Difference between revisions of "Gadidae" - New World Encyclopedia

From New World Encyclopedia
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The subfamily '''Ranicipitinae''' has only one extant member, the tadpole cod (''Raniceps raninus''). It has two dorsal fins with the first dorsal fin having three short rays, one anal fin with a long base, the head without pores or a lateral line, a chin barbel present, and the upper jaw protruding beyond the lower. Most authors place it in Gadidae, although some place it in its own family (Nelson 2006).
 
The subfamily '''Ranicipitinae''' has only one extant member, the tadpole cod (''Raniceps raninus''). It has two dorsal fins with the first dorsal fin having three short rays, one anal fin with a long base, the head without pores or a lateral line, a chin barbel present, and the upper jaw protruding beyond the lower. Most authors place it in Gadidae, although some place it in its own family (Nelson 2006).
 +
 +
ITIS (2004a) recognizes yet another subfamily within Gadidae, along with Gadinae and Lotinae. This is Phycinae, the phycine hakes.
  
 
Members of Gadidae are  generally medium sized fish. Almost all species have [[barbel (anatomy)|barbel]]s on their chin (all Lotinae, almost all Gadidae, and the tadpole cod). This barbel is used while browsing on the sea floor. Gadids are carnivorous, feeding on smaller fish and [[crustacean]]s.
 
Members of Gadidae are  generally medium sized fish. Almost all species have [[barbel (anatomy)|barbel]]s on their chin (all Lotinae, almost all Gadidae, and the tadpole cod). This barbel is used while browsing on the sea floor. Gadids are carnivorous, feeding on smaller fish and [[crustacean]]s.
  
 
Gadids tend to be highly prolific, with many species producing several million eggs at each spawning. This contributes to their high population numbers, which, in turn, makes commercial fishing relatively easy (Cohen 1998).
 
Gadids tend to be highly prolific, with many species producing several million eggs at each spawning. This contributes to their high population numbers, which, in turn, makes commercial fishing relatively easy (Cohen 1998).
 
  
 
==Distribution==
 
==Distribution==
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==Genera and species==
 
==Genera and species==
The following taxonomy follows (Nelson 2006). However, as noted above, many authorities raise Gadinae to the family level, and the cuskfishes (subfamily Lotinae) then are removed from the Gadidae family. While the Ranicipinae subfamily is sometimes made into its own family, generally its one extant member, ''Rancipes raninus'' is considered part of Gadidae whether or not Gadinae is raised to the family level.
+
The following taxonomy follows (Nelson 2006). However, as noted above, many authorities raise Gadinae to the family level, and the cuskfishes (subfamily Lotinae) then are removed from the Gadidae family. While the Ranicipinae subfamily is sometimes made into its own family, generally its one extant member, ''Rancipes raninus'' is considered part of Gadidae whether or not Gadinae is raised to the family level. When Gadinae is raised to the family level, some authoritires recognize several other subfamilies within Gadidae (Nelson 2006).
  
 
Family '''Gadidae'''
 
Family '''Gadidae'''
* Genus ''[[Arctogadus]]'' <small> [[P. A. Dryagin|Dryagin]], 1932. </small>  
+
*'''Subfamily Gadinae'''
** [[East Siberian cod]] (''[[Arctogadus borisovi]]'') <small> [[P. A. Dryagin|Dryagin]], 1932 </small>
+
** Genus ''[[Arctogadus]]'' <small> [[P. A. Dryagin|Dryagin]], 1932. </small>  
** [[Arctic cod]] (''[[Arctogadus glacialis]]'') <small> ([[Wilhelm Peters|Peters]], 1872) </small>
+
*** [[East Siberian cod]] (''[[Arctogadus borisovi]]'') <small> [[P. A. Dryagin|Dryagin]], 1932 </small>
* Genus ''[[Boreogadus]]'' <small> [[Albert Günther|Günther]], 1862. </small>  
+
*** [[Arctic cod]] (''[[Arctogadus glacialis]]'') <small> ([[Wilhelm Peters|Peters]], 1872) </small>
** [[Polar cod]] (''[[Boreogadus saida]]'') <small> ([[Ivan Ivanovich Lepekhin|Lepechin]], 1774) </small>
+
** Genus ''[[Boreogadus]]'' <small> [[Albert Günther|Günther]], 1862. </small>  
* Genus ''[[Eleginus]]'' <small> [[G. Fischer|Fischer]], 1813. </small>  
+
*** [[Polar cod]] (''[[Boreogadus saida]]'') <small> ([[Ivan Ivanovich Lepekhin|Lepechin]], 1774) </small>
** [[Saffron cod]] (''[[Eleginus gracilis]]'') <small> ([[Tilesius]], 1810) </small>
+
** Genus ''[[Eleginus]]'' <small> [[G. Fischer|Fischer]], 1813. </small>  
** [[Navaga]] (''[[Eleginus nawaga]]'') <small> ([[Joseph Gottlieb Kölreuter|Koelreuter]], 1770) </small>
+
*** [[Saffron cod]] (''[[Eleginus gracilis]]'') <small> ([[Tilesius]], 1810) </small>
* Genus ''[[Gadiculus]]'' <small> [[Alphone Guichenot|Guichenot]], 1850. </small>  
+
*** [[Navaga]] (''[[Eleginus nawaga]]'') <small> ([[Joseph Gottlieb Kölreuter|Koelreuter]], 1770) </small>
** [[Silvery cod]] (''[[Gadiculus argenteus]] argentus'') <small> [[Alphone Guichenot|Guichenot]], 1850 </small>   
+
** Genus ''[[Gadiculus]]'' <small> [[Alphone Guichenot|Guichenot]], 1850. </small>  
** ''[[Gadiculus argenteus thori]]'' <small> [[Johannes Schmidt|Schmidt]], 1914 </small>   
+
*** [[Silvery cod]] (''[[Gadiculus argenteus]] argentus'') <small> [[Alphone Guichenot|Guichenot]], 1850 </small>   
* Genus ''[[Cod|Gadus]]'' <small> [[Carl von Linné|Linné]], 1758. </small>  
+
*** ''[[Gadiculus argenteus thori]]'' <small> [[Johannes Schmidt|Schmidt]], 1914 </small>   
** [[Pacific cod]] (''[[Gadus macrocephalus]]'') <small> [[Wilhelm Gottlieb Tilesius|Tilesius]], 1810 </small>
+
** Genus ''[[Cod|Gadus]]'' <small> [[Carl von Linné|Linné]], 1758. </small>  
** [[Atlantic cod]] (''[[Gadus morhua]]'') <small> [[Linnaeus]], 1758 </small>
+
*** [[Pacific cod]] (''[[Gadus macrocephalus]]'') <small> [[Wilhelm Gottlieb Tilesius|Tilesius]], 1810 </small>
** [[Greenland cod]] (''[[Gadus ogac]]'') <small> [[John Richardson|Richardson]], 1836 </small>
+
*** [[Atlantic cod]] (''[[Gadus morhua]]'') <small> [[Linnaeus]], 1758 </small>
* Genus ''[[Melanogrammus]]'' <small> [[Theodore Nicholas Gill|Gill]], 1862. </small>
+
*** [[Greenland cod]] (''[[Gadus ogac]]'') <small> [[John Richardson|Richardson]], 1836 </small>
** [[Haddock]] (''[[Melanogrammus aeglefinus]]'') <small> ([[Linnaeus]], 1758) </small>
+
** Genus ''[[Melanogrammus]]'' <small> [[Theodore Nicholas Gill|Gill]], 1862. </small>
* Genus ''[[Merlangius]]'' <small> [[F. A. P. de Garsault|Garsault]], 1764. </small>  
+
*** [[Haddock]] (''[[Melanogrammus aeglefinus]]'') <small> ([[Linnaeus]], 1758) </small>
** [[Whiting (fish)|Whiting]] (''[[Merlangius merlangus]]'') <small> ([[Linnaeus]], 1758) </small>
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** Genus ''[[Merlangius]]'' <small> [[F. A. P. de Garsault|Garsault]], 1764. </small>  
* Genus ''[[Microgadus]]'' <small> [[Theodore Nicholas Gill|Gill]], 1865. </small>  
+
*** [[Whiting (fish)|Whiting]] (''[[Merlangius merlangus]]'') <small> ([[Linnaeus]], 1758) </small>
** [[Pacific tomcod]] (''[[Microgadus proximus]]'') <small> ([[Charles Frédéric Girard|Girard]], 1854) </small>  
+
** Genus ''[[Microgadus]]'' <small> [[Theodore Nicholas Gill|Gill]], 1865. </small>  
** [[Atlantic tomcod]] (''[[Microgadus tomcod]]'') <small> ([[Johann Julius Walbaum|Walbaum]], 1792) </small>
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*** [[Pacific tomcod]] (''[[Microgadus proximus]]'') <small> ([[Charles Frédéric Girard|Girard]], 1854) </small>  
* Genus ''[[Micromesistius]]'' <small> [[Theodore Nicholas Gill|Gill]], 1863. </small>  
+
*** [[Atlantic tomcod]] (''[[Microgadus tomcod]]'') <small> ([[Johann Julius Walbaum|Walbaum]], 1792) </small>
 +
** Genus ''[[Micromesistius]]'' <small> [[Theodore Nicholas Gill|Gill]], 1863. </small>  
 
** [[Southern blue whiting]] (''[[Micromesistius australis]]'') <small> [[John Roxbrough Norman|Norman]], 1937 </small>  
 
** [[Southern blue whiting]] (''[[Micromesistius australis]]'') <small> [[John Roxbrough Norman|Norman]], 1937 </small>  
** [[Blue whiting]] (''[[Micromesistius poutassou]]'') <small> ([[Antoine Risso|Risso]], 1827) </small>  
+
*** [[Blue whiting]] (''[[Micromesistius poutassou]]'') <small> ([[Antoine Risso|Risso]], 1827) </small>  
* Genus ''[[Pollachius]]'' <small> [[Sven Nilsson|Nilsson]], 1832. </small>  
+
** Genus ''[[Pollachius]]'' <small> [[Sven Nilsson|Nilsson]], 1832. </small>  
** [[Atlantic pollock]] (''[[Pollachius pollachius]]'') <small> ([[Linnaeus]], 1758) </small>  
+
*** [[Atlantic pollock]] (''[[Pollachius pollachius]]'') <small> ([[Linnaeus]], 1758) </small>  
** [[Saithe]] (''[[Pollachius virens]]'') <small> ([[Linnaeus]], 1758) </small>
+
*** [[Saithe]] (''[[Pollachius virens]]'') <small> ([[Linnaeus]], 1758) </small>
* Genus ''[[Raniceps]]'' <small> [[Lorenz Oken|Oken]], 1817. </small>
+
** Genus ''[[Theragra]]'' <small> [[Frederic Augustus Lucas|Lucas]] in [[David Starr Jordan|Jordan]] & [[Barton Warren Evermann|Evermann]], 1898. </small>  
** [[Tadpole fish]] (''[[Raniceps raninus]]'') <small> ([[Linnaeus]], 1758) </small>
+
*** [[Alaska pollock]] (''[[Theragra chalcogramma]]'') <small> ([[Peter Simon Pallas|Pallas]], 1814) </small>  
* Genus ''[[Theragra]]'' <small> [[Frederic Augustus Lucas|Lucas]] in [[David Starr Jordan|Jordan]] & [[Barton Warren Evermann|Evermann]], 1898. </small>  
+
*** [[Norwegian pollock]] (''[[Theragra finnmarchica]]'') <small> [[Einar Koefoed|Koefoed]], 1956 </small>
** [[Alaska pollock]] (''[[Theragra chalcogramma]]'') <small> ([[Peter Simon Pallas|Pallas]], 1814) </small>  
+
** Genus ''[[Trisopterus]]'' <small> [[Constantine Samuel Rafinesque|Rafinesque]], 1814. </small>  
** [[Norwegian pollock]] (''[[Theragra finnmarchica]]'') <small> [[Einar Koefoed|Koefoed]], 1956 </small>
+
*** [[Norway pout]] (''[[Trisopterus esmarkii]]'') <small> ([[Sven Nilsson|Nilsson]], 1855) </small>
* Genus ''[[Trisopterus]]'' <small> [[Constantine Samuel Rafinesque|Rafinesque]], 1814. </small>  
+
*** [[Trisopterus luscus|Pouting]] (''[[Trisopterus luscus]]'') <small> ([[Linnaeus]], 1758) </small>  
** [[Norway pout]] (''[[Trisopterus esmarkii]]'') <small> ([[Sven Nilsson|Nilsson]], 1855) </small>
+
*** [[Poor cod]] (''[[Trisopterus minutus]]'') <small> ([[Linnaeus]], 1758) </small>
** [[Trisopterus luscus|Pouting]] (''[[Trisopterus luscus]]'') <small> ([[Linnaeus]], 1758) </small>  
+
*'''Subfamily Ranicipitinae'''
** [[Poor cod]] (''[[Trisopterus minutus]]'') <small> ([[Linnaeus]], 1758) </small>
+
** Genus ''[[Raniceps]]'' <small> [[Lorenz Oken|Oken]], 1817. </small>
 +
*** [[Tadpole fish]] or tadpole cod (''[[Raniceps raninus]]'') <small> ([[Linnaeus]], 1758) </small>
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*'''Subfamily '''Lotinae''' (cuskfishes)
 +
** Genus ''Lota'' <small> [[Lorenz Oken|Oken]], 1817. </small>
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***[[Burbot]] (''Lota lota'') <small> ([[Linnaeus]], 1758) </small>
 +
**Genus ''Brosme'' <small> [[Lorenz Oken|Oken]], 1817. </small>
 +
*** [[Brosme]] (cusk) (''Brosme brosme'') <small> [[Ascanius]], 1772. </small>
 +
** Genus ''Molva'' (three species) <small> [[Lesueur]], 1819. </small>
 +
*** Blue ling (''Molva dypterygia'') <small> Pennant, 1784) </small>
 +
*** Mediterranean ling (''Molva macrophthalma'' <small>(Rafinesque, 1810) </small>
 +
*** European ling, drizzie (''Molva molva'' <small> ([[Linnaeus]], 1758) </small>
  
 
== References ==
 
== References ==
{{reflist}}
+
 
 
*{{FishBase family|family=Gadidae|year=2005|month=December}}
 
*{{FishBase family|family=Gadidae|year=2005|month=December}}
 
*''FishBase'': [http://www.fishbase.org/identification/specieslist.cfm?famcode=183&areacode= Gadidae species photo gallery]  
 
*''FishBase'': [http://www.fishbase.org/identification/specieslist.cfm?famcode=183&areacode= Gadidae species photo gallery]  
  
 
<ref name=EoF>{{cite book |editor=Paxton, J.R. & Eschmeyer, W.N.|author= Cohen, Daniel M.|year=1998|title=Encyclopedia of Fishes|publisher= Academic Press|location=San Diego|pages= 130-131|isbn= 0-12-547665-5}}</ref>
 
<ref name=EoF>{{cite book |editor=Paxton, J.R. & Eschmeyer, W.N.|author= Cohen, Daniel M.|year=1998|title=Encyclopedia of Fishes|publisher= Academic Press|location=San Diego|pages= 130-131|isbn= 0-12-547665-5}}</ref>
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 +
ITIS 2004a
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http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=164701
 +
Gadidae  Rafinesque, 1810
 +
Taxonomic Serial No.: 164701
 +
 +
ITIS 2004b
 +
http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=550570
 +
Lotinae 
 +
Taxonomic Serial No.: 550570
 +
  
  

Revision as of 02:41, 26 December 2008

Gadidae
Fossil range: Oligocene–Recent
Atlantic cod, Gadus morhua
Atlantic cod, Gadus morhua
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Gadiformes
Family: Gadidae
Genera

See text

Gadidae is a family of almost exclusively marine fish in the order Gadiformes, characterized by the first dorsal fin being posterior to the head, the lack of true spines in the fins, and most with long dorsal and anal fins. Known as the "cod family," Gadidae includes cods, haddocks, whiting, pollock, and, in some taxonomic schemes, the cuskfishes. The familiar cods and haddocks are characterized by three dorsal fins and two anal fins. Members of Gadidae are found in the Arctic, Atlantic, and Pacific Oceans, with one of the cuskfishes a freshwater species.


Overview and description

Gadiformes, the order in class Actinopterygii to which Gadidae belongs, includes many fish of commercial importance, constituting over one quarter of the marine fish catch of the world (Nelson 2006). Members of Gadiformes are characterized by the pelvic fins (when presence) havin up to 11 rays and being below or in front of the pectoral fins, generally long dorsal and anal fins, no true spines in the fins, usually cycloid (rarely ctenoid) scales, and a swim bladder without pneumatic duct (Nelson 2006).

Gadidae, the cod family, has about 16 genera and 31 species according to the classification used by Nelson (2006). According to his system, members of Gadidae are characterized by having the first dorsal fin posterior to the head, the head of the vomer toothed, and the swim bladder not connected with auditory capsules. Nelson places the members of Gadiformes into three subfamilies: Lotinae (cuskfishes), Gadinae (cods and haddocks), and Ranicipitinae (tadpole cods). However, different classification schemes are recognized, including those that elevate the first two subfamilies to the family level, and even some that elevate the third subfamily to family status.

Members of Gadinae subfamily, which some elevate to family status (as Gadidae), is characterized by having three dorsal and two anal fins, an egg without an oil globule, usually a chin barbel, and a slightly forked or truncate caudal fin (Nelson 2006). This subfamily includes the cods, haddocks, pollocks, and the whiting (Merlangius merlangus). Twelve genera are included in this subfamily, with about 25 species. Note that P. pollachius, one of the two species known as pollocks (or pollacks) lacks a chin barbel.

Members of the Lotinae subfamily, which some also elevate to family status, is characterized by having one or two dorsal fins and one anal fin. Members of this subfamily always have a chin barbel, but there are not barbels on the snout. The caudal fin is rounded and the egg has an oil globule. Known as the cuskfishes, there are five species in the taxon, which are arranged into three genera.

The subfamily Ranicipitinae has only one extant member, the tadpole cod (Raniceps raninus). It has two dorsal fins with the first dorsal fin having three short rays, one anal fin with a long base, the head without pores or a lateral line, a chin barbel present, and the upper jaw protruding beyond the lower. Most authors place it in Gadidae, although some place it in its own family (Nelson 2006).

ITIS (2004a) recognizes yet another subfamily within Gadidae, along with Gadinae and Lotinae. This is Phycinae, the phycine hakes.

Members of Gadidae are generally medium sized fish. Almost all species have barbels on their chin (all Lotinae, almost all Gadidae, and the tadpole cod). This barbel is used while browsing on the sea floor. Gadids are carnivorous, feeding on smaller fish and crustaceans.

Gadids tend to be highly prolific, with many species producing several million eggs at each spawning. This contributes to their high population numbers, which, in turn, makes commercial fishing relatively easy (Cohen 1998).

Distribution

Only one species, Lota lota, the burbot, is a completely freshwater member of the family and order. It is found in northern parts of North America and Eurasia. The burbot is part of the subfamily Lotinae (cuskfishes). Other members of Lotinae are marine and found in the northern Atlantic.

Members of the Gadinae subfamily (which a number of authorities place at the family level, with the cuskfishes in their own family) are marine and found in the Arctic, Atlantic, and Pacific Oceans. There is one member, Microgadus tomcod, that is marine (off the Atlantic coast of North America) but can spawn in freshwater, and some freshwater populations are known (Nelson 2006).

The tadpole cod, Ranicpes raninux, in subfamily Ranicipitinae, is found in shallow coastal waters of the northeatern Atlantic, from the Norwegian coast to the the Bay of Biscay and the British Isles (Nelson 2006).

Genera and species

The following taxonomy follows (Nelson 2006). However, as noted above, many authorities raise Gadinae to the family level, and the cuskfishes (subfamily Lotinae) then are removed from the Gadidae family. While the Ranicipinae subfamily is sometimes made into its own family, generally its one extant member, Rancipes raninus is considered part of Gadidae whether or not Gadinae is raised to the family level. When Gadinae is raised to the family level, some authoritires recognize several other subfamilies within Gadidae (Nelson 2006).

Family Gadidae

  • Subfamily Gadinae
    • Genus Arctogadus Dryagin, 1932.
      • East Siberian cod (Arctogadus borisovi) Dryagin, 1932
      • Arctic cod (Arctogadus glacialis) (Peters, 1872)
    • Genus Boreogadus Günther, 1862.
      • Polar cod (Boreogadus saida) (Lepechin, 1774)
    • Genus Eleginus Fischer, 1813.
      • Saffron cod (Eleginus gracilis) (Tilesius, 1810)
      • Navaga (Eleginus nawaga) (Koelreuter, 1770)
    • Genus Gadiculus Guichenot, 1850.
      • Silvery cod (Gadiculus argenteus argentus) Guichenot, 1850
      • Gadiculus argenteus thori Schmidt, 1914
    • Genus Gadus Linné, 1758.
      • Pacific cod (Gadus macrocephalus) Tilesius, 1810
      • Atlantic cod (Gadus morhua) Linnaeus, 1758
      • Greenland cod (Gadus ogac) Richardson, 1836
    • Genus Melanogrammus Gill, 1862.
    • Genus Merlangius Garsault, 1764.
      • Whiting (Merlangius merlangus) (Linnaeus, 1758)
    • Genus Microgadus Gill, 1865.
      • Pacific tomcod (Microgadus proximus) (Girard, 1854)
      • Atlantic tomcod (Microgadus tomcod) (Walbaum, 1792)
    • Genus Micromesistius Gill, 1863.
    • Southern blue whiting (Micromesistius australis) Norman, 1937
      • Blue whiting (Micromesistius poutassou) (Risso, 1827)
    • Genus Pollachius Nilsson, 1832.
      • Atlantic pollock (Pollachius pollachius) (Linnaeus, 1758)
      • Saithe (Pollachius virens) (Linnaeus, 1758)
    • Genus Theragra Lucas in Jordan & Evermann, 1898.
      • Alaska pollock (Theragra chalcogramma) (Pallas, 1814)
      • Norwegian pollock (Theragra finnmarchica) Koefoed, 1956
    • Genus Trisopterus Rafinesque, 1814.
      • Norway pout (Trisopterus esmarkii) (Nilsson, 1855)
      • Pouting (Trisopterus luscus) (Linnaeus, 1758)
      • Poor cod (Trisopterus minutus) (Linnaeus, 1758)
  • Subfamily Ranicipitinae
    • Genus Raniceps Oken, 1817.
      • Tadpole fish or tadpole cod (Raniceps raninus) (Linnaeus, 1758)
  • Subfamily Lotinae (cuskfishes)
    • Genus Lota Oken, 1817.
    • Genus Brosme Oken, 1817.
      • Brosme (cusk) (Brosme brosme) Ascanius, 1772.
    • Genus Molva (three species) Lesueur, 1819.
      • Blue ling (Molva dypterygia) Pennant, 1784)
      • Mediterranean ling (Molva macrophthalma (Rafinesque, 1810)
      • European ling, drizzie (Molva molva (Linnaeus, 1758)

References
ISBN links support NWE through referral fees

[1]

ITIS 2004a http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=164701 Gadidae Rafinesque, 1810 Taxonomic Serial No.: 164701

ITIS 2004b http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=550570 Lotinae Taxonomic Serial No.: 550570

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  1. Cohen, Daniel M. (1998). in Paxton, J.R. & Eschmeyer, W.N.: Encyclopedia of Fishes. San Diego: Academic Press, 130-131. ISBN 0-12-547665-5.