Difference between revisions of "Esox" - New World Encyclopedia

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== Overview and description ==
 
== Overview and description ==
The ''Esox'' genus is the only living genus in the pike family, Esocidae, in the order Esociformes of the class Actinopterygii.  Members of the Esociformes, which also include the mudminnows (family Umbridae), are characterized by posteriorly located dorsal and anal fins, the lack of an adipose fin, toothless maxilla but in the gape of the mouth, and no pyloric caeca. Members of the family Esocidae, the esocids, are characterized by a forked caudal fin with 40 to 50 rays, a complete lateral line, 10 to 20 branchiostegal rays, nasals present, snout produced, and 43 to 67 vertebrae. The other family of Esociformes, the Umbridae, differs in that the mudminnows have a rounded caudal fin with 20 to 30 rays, a lateral line that is faint or absent, nasals absent, snout not produced, only 32 to 42 vertebrae, and 5 to 8 branchiostegal rays (Nelson 2006).
+
The ''Esox'' genus is the only living genus in the pike family, Esocidae, in the order Esociformes of the class Actinopterygii.   
 
[[File:Muskellunge USFWS.jpg|thumb|320px|right|Muskellunge (''E. masquinongy'')]]
 
[[File:Muskellunge USFWS.jpg|thumb|320px|right|Muskellunge (''E. masquinongy'')]]
 +
Members of the Esociformes, which also include the mudminnows (family Umbridae), are characterized by posteriorly located dorsal and anal fins, the lack of an adipose fin, toothless maxilla but in the gape of the mouth, and no pyloric caeca. Members of the family Esocidae, the esocids, are characterized by a forked caudal fin with 40 to 50 rays, a complete lateral line, 10 to 20 branchiostegal rays, nasals present, snout produced, and 43 to 67 vertebrae. The other family of Esociformes, the Umbridae, differs in that the mudminnows have a rounded caudal fin with 20 to 30 rays, a lateral line that is faint or absent, nasals absent, snout not produced, only 32 to 42 vertebrae, and 5 to 8 branchiostegal rays (Nelson 2006).
 +
 
The only extant genus of Esocidae is ''Esox''.  
 
The only extant genus of Esocidae is ''Esox''.  
  

Revision as of 20:17, 27 June 2013

Esox
Fossil range: Upper Cretaceous–Recent
[1]
Northern pike (E. lucius)
Northern pike (E. lucius)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Superorder: Protacanthopterygii
Order: Esociformes
Family: Esocidae
G. Cuvier, 1817
Genus: Esox
Linnaeus, 1758
Type species
Esox lucius
Linnaeus, 1758

Esox is a genus of freshwater ray-finned fish, whose members are known as pikes, pickerels, and muskellunge, and are characterized by pronounced snout, forked caudal fin, complete lateral line, posteriorly located dorsal and anal fins, and no adipose fin. Esox is the only extant genus in the pike family Esocidae of the order Esociformes (with the mudminnows, family Umbridae, also placed in the order).

Of the five traditionally recognized extant species in Esox, the type species, E. lucius (the northern pike) is the only circumpolar member, while E. richerti (Amur pike) is located in Siberia, and three species, E. masquinongy (muskellunge), E. niger (chain pickerel), and E. americanus (redfin and grass pickerel) are restricted to eastern North America. However, in 2011 it was claimed that the northern pike was not the only extant esocid species in Europe, with at least one new species being delineated as distinct. This species, which was restricted to freshwater habitats in Southern Europe, was variously identified as E, flaviae or E. cisalpinus and sometimes called the southern pike.

The largest member of the genus and family is the muskellunge (E. masquinongy) which reaches 1.8 meters (6 feet).

sports fishing, food, etc.

Overview and description

The Esox genus is the only living genus in the pike family, Esocidae, in the order Esociformes of the class Actinopterygii.

Muskellunge (E. masquinongy)

Members of the Esociformes, which also include the mudminnows (family Umbridae), are characterized by posteriorly located dorsal and anal fins, the lack of an adipose fin, toothless maxilla but in the gape of the mouth, and no pyloric caeca. Members of the family Esocidae, the esocids, are characterized by a forked caudal fin with 40 to 50 rays, a complete lateral line, 10 to 20 branchiostegal rays, nasals present, snout produced, and 43 to 67 vertebrae. The other family of Esociformes, the Umbridae, differs in that the mudminnows have a rounded caudal fin with 20 to 30 rays, a lateral line that is faint or absent, nasals absent, snout not produced, only 32 to 42 vertebrae, and 5 to 8 branchiostegal rays (Nelson 2006).

The only extant genus of Esocidae is Esox.

Esocids

Esox is a genus of freshwater fish, the only living genus in the family Esocidae—the esocids which were endemic to North America, Europe and Eurasia during the Paleogene through present.[2]

The type species is E. lucius, the northern pike. The species of this genus are known as pike and pickerel, and in heraldry they are usually called lucy.[3][4]

        • The Pollachius genus is one of twelve genera placed in the subfamily Gadinae (cods and haddock) of the cod family Gadidae (16 genera) of the order Gadiformes and the class Actinopterygii. Members of the Gadiformes are characterized by lacking true spines in the fins, generally long dorsal and anal fins, usually cycloid (rarely ctenoid) scales, a swim bladder without pneumatic duct, and pelvic fins (when present) below or in front of the pectorals. Members of the almost exclusively marine family Gadidae (one Holarctic freshwater species) are typified by having the first dorsal fin posterior to the head, the head of the vomer toothed, and the swim bladder not connected with auditory capsules. Members of the Gadinae subfamily are characterized by having three dorsal and two anal fins, an egg without an oil globule, usually a chin barbel, and a slightly forked or truncate caudal fin (Nelson 2006). In some taxonomic schemes, Gadinae is raised to the family level.

http://www.fishbase.org/summary/258



The big pike species are native to the Palearctic and Nearctic ecozones, ranging across northern North America and from Western Europe to Siberia in Eurasia. They have been found in many urban lakes in Western Europe, reported to be in the Rostrum (Lucerne) and the Serpentine, (London).

Pike can grow to a maximum recorded length of 1.83 meters (6.0 ft), reaching a maximum recorded weight of 35 kilograms (77 lb). The UK record pike of 21 kilograms (46 lb) was caught on a Creek Chub Pikie lure by Roy Lewis at Llandegfedd Reservoir in Wales in 1992.[5] Individuals have been reported to reach 30 years in age. All pike over 5 kilograms (11 lb) in body weight are females. They have the elongated, torpedo-like form of predatory fishes, with sharply-pointed heads and sharp teeth. Their coloration is typically grey-green with a mottled or spotted appearance with stripes along their back, perfectly camouflaged among weeds. Individual pike marking patterns are unique, like fingerprints.

Taxonomy

traditionally, five species recogtnized. http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=162138 recognizes only 5 species


However, the taxonomy remains a bit unclear with the delineation of a new species of Esox in 2011 by *****. This species was held to be different from the northern pike (E. ) according to . Named Esox flaviae It is located in http://www.plosone.org/article/info:doi/10.1371/journal.pone.0025218

In 2011, another species was likewise identified in Southern Europe by *** and labelled ****. However, it appears that these species are synonyms (ITIS ***). Fishbase *** http://www.fishbase.org/summary/FamilySummary.php?ID=74 Esox cisalpinus Bianco & Delmastro, 2011 accepted senior original 1 Esox flaviae Lucentini, Puletti, Ricciolini, Gigliarelli, Fontaneto, Lanfaloni, Bilò, Natali & Panara, 2011 ambiguous questionable original 1


Most generally there are currently six recognized extant species (references):

There are currently seven recognized species in this genus: [6][7]

  • Esox americanus
    • Esox americanus americanus J. F. Gmelin, 1789 (Redfin pickerel)
    • Esox americanus vermiculatus Lesueur, 1846 (Grass pickerel)
  • Esox cisalpinus Bianco & Delmastro, 2011
  • Esox flaviae Lucentini, Puletti, Ricciolini, Gigliarelli, Fontaneto, 2011 (Southern Pike)
  • Esox lucius Linnaeus, 1758 (Northern pike)
  • Esox masquinongy Mitchill, 1824 (Muskellunge)
  • Esox niger Lesueur, 1818 (Chain pickerel)
  • Esox reichertii Dybowski, 1869 (Amur pike)


northern pike is the only native esocid in Europe


However, in 2011, two new species were claimed, Esox flaviae (southern pike) and Esox cisalpinus (), both restricted to freshwater habitats in Southern Europe, although these may be synonymous species.

Southern pike (Esox flaviae) is a freshwater fish restricted to freshwater habitats in Southern Europe.

Esox cisalpinus

Esox cisalpinus  Bianco & Delmastro, 2011	

http://www.fishbase.org/summary/66315 Europe: Central and northern Italy; potentially present in other European water bodies in the Mediterranean area as those on the NE shores of the Adriatic and in Mediterranean France.

File:Esox cisalpinus.jpg

E. flaviae Lucentini et al., 2011 is probably a synonym of this species.

http://www.plosone.org/article/info:doi/10.1371/journal.pone.0025218 tion: Lucentini L, Puletti ME, Ricciolini C, Gigliarelli L, Fontaneto D, et al. (2011) Molecular and Phenotypic Evidence of a New Species of Genus Esox (Esocidae, Esociformes, Actinopterygii): The Southern Pike, Esox flaviae. PLoS ONE 6(12): e25218. doi:10.1371/journal.pone.0025218

(2011) Molecular and Phenotypic Evidence of a New Species of Genus Esox (Esocidae, Esociformes, Actinopterygii): The Southern Pike, Esox flaviae.

There is one fossil species, Esox kronneri Grande, 1999 known from the Eocene of the Green River formation.[8]

Etymology

The generic name Esox (pike fish) derives from the Greek ίσοξ (a kind of fish), itself a word of Celtic origin related to the Welsh eog and Irish Gaelic iasc (fish). Pliny uses the Latin form esox in reference to a large fish in the Rhine normally identified with lax (salmon). It is likely that Carolus Linnaeus's application of Esox to the pike is thus a misnomer.

A young E. lucius specimen—a "Chain pickerel" in the original sense—in an aquarium.

The English common name "pike" is an apparent shortening of "pike-fish", in reference to its pointed head, Old English píc originally referring to a pickaxe. The plural of pike is pike.

A northern English and Lowland Scots name for the pike, ged, similarly derives from Old Norse gaddr (spike) (cf. the modern Swedish name for the pike, gädda, the Danish "gedde", the Norwegian "gjedde" and Scottish Gaelic: geadais). The Dutch name for the pike (snoek) has been given to a wide variety of fish reminding sailors of the pike (see snoek, snook).

The English "pike" originally referred specifically to the adult fish, the diminutive form "pickerel" (now used to name some of the smaller pike, E. americanus and E. niger) referring to the young. The walleye (Sander vitreus) is sometimes called a pickerel by Gerard, but it is unrelated to the pike, being a member of the perch family (family Percidae). Pike are not to be confused with the unrelated pikeminnows (traditionally, and perhaps better, known as squawfish) of genus Ptychocheilus (family Cyprinidae) or pikeperch (Sander lucioperca) which is more akin to walleye than to pike. Pike are also called "Jackfish" in North America and informally "Slough Shark" in Western Canada.

Diet

Pike in Haus des Meeres, Vienna

Pike feed on a wide range of food sources, predominantly smaller shoal fish. Pike are also cannibalistic, sometimes preying upon smaller members of their own species.

They will also prey on insects and amphibians such as newts or frogs in times when their usual food is scarce, and occasionally on small mammals like moles or mice when caught water-borne. Small birds such as ducklings may become a target for hungry pike. Pike are also known to prey on swimming snakes.

They are, however, undeserving of their reputation for being overly vicious predators. There are few substantiated incidents of pike 'attacks' on people. Pike's further reputation as a pest seems to lie predominately amongst anglers who seek more desirable species.

Angling

Sign on the River Shannon, Ireland
A caught pike

Effective methods for catching this hard-fighting fish include dead baits, live baits, and lure fishing. Pike can easily be damaged when handled since they are not as robust as their reputation would suggest. Colour of lure can be influenced by water clarity and weather conditions. Since pike have numerous sharp teeth it is wise to take extreme care when unhooking them. The use of a wet leather gauntlet and surgical forceps to remove hooks is highly recommended on safety grounds. If practising catch and release fishing, care for the pike should be the pike angler's utmost concern. The formerly recommended practice of grasping a pike by its eye sockets (tragically interpreted as "its eyes") resulted in countless released pike that quickly died from inability to see prey any longer.

The current recommended method of grasping pike is to close the hand firmly over the gill covers, and to make the period of handling as short as possible before release. Grabbing a pike by the gill covers is not feasible when a pike is very big, but it is easy to handle a pike by inserting the fingers at the bottom of the gill opening and grabbing the lower jaw. Big pike should also be supported at the belly. When a pike is held this way it is also easier to keep the mouth open to remove a hook. Many anglers now use special grips to grab the pike's front lower jaw, which can add to the safety of an anglers because of the danger imposed by the hooks of the lure or tackle and the pike's teeth. The Pike Anglers Club was formed in 1977 to campaign for the preservation of pike and the sport of pike fishing.

A practice known as gut hooking was previously widely used in catching pike. Upon taking the bait, the pike will hold it for a short time in its mouth as it moves off. The pike will then, usually, turn the bait in its mouth, so that it sits in alignment with its throat to ease swallowing. It is recommended that when pike fishing the process is not allowed to go this far and a strike is recommended as soon as a bite is indicated. For this it is necessary to attach hooks on the head side as well as the middle of the baitfish. Otherwise, what is known as gut hooking will result, which will normally kill or seriously injure the fish. Dutch research shows that cutting the line immediately when the fish is gut hooked will still give low mortality (14%). The hooks in the gut or stomach were either encapsulated or removed from the body.[9]

Other methods of catching and handing pike that are now frowned upon are the gaff and the gag. The gaff is a metal hook on the end of a pole used to hook through the fish's body in place of a more humane landing net. A gag is a device for holding open the pike's mouth whilst unhooking. These are now illegal in Scotland, as they put a huge amount of pressure on a pike's jaw, thus causing irreparable damage.

Catching a 10 kg pike, kymppihauki (Finnish for "ten pike"), is considered as "earning one's spurs" as a fisherman in Finland.

In mythology and literature

In the Finnish Kalevala, Väinämöinen creates a kantele (string instrument) from the jawbone of the pike.

Russian mythology holds that the pike is one of several forms assumed by evil water spirits called vodyanoy, and a ravenous mythical pike is traditionally blamed for decimating the fish population in the Sheksna River. Russian fairy tales, on the other hand, also tell about an old wise pike that can fulfil wishes of the one who catches it, if its catcher releases it back into its habitat. [citation needed]

In the beginning of Boris and Arkady Strugatsky's book, Monday Begins on Saturday, a wise centuries-old pike appears in the story when main character Sasha arrives at Baba Yaga's house in Solovets.

In the 10th episode of Monarch of the Glen series 3, Hector MacDonald fights a pike that has infiltrated his loch and has eaten all the fish.

In the book Redwall, Cluny the Scourge has only one eye, the other having been lost to a pike who was then killed by Cluny.

Submarines and tanks

Two United States Navy submarines have been named Pike, SS-6 of 1903 and SS-173 of 1935, and three, SS-22 of 1912, SS-177 of 1936, and SS-524 of 1944, named Pickerel. In addition, the Soviet submarines known to NATO as the Victor III class were called the Shchuka (Щука, "pike") class, in Russian. The Soviet Iosif Stalin tank (IS-3) was also nicknamed Shchuka, in reference to its sharply pointed hull front.

Heraldry

In heraldry, the pike is called a lucy.[3] It is usually blazoned either naiant (swimming), embowed (bowed) or hauriant (jumping), though pairs of lucies may appear addorsed (back to back), as in the arms of the Finnish town of Uusikaupunki, Finland. (Argent, two lucies addorsed azure).[citation needed]

Uusikaupunki coat of arms
Coat of arms of Gimte, in Lower Saxony, Germany.

References
ISBN links support NWE through referral fees

  1. "Esocidae". FishBase. Ed. Ranier Froese and Daniel Pauly. December 2008 version. N.p.: FishBase, 2008.
  2. PaleoBiology Database: Esox, basic info
  3. 3.0 3.1 Fox-Davies, Arthur Charles (1909). A Complete Guide to Heraldry. London: T.C. & E.C. Jack. 
  4. Woodward, John (1892). A treatise on heraldry, British and foreign. Edinburgh: W. & A.K. Johnston. 
  5. Record pike
  6. "Esox". FishBase. Ed. Ranier Froese and Daniel Pauly. February 2012 version. N.p.: FishBase, 2012.
  7. Lucentini, Puletti, Ricciolini, Gigliarelli, Fontaneto, Lanfaloni, Bilò, Natali, Panara (2011). Molecular and Phenotypic Evidence of a New Species of Genus Esox (Esocidae, Esociformes, Actinopterygii): The Southern Pike, Esox flaviae. PLoS ONE 6(12): e25218. doi:10.1371/journal.pone.0025218
  8. Grande, L. (1999): The First Esox (Esocidae: Teleostei) from the Eocene Green River Formation, and a Brief Review of Esocid Fishes. Journal of Vertebrate Paleontology Vol. 19, No. 2 (Jun. 14, 1999), pp. 271-292
  9. http://www.hsvdekleuter.nl/documents/snoek-kennis.pdf
  • Nelson, J. S. 2006. Fishes of the World, 4th edition. Hoboken, NJ: John Wiley & Sons. ISBN 0471250317.

External links

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