the Tower of London]] in a seraglio established by [[John I of England|King John]] in the 13th century,<ref>Baratay & Hardouin-Fugier, p. 20.</ref><ref>{{cite web | last=Owen | first=James | title=Medieval Lion Skulls Reveal Secrets of Tower of London "Zoo" | work=National Geographic Magazine | publisher=National Geographic | date=November 3, 2005 | url=http://news.nationalgeographic.com/news/2005/11/1103_051103_tower_lions.html | accessdate=December 10, 2007}}</ref> probably stocked with animals from an earlier menagerie started in 1125 by [[Henry I of England|Henry I]] at his palace in [[Woodstock, Oxfordshire|Woodstock]], near [[Oxford]]; where lions had been reported stocked by [[William of Malmesbury]].<ref name ="Blunt15">Blunt, p. 15</ref>
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| − | [[Image:Durer lions (sketch).jpg|thumb|[[Albrecht Dürer]], Lions sketch. Circa 1520.]]
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| − | Seraglios served as expressions of the nobility's power and wealth. Animals like big cats and [[elephant]]s, in particular, symbolized power, and would be pitted in fights against each other or domesticated animals. By extension, menageries and seraglios served as demonstrations of the dominance of man over nature. Consequently, the defeat of such natural "lords" by a cow in 1682 astonished the spectators, and the flight of an elephant before a rhinoceros drew jeers. Such fights would slowly fade out in the 17th century with the spread of the menagerie and their appropriation by the commoners. The tradition of keeping big cats as pets would last into the 19th century, at which time it was seen as highly eccentric.<ref>Baratay & Hardouin-Fugier, pp. 24–28.</ref>
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| − | The presence of lions at the Tower of London was intermittent, being restocked when a monarch or his consort, such as [[Margaret of Anjou]] the wife of [[Henry VI of England|Henry VI]], either sought or were given animals. Records indicate they were kept in poor conditions there in the 17th century, in contrast to more open conditions in [[Florence]] at the time.<ref name ="Blunt16">Blunt, p. 16</ref> The menagerie was open to the public by the 18th century; admission was a sum of three half-pence or the supply of a cat or dog for feeding to the lions.<ref name ="Blunt17">Blunt, p. 17</ref> A rival menagerie at the [[Exeter Exchange]] also exhibited lions until the early 19th century.<ref>de Courcy, p. 8–9</ref> The Tower menagerie was closed down by [[William IV of the United Kingdom|William IV]],<ref name ="Blunt17"/> and animals transferred to the [[London Zoo]] which opened its gates to the public on 27 April 1828.<ref name ="Blunt32">Blunt, p. 32</ref>
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| − | {{quotebox|align=right|quote=Animal species disappear when they cannot peacefully orbit the center of gravity that is man.|source=Pierre-Amédée Pichot, 1891<ref>Son of anglophile Amédée Pichot (Baratay & Hardouin-Fugier, p. 114.)</ref>|width=240px}}
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| − | The wild animals trade flourished alongside improved colonial trade of the 19th century. Lions were considered fairly common and inexpensive. Although they would barter higher than tigers, they were less costly than larger, or more difficult to transport animals like the giraffe and hippopotamus, and much less than [[panda]]s.<ref>Baratay & Hardouin-Fugier, p. 122.</ref> Like other animals, lions were seen as little more than a natural, boundless commodity that was mercilessly exploited with terrible losses in capture and transportation.<ref>Baratay & Hardouin-Fugier, pp. 114, 117.</ref> The widely reproduced imagery of the heroic hunter chasing lions would dominate a large part of the century.<ref>Baratay & Hardouin-Fugier, p. 113.</ref> Explorers and hunters exploited a popular [[manicheism|manichean]] division of animals into "good" and "evil" to add thrilling value to their adventures, casting themselves as heroic figures. This resulted in big cats, always suspected of being man-eaters, representing "both the fear of nature and the satisfaction of having overcome it."<ref>Baratay & Hardouin-Fugier, pp. 173, 180–183.</ref>
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| − | [[Image:Lion - melbourne zoo.jpg|thumb|left|Lion at [[Melbourne Zoo]] enjoying an elevated grassy area with some tree shelter]]
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| − | Lions were kept in cramped and squalid conditions at London Zoo until a larger lion house with roomier cages was built in the 1870s.<ref name ="Blunt208">Blunt, p. 208</ref> Further changes took place in the early 20th century, when [[Carl Hagenbeck]] designed enclosures more closely resembling a natural habitat, with concrete 'rocks', more open space and a moat instead of bars. He designed lion enclosures for both [[Melbourne Zoo]] and Sydney's [[Taronga Zoo]], among others, in the early 20th century. Though his designs were popular, the old bars and cage enclosures prevailed until the 1960s in many zoos.<ref name ="dc69">de Courcy, p. 69</ref> In the later decades of the 20th century, larger, more natural enclosures and the use of [[wire mesh]] or [[laminated glass]] instead of lowered dens allowed visitors to come closer than ever to the animals, with some attractions even placing the den on ground higher than visitors, such as the Cat Forest/Lion Overlook of [[Oklahoma City Zoological Park]].<ref name="zoos_encyclopedia"/> Lions are now housed in much larger naturalistic areas; modern recommended guidelines more closely approximate conditions in the wild with closer attention to the lions' needs, highlighting the need for dens in separate areas, elevated positions in both sun and shade where lions can sit and adequate ground cover and drainage as well as sufficient space to roam.<ref name ="WAZA"/>
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| − | ===Baiting and taming===
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| − | [[Image:Circus Lion Tamer.jpg|right|thumb|19th century Etching of a lion tamer in a cage of lions.]]
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| − | {{main|Lion-baiting|Lion taming}}
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| − | Lion-baiting is a [[blood sport]] involving the [[Bait (dogs)|baiting]] of lions in combat with other animals, usually dogs. Records of it exist in ancient times through until the 17th century. It was finally banned in [[Vienna]] by 1800 and England in 1825.<ref>{{cite book |last=Hone |first= William |editor= Kyle Grimes|title= The Every-Day Book |origyear=1825–26 |url=http://www.uab.edu/english/hone/etexts/edb/day-pages/207-july26.html |accessdate=December 10, 2007 |year=2004 |publisher=University of Alabama at Birmingham |pages=26 |chapter=July |chapterurl=http://www.uab.edu/english/hone/etexts/edb/indices/index.html#jul}}</ref><ref>{{cite journal |last=Blaisdell |first=Warren H. |year=1997 |month=November |title= How A Lion Fight Caused England To Stop The Breeding Of Both Ring And Pit Bulldogs|journal=American Bulldog Review |volume=3 |issue=4 |url=http://www.american-bulldog.com/how_a_lion.htm |accessdate= December 10, 2007}}</ref>
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| − | Lion taming refers to the practice of taming lions for entertainment, either as part of an established [[circus]] or as an individual act, such as [[Siegfried & Roy]]. The term is also often used for the taming and display of other big cats such as [[tigers]], [[leopards]] and [[cougars]]. The practice was pioneered in the first half of the 19th century by Frenchman [[Henri Martin (tamer)|Henri Martin]] and American [[Isaac Van Amburgh]] who both toured widely, and whose techniques were copied by a number of followers.<ref name="baratay187">Baratay & Hardouin-Fugier, p. 187.</ref> Van Amburgh performed before Queen [[Victoria of the United Kingdom]] in 1838 when he toured [[Great Britain]]. Martin composed a [[pantomime]] titled ''Les Lions de Mysore'' ("the lions of Mysore"), an idea that Amburgh quickly borrowed. These acts eclipsed [[equestrianism]] acts as the central display of circus shows, but truly entered public consciousness in the early 20th century with cinema. In demonstrating the superiority of man over animal, lion taming served a purpose similar to animal fights of previous centuries.<ref name="baratay187">Baratay & Hardouin-Fugier, p. 187.</ref> The now iconic lion tamer's chair was possibly first used by American [[Clyde Beatty]] (1903–1965).<ref>{{cite book | first=David | last=Feldman | authorlink=David Feldman (author) | coauthors= | year=1993 | title=How Does Aspirin Find a Headache? | edition= | publisher=HarperCollins | location= | isbn= 0-06-016923-0 }}</ref>
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| − | ===Cultural depictions===
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| − | {{details|Cultural depictions of lions}}
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| − | [[Image:Nineve.jpg|thumb|upright|Bronze Lion from [[Nineveh]].]]
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| − | [[Image:GD-EG-KomOmbo016.JPG|thumb|upright|left|The warrior goddess [[Sekhmet]], shown with her sun disk and cobra crown]]
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| − | The Lion has been an icon for humanity for thousands of years, appearing in cultures across Europe, Asia, and Africa. Despite incidents of attacks on humans, lions have enjoyed a positive depiction in culture as strong but noble. A common depiction is their representation as "[[Monarch|king]] of the jungle" or "king of the beasts"; hence, the lion has been a popular symbol of royalty and stateliness,<ref name ="Garai73">{{cite book |last=Garai |first=Jana |title=The Book of Symbols |year=1973 |publisher=Simon & Schuster |location=New York |isbn=671-21773-9}}</ref> as well as a symbol of bravery; it is featured in several [[Aesop's Fables|fables]] of the
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| − | [[6th century B.C.E.|sixth century B.C.E.]] [[Ancient Greece|Greek]] storyteller [[Aesop]].<ref>{{cite book| author=Aesop|coauthors= Gibbs L |title=Aesop's Fables |series=Oxford World's Classics |year=2002 |publisher=Oxford University Press |location=Oxford |isbn=0192840509}}</ref>
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| − | Representations of lions date back 32,000 years; the [[Lion man]] ivory carving from Vogelherd cave in the [[Swabian Alb]] in southwestern [[Germany]] has been determined to be about 32,000 years old from the [[Aurignacian]] culture.<ref name="BurgerJ-Molecular-phylogeny"/> Two lions were depicted mating in the Chamber of Felines in 15,000 year old [[Paleolithic]] [[cave painting]]s in the [[Lascaux]] caves. Cave lions are also depicted in the [[Chauvet Cave]], discovered in 1994; this has been dated at 32,000 years of age,<ref name="Packer00"/> though it may be of similar or younger age to Lascaux.<ref>{{cite conference| first=Christian | last=Züchner | title=Grotte Chauvet Archaeologically Dated | date=September 1998 | location=International Rock Art Congress IRAC ´98 - Vila Real – Portugal | url=http://www.uf.uni-erlangen.de/chauvet/chauvet.html | accessdate =December 10, 2007}}</ref>
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| − | [[Ancient Egypt]] venerated the lion, with the [[Sphinx]] and the lion-headed deity [[Sekhmet]];<ref name ="Garai73"/> [[Maahes]] and [[Dedun]] were Egyptian deities in full lion form.<ref>{{cite web | author=Cass S | title=Maahes | work=Encyclopedia Mythica | publisher=Encyclopedia Mythica | date=1998| url=http://www.pantheon.org/articles/m/maahes.html | accessdate=December 10, 2007}}</ref><ref>{{cite web | author=Lindemans MF | title=Dedun | work=Encyclopedia Mythica | publisher=Encyclopedia Mythica | date=1997 | url=http://www.pantheon.org/articles/d/dedun.html | accessdate=December 10, 2007}}</ref> The [[Nemean Lion]] was symbolic in Ancient Greece and Rome, represented as the constellation and zodiac sign [[Leo (astrology)|Leo]], and described in mythology, where its skin was borne by the hero [[Heracles]].<ref>{{cite book |last=Graves |first=R |authorlink=Robert Graves |title=Greek Myths |year=1955|publisher=Penguin |location=London |isbn=0-14-001026-2|pages=465–469 |chapter=The First Labour:The Nemean Lion}}</ref>
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| − | The lion was a prominent symbol in both the [[Old Babylonian]] and [[Neo-Babylonian Empire]] periods. The classic Babylonian lion motif, found as a statue, carved or painted on walls, is often referred to as the ''striding lion of Babylon''. It is in Babylon that the biblical [[Daniel]] is said to have been delivered from the lion's den.<ref>[[Book of Daniel|Daniel]] 6</ref> Such symbolism was appropriated by Saddam Hussein's regime in Iraq for their [[Lion of Babylon tank]], with the technology adapted from a Russian model.
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| − | <!-- Image with unknown copyright status removed: [[Image:Babylonlion.JPG|thumb|right|upright|Lion of [[Babylon]].]] —>
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| − | [[Narasimha]] ("man-lion") is described as an incarnation ([[avatara]]) of [[Vishnu]] within the [[Puranic]] texts of [[Hinduism]];<ref> [http://srimadbhagavatam.com/1/3/18/en1 Bhag-P 1.3.18] "In the fourteenth incarnation, the Lord appeared as Nrisimha and bifurcated the strong body of the atheist Hiranyakasipu with His nails, just as a carpenter pierces cane." </ref> who takes the form of half-man / half-[[Asiatic Lions|lion]], having a human torso and lower body, but with a lion-like face and claws.<ref> [http://srimadbhagavatam.com/7/8/19-22/en1 Bhag-P 7.8.19–22] </ref> it is worshiped as "Lion God" thus Indian or [[Asiatic Lion]]s which were commonly found throughout most of India in ancient times are considered sacred by all [[Hindus]] in India.
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| − | [[Image:matha.png|right|thumb|180px|"[[Bharat Mata]]" ("Mother India"), [[National personification]] of [[India]], depicted with an Asiatic / [[Asiatic lion|Indian lion]] at her side]]
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| − | [[Singh]] is an [[ancient]] [[India]]n [[Vedic Sanskrit|vedic]] name meaning "Lion" ([[Asiatic Lion]]), dating back over 2000 years to [[ancient India]]. It was originally only used by [[Rajputs]] a [[Hindu]] [[Kshatriya]] or military [[caste]] in India. After the birth of the [[Khalsa]] brotherhood in 1699, the [[Sikh]]s also adopted the name "Singh" due to the wishes of [[Guru Gobind Singh]]. Along with millions of Hindu Rajputs today, it is also used by over 20 million [[Sikhs]] worldwide.<ref>Dr. McCleod, Head of Sikh Studies, Department of South Asian Studies, McMaster University, Hamilton, Ontario, Canada.<!--I thought we only cited published sources?—></ref><ref>Khushwant Singh, ''A History of the Sikhs, Volume I''</ref>
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| − | Found famously on numerous [[Flags]] and [[Coat of Arms]] all across Asia and Europe, the Asiatic Lions also stand firm on the [[National Emblem of India]].<ref>{{cite web | author=Government of India | title=Know India: State Emblem | work=National Portal of India | publisher=National Informatics Centre | date=2005 | url=http://india.gov.in/knowindia/state_emblem.php | accessdate =December 10, 2007}}</ref>.
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| − | Further south on the [[Indian subcontinent]], the Asiatic lion is symbolic for the [[Sinhalese people|Sinhalese]],<ref>{{cite web | author=Government of Sri Lanka | title=Sri Lanka National Flag | publisher=Government of Sri Lanka | date=| url=http://www.gov.lk/info/index.asp?mi=19&xp=52&xi=54&xl=3&o=0&t= | accessdate=December 10, 2007}}</ref> [[Sri Lanka]]'s ethnic majority; the term derived from the Indo-Aryan ''Sinhala'', meaning the "lion people" or "people with lion blood," while a sword wielding lion is the central figure on the national [[flag of Sri Lanka]].<ref>{{cite web | author=Government of Sri Lanka | title=Article 6: The National Flag | work=Official Website of the Government of Sri Lanka | publisher=Government of Sri Lanka | date=| url=http://www.priu.gov.lk/Cons/1978Constitution/Schedle_2_Amd.html | accessdate=December 10, 2007}}</ref>
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| − | The Asiatic Lion is a common motif in [[Chinese art]]. They were first used in art during the late [[Spring and Autumn Period]] (5th or 6th century B.C.E.), and became much more popular during the [[Han Dynasty]] (206 B.C.E.-AD 220), when [[imperial guardian lions]] started to be placed in front of imperial palaces for protection. Because lions have never been native to China, early depictions were somewhat unrealistic; after the introduction of [[Buddhist art]] to China in the [[Tang Dynasty]] (after the 6th century AD), lions were usually depicted without wings, their bodies became thicker and shorter, and their manes became curly.<ref>Li Ling (May 2002). "[http://www.cityu.edu.hk/ccs/Newsletter/newsletter4/Lion/Lion.htm The Two-Way Process in the Age of Globalization]," translated by Ronald Egan. Ex/Change Newsletter from [[City University of Hong Kong]], Issue 4. Retrieved December 10, 2007.</ref> The [[lion dance]] is a form of traditional dance in [[Chinese culture]] in which performers mimic a lion's movements in a lion costume, often with musical accompaniment from cymbals, drums and gongs. They are performed at [[Chinese New Year]], the [[Mid-Autumn Festival|August Moon Festival]] and other celebratory occasions for good luck.<ref>[http://web.mit.edu/lion-dance/www/about/index.html MIT Lion Dance Club - about], Retrieved December 10, 2007.</ref>
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| − | The [[island nation]] of [[Singapore]] ''(Singapura)'' derives its name from the [[Malay language|Malay]] words {{lang|ms|''singa''}} (lion) and {{lang|ms|''pura''}} (city), which in turn is from the [[Tamil language|Tamil]]-[[Sanskrit]] சிங்க ''singa'' {{lang|sa|सिंह}} {{IAST|''siṃha''}} and {{lang|sa|पुर}} புர {{IAST|''pura''}}, which is cognate to the [[Greek language|Greek]] {{lang|el|''πόλις''}}, ''pólis''.<ref> {{cite web | url=http://www.bartleby.com/61/46/S0424600.html|title=Singapore| work=The American Heritage® Dictionary of the English Language: Fourth Edition|year=2000|publisher=bartleby.com|accessdate=December 10, 2007 }}</ref> According to the [[Malay Annals]], this name was given by a 14th century [[Sumatra]]n Malay prince named [[Sang Nila Utama]], who, on alighting the island after a thunderstorm, spotted an auspicious beast on shore that his chief minister identified as a lion (Asiatic Lion).<ref> {{cite web| title=Early History | url=http://www.sg/explore/history.htm | publisher=Ministry of Information, Communications and the Arts, Singapore | accessdate=December 10, 2007 }}</ref> Recent studies of Singapore indicate that lions have never lived there, and the beast seen by Sang Nila Utama was more likely to have been a [[tiger]].
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| − | "[[Aslan]]" or "[[Arslan]] (Ottoman ارسلان ''arslān'' and اصلان ''aṣlān'') is the [[Turkish language|Turkish]] and [[Mongolian language|Mongolian]] word for "[[lion]]." It was used as a title by a number of [[Seljuk Turks|Seljuk]] and [[Ottoman Dynasty|Ottoman]] rulers, including [[Alp Arslan]] and [[Ali Pasha]], and is a [[Turkic languages|Turkic]]/[[Iranian languages|Iranian]] name.
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| − | [[Image:Royal Arms of Scotland.png|thumb|left|Lion rampant on the [[royal coat of arms of Scotland]]]]
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| − | "Lion" was the nickname of medieval warrior rulers with a reputation for bravery, such as [[Richard I of England]], known as Richard the Lionheart,<ref name ="Garai73"/>, [[Henry the Lion]] ([[German language|German]]: ''Heinrich der Löwe''), [[Duke of Saxony]] and [[Robert III of Flanders]] nicknamed "The Lion of Flanders" - a major [[Flemish]] national hero up to the present. Lions were frequently depicted on [[coat of arms|coats of arms]], either as a device on shields themselves, or as [[supporters]]. The formal language of [[heraldry]], called [[blazon]], employs French terms to describe the images precisely. Such descriptions specified whether lions or other creatures were "rampant" or "passant," that is whether they were rearing or crouching.<ref name="Notre Dame">{{cite web | title =Heraldic Dictionary: Beasts | publisher=University of Notre Dame | url =http://www.rarebooks.nd.edu/digital/heraldry/charges/lions.html | accessdate=December 10, 2007 }}</ref> As a heraldic symbol or epithet, the lion continues to be used for modern sporting teams, from national soccer teams such as [[England national football team|England]], [[Scotland national football team|Scotland]] and [[Singapore national football team|Singapore]] to famous clubs such as the [[Detroit Lions]]<ref>{{cite web | author=| title=Official Website of the Detroit Lions | publisher=Detroit Lions | date=2001 | url=http://www.detroitlions.com/index.cfm?homelink=y | accessdate=December 10, 2007}}</ref> of the NFL, [[Chelsea F.C.|Chelsea]]<ref name="new crest">{{cite news | date=2004-11-12 | url=http://news.bbc.co.uk/sport1/hi/football/teams/c/chelsea/4008257.stm | title=Chelsea centenary crest unveiled | publisher=BBC | accessdate=December 10, 2007| author=}}</ref> and [[Aston Villa F.C.|Aston Villa]] of the [[English Premier League]],<ref>{{cite web | author=Aston Villa F.C. | title=The Aston Villa Crest: 2007 Onwards... | publisher=Aston Villa F.C. | date=2007 | url=http://www.avfc.premiumtv.co.uk/page/CrestTest/0,,10265,00.html | accessdate=December 10, 2007}}</ref> (and the Premiership itself) to a [[The Lions|host of smaller clubs]] around the world. Villa sport a Scottish [[Lion Rampant]] on their crest, as do [[Rangers F.C.|Rangers]] and [[Dundee United F.C.|Dundee United]] of the [[Scottish Premier League]].
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| − | [[Image:Una-lion.jpg|thumb|right|225| The lion is a popular symbol and mascot of high schools, colleges and universities throughout the United States. This statue is on the campus of the [[University of North Alabama]].]]Lions continue to feature in modern literature, from the messianic [[Aslan]] in ''[[The Lion, the Witch and the Wardrobe]]'' and following books from the [[Narnia]] series written by [[C.S. Lewis]],<ref>{{cite book |last=Lewis |first=C.S. |authorlink=C. S. Lewis |title= The Lion, the Witch and the Wardrobe |year= 1950 |publisher= HarperCollins |isbn= 0-06-023481-4}}</ref> to the comedic [[Cowardly Lion]] in ''[[The Wonderful Wizard of Oz]]''.<ref>L. Frank Baum, Michael Patrick Hearn, ''The Annotated Wizard of Oz'', p 148, ISBN 0-517-500868 </ref> The advent of moving pictures saw the continued presence of lion symbolism; one of the most iconic and widely recognized lions is [[Leo the Lion (MGM)|Leo the Lion]], the mascot for [[Metro-Goldwyn-Mayer]] (MGM) studios, which has been in use since the 1920s.<ref name="tvacres">{{cite web | url=http://www.tvacres.com/adanimals_leolion.htm | title=TV ACRES: Advertising Mascots - Animals - Leo the MGM Lion (MGM Studios) | work=TV Acres}}</ref> The 1960s saw the appearance of what is possibly the most famous lioness, the Kenyan animal [[Elsa the lioness|Elsa]] in the movie ''[[Born Free]]'',<ref>{{cite book |last=Adamson |first=George |authorlink=George Adamson |title= Bwana Game : the life story of George Adamson|year= 1969 |publisher=Fontana |location= |isbn=0006121454 }}</ref> based on the true-life international bestselling book of the same title.<ref>{{cite book |last=Adamson |first=Joy |authorlink=Joy Adamson |title=Born Free: A Lioness of Two Worlds |year=2000|origyear=1960 |publisher=Pantheon |location= |isbn=0375714383 }}</ref> The lion's role as King of the Beasts has been used in cartoons, from the 1950s manga which gave rise to the first Japanese color TV animation series, ''[[Kimba the White Lion]]'', Leonardo Lion of ''[[King Leonardo and his Short Subjects]]'', both from the 1960s, up to the 1994 [[Walt Disney Pictures|Disney]] animated feature film ''[[The Lion King]]'',<ref>{{cite book |last=Schweizer |first=Peter |title=Disney: The Mouse Betrayed |year=1998 |pages=164-69|publisher=Regnery Publishing |location=Washington D.C. |isbn=0-89526-387-4}}</ref><ref>{{cite web | title=King Leonardo and His Short Subjects | work=Internet Movie Database | publisher=Internet Movie Database Inc. | date=2007 | url=http://www.imdb.com/title/tt0053515/maindetails | accessdate=December 10, 2007}}</ref> which also featured the popular song "[[The Lion Sleeps Tonight]]" in its soundtrack.
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| − | {{clear}}
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| − | ==Notes==
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| − | {{reflist|3}}
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| − | ==References==
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| − | {{refbegin}}
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| − | *{{cite book |last=Baratay |first=Eric |coauthors=Elisabeth Hardouin-Fugier |title=Zoo : a history of zoological gardens in the West |year=2002 |publisher=Reaktion Books |location=London |isbn=1861891113}}
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| − | *{{cite book |last=Blunt |first=Wilfred |title=The Ark in the Park: The Zoo in the Nineteenth Century |year=1975 |publisher=Hamish Hamilton |location=London |isbn=0241893313}}
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| − | *{{cite book |last=de Courcy |first=Catherine |title=The Zoo Story |year=1995 |publisher=Penguin Books |location=Ringwood, Victoria |isbn=0140239197}}
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| − | *{{cite book |last=Schaller |first=George B. |authorlink=George Schaller|year=1972 |title= The Serengeti lion: A study of predator-prey relations|publisher= University of Chicago Press|location=Chicago |isbn=0226736393}}
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| − | {{refend}}
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| − | ==External links==
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| − | All links Retrieved December 10, 2007.
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| − | *[http://www.bigcatcare.org Center for Animal Research and Education] Providing Sanctuary for over 50 big cats
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| − | * [http://www.awf.org/wildlives/148 Lion: Wildlife summary from the African Wildlife Foundation]
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| − | * [http://animaldiversity.ummz.umich.edu/site/accounts/information/Panthera_leo.html Panthera leo (lion) from "Animal Diversity Web"]
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| − | * [http://www.lionresearch.org/ Lion Research Center] – The official website of a research group at the U. of Minnesota that has conducted extensive field research on lions and has published over 50 peer-reviewed scientific articles on lions.
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| − | * [http://www.lionconservationfund.org/ Lion Conservation Fund] – A fund dedicated to the research and conservation of the lion
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| − | * [http://www.pbs.org/wnet/nature/vanishinglions/ Nature Documentary: "The Vanishing Lions"]
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| − | * [http://www.asterpix.com/console?avi=7660461 Hypervideo showing buffalos fending off lion attack]
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| − | [[Category:Life sciences]]
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| − | [[Category:Animals]]
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